Australasian/Oceanian Diptera Catalog -- Web Version

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40. Family BOMBYLIIDAE

By Neal L. Evenhuis

Members of the Bombyliidae, commonly called bee flies, are most frequently found in arid and semiarid portions of the world, with a few specialized forms occurring in moist tropical climates. Adults range in size from 1 to 21 mm in length and are commonly found on flowers, hovering in bright sunlight, or resting on warm, bare ground. Many, if not all, are pollen and nectar feeders, with some acting as primary pollinators of certain species of desert wildflowers.

Species found in Australia include some of the most striking species known; many have large, pictured wings and brightly colored cuticular pigmentation or vestiture. There is a great deal of endemism exhibited in the taxa from Australia and the Oceanic islands (Evenhuis 1982b). The Oriental Region seems to be a source for many of the Oceanic taxa, but some of the Australian fauna (viz., Comptosiini and Dischistini) show possible Transantarctic relationships because related genera of the 2 tribes are also found in central and southern Chile. There are over 400 species known from the areas covered in this catalog. As the fauna from these areas, especially the Australian, becomes better known, this number could easily double or even triple.

Immatures are infrequently encountered. Larvae of some genera are predators of orthopteroid egg pods, most others are parasitoids on immature stages of holometabolous orders of insects (DuMerle 1975) or are free-living in soil. Ectoparasitic representatives from the regions covered here include Exhyalanthrax on pupiparous higher Diptera, Systropus on saddleback moths (Limacodidae), Geron on Psychidae, and the ubiquitous Anthrax on a variety of hosts, primarily aculeate Hymenoptera. Hyperparasites occurring in these areas include Exoprosopa and Ligyra on Tiphiidae and Scoliidae (Hymenoptera), which are parasites of grubs of Scarabaeoidea. Hardy (1921c) recorded Ligyra satyrus as hyperparasitic on an unidentified asilid.

Classification of the family follows, for the most part, that provided by Yeates (1994) with some exception, which follow that of Evenhuis (1991). The Mythicomyiinae of Yeates has been raised to family level here following Zaitzev (1991) and Evenhuis (1993, 1994). The subfamily Bombyliinae is represented in these regions only by the tribe Dischistini. Species previously described in the genus Bombylius should be transferred to other genera after types have been examined. Due to the poor level of taxonomic knowledge of these flies, previous synonymies for species in many genera have been included here only when substantiated by examination of type material.

The only comprehensive work on the bombyliid fauna from these regions is the revisionary work of Roberts (1928a,b, 1929). The late S.J. Paramonov contributed many papers on the Australian Bombyliidae, and many of his unfinished manuscripts on various genera are deposited in CSIRO, Canberra. David Yeates has begun a series of studies at the subfamilial and generic level of the Australian Bombyliidae.Chris Lambkin et al.(2003) revised the Australian Exoprosopini.
Refs.: Roberts (1928a,b, 1929, keys, rev. Australian spp.), Edwards (1934, rev. Lomatiinae), Paramonov (1967. rev. Australian Ligyra), Hull (1973, world gen.), DuMerle (1975, immat. biol., bibliogr.), Evenhuis (1979b, rev. Australasian/Oceanian Geron; 1982, Oceanian zoogeogr.; 1983b, bibliogr.; 1988a, rev. Brachyanax; 1988b, phylog. Phthiriinae; 1991, classif., world gen.; 1994, cat. fossil Bombyliidae), Greathead (1988, phylog. Ecliminae), Yeates (1988, rev. Oncodosia; 1994, fam. classif.), Yeates & Hagen (1988; rev. Australian Systropus); Yeates & Lambkin (1998, rev. Australian Anthracini); Evenhuis & Greathead (1999, world cat.); Lambkin et al. (2003, rev. Australian Exoprosopini).

Subfamily TOXOPHORINAE
Tribe SYSTROPODINI

Genus SYSTROPUS Wiedemann

SYSTROPUS Wiedemann, 1820a: 18. Type species: Systropus macilentus Wiedemann, 1820, orig. des. (on plate).
SYSTROPUS Henning, 1832: 335. Type species: Systropus macilentus Henning, 1832 [preocc., = Systropus macilentus Wiedemann, 1820], mon. [Preocc. Wiedemann, 1820.]
CEPHENE Latreille, 1825: 496. Vernacular name without nomenclatural status.
CEPHENUS Berthold, 1827: 506 (unnec. n. n. for Systropus).
XYSTROPHUS Agassiz, 1846b: 393 (unnec. emend. of Systropus).
CEPHENIUS Enderlein, 1926b: 70 (as gen.). Type species: Systropus studyi Enderlein, 1926, orig. des.
SYSTROPHUS, error for Systropus.
SYSTROPHOPUS, error for Systropus.
doddi Roberts, 1929: 555. Australia (Qld).
flavoornatus Roberts, 1929: 554. Australia (Qld).
tipuloides Westwood, 1876c: 576. Indonesia (Maluku).

Genus ZACLAVA Hull

ZACLAVA Hull, 1973: 250. Type species: Systropus clavifemoratus Hardy, 1921, orig. des.
clavifemorata Hardy, 1921c: 61 (Systropus). Australia (NSW); Australia (Qld).
kraussi Evenhuis, 1979a: 87. New Caledonia (Loyalty Is); New Caledonia (New Caledonia).
minima Roberts, 1929: 557 (Dolichomyia). Australia (Qld); Australia (NSW).
occidentis Roberts, 1929: 557 (Dolichomyia). Australia (WA).

Tribe TOXOPHORINI

Genus TOXOPHORA Meigen

TOXOPHORA Meigen, 1803: 270. Type species: Toxophora maculata Meigen, 1804 [preocc., = ?Asilus fasciculatus Villers, 1789], subs. mon. Meigen, 1804: 273.
ENICONEVRA Macquart, 1840: 110 [1841: 388]. Type species: Eniconevra fuscipennis Macquart, 1840, mon.
ENICONEURA Agassiz, 1846a: 13 (unjust. emend. of Eniconevra).
HENICONEVRA Agassiz, 1846b: 138 (unjust. emend. of Eniconevra).
HENICONEURA Scudder, 1882: 159 (unjust. emend. of Eniconevra).
compta Roberts, 1929: 559. Australia (Qld).

Tribe GERONTINI

Genus GERON Meigen

GERON Meigen, 1820: 223. Type species: Geron gibbosus Meigen, 1820 [= Bombylius gibbosus Olivier, 1789], des. Duponchel in d'Orbigny, 1845b: 206.

Subgenus GERON Meigen

GERON Meigen, 1820: 223 (as gen.). Type species: Geron gibbosus Meigen, 1820 [= Bombylius gibbosus Olivier, 1789], des. Duponchel in d'Orbigny, 1845b: 206.
aaptis Evenhuis, 1979b: 19. New Caledonia (Loyalty Is).
aesion Evenhuis, 1979b: 20. New Caledonia (New Caledonia); PNG (PNG).
australis Macquart, 1840: 118 [1841: 396]. Australia (NSW); widesp. Australia.
     hilaris White, 1916b: 188. Australia (Tas).
balpi Evenhuis, 1979b: 23. Australia (WA); Australia (NT).
cabon Evenhuis, 1979b: 24. Australia (WA).
chrysonotum Evenhuis, 1979b: 25. Australia (WA).
europacificus Evenhuis, 1979b: 27. French Polynesia (Society Is).
flavocciput Evenhuis, 1979b: 24. Australia (ACT).
fumipennis Evenhuis, 1979b: 30. Australia (Qld); Australia (ACT, NSW).
fusciscelis Evenhuis, 1979b: 30. French Polynesia (Society Is).
inflatus Evenhuis, 1979b: 31. Australia (WA).
karakara Evenhuis, 1979b: 32. Australia (WA).
malekulanus Evenhuis, 1979b: 33. Vanuatu.
mononesensis Evenhuis, 1979b: 33. Indonesia (Maluku).
mysticus Evenhuis, 1979b: 33. Vanuatu.
neutralis Evenhuis, 1979b: 35. New Caledonia (New Caledonia); New Caledonia (Loyalty Is).
paramonovi Evenhuis, 1979b: 36. Australia (NSW).
terminatus Evenhuis, 1979b: 37. Vanuatu.
validus Evenhuis, 1989: 362 (n. n. for robustus Evenhuis). Australia (NT); Australia (Qld).
     robustus Evenhuis, 1979b: 37. Australia (NT). [Preocc. Cresson, 1919.]

Subgenus PLICHTAMYIA Evenhuis

PLICHTAMYIA Evenhuis, 1979b: 39. Type species: Geron dispar Macquart, 1850, orig. des.
albihalteris Evenhuis, 1979b: 39. Solomon Is; PNG (Bismarck Arch).
amboinensis Evenhuis, 1979b: 40. Indonesia (Maluku).
bowdeni Evenhuis, 1979b: 41. PNG (Bismarck Arch).
dispar Macquart, 1850: 426(122). Australia (Tas); Australia (ACT, NSW, Qld, Vic).
     cothurnatus Bigot, 1892b: 374. Australia (Tas).
erythroccipitalis Evenhuis, 1979b: 44. Solomon Is; PNG (Bougainville I).
hemifuscis Evenhuis, 1979b: 45. PNG (PNG).
nigralis Roberts, 1929: 571. Australia (NSW); Australia (Qld).
nigrocciput Evenhuis, 1979b: 48. Australia (Qld); Australia (ACT, NSW, SA, Tas), PNG (PNG).
opacus Bowden, 1971d: 312. PNG (Bismarck Arch); Solomon Is.
paraustralicus Evenhuis, 1979b: 51. Indonesia (Maluku).
robertsi Evenhuis, 1979b: 52. Australia (NSW); Australia (Qld).
sparsus Bowden, 1971d: 312. PNG (Bismarck Arch).
westralicus Evenhuis, 1979b: 54. Australia (WA).

Unplaced species of Geron Meigen

simplex Walker, 1858b: 90. Indonesia (Maluku); ?PNG (PNG).
tenue Walker, 1857: 147. "Country unknown" [? Australasian].

Subfamily BOMBYLIINAE
Tribe DISCHISTINI

Genus APIFORMYIA Yeates

APIFORMYIA Yeates, 2008: 32. Type species: Apiformyia Yeates, 2008, orig. des..
australis Yeates, 2008: 32. Australia (NT).

Genus BROMOGLYCIS Hull

BROMOGLYCIS Hull, 1971: 5. Type species: Bromoglycis robustus Hull, 1971, mon.
robustus Hull, 1971: 7. Australia (Qld).

Genus BRYCHOSOMA Hull

BRYCHOSOMA Hull, 1973: 102. Type species: Brychosoma pulchrum Hull, 1973 [= Bombylius pictipennis Macquart, 1850], mon.
areolatus, error for aureolatum.
hilare Walker, 1849: 274 (Bombylius). Australia (WA); Australia (NSW, Qld).
pictipenne Macquart, 1850: 422(118) (Bombylius). Australia, "côte orientale"; Australia (NSW, Qld).
     pulchrum Hull, 1973: 104. Australia (Qld).

Genus CHORISTUS Walker

CHORISTUS Walker, 1852: 197. Type species: Choristus bifrons Walker, 1852, mon.
ANASTOECHUS, authors, not Osten Sacken, 1877, misid.
annexus Roberts, 1928b: 434 (Anastoechus). Australia (WA).
bifrons Walker, 1852: 198. Australia (NSW); Australia (WA).
     perspicuus Roberts, 1928b: 433 (Anastoechus). Australia (WA).

Genus CRYOMYIA Hull

CRYOMYIA Hull, 1973: 131. Type species: Cryomyia argyropila Hull, 1973, mon.
argyropila Hull, 1973: 131. Australia (WA).
perparva Roberts, 1928b: 453 (Anastoechus). Australia (Qld).

Genus ERISTALOPSIS Evenhuis

SYRPHOIDES Evenhuis, 1983a: 206. Type species: Bombylius byrrhus Bowden, 1971, orig. des. [Preocc. Hippa, 1968.]
ERISTALOPSIS Evenhuis, 1985a: 289 (n. n. for Syrphoides).
byrrha Bowden, 1971c: 297 (Bombylius). Australia (WA).
robertsi Paramonov, 1934: 31 (Bombylius). Australia (Qld).
rubriventris Bigot, 1892b: 365 (Bombylius). Australia (NSW).

Genus EUSURBUS Roberts

EUSURBUS Roberts, 1929: 580. Type species: Bombylius crassilabris Macquart, 1855, orig. des.
altus Walker, 1849: 288 (Bombylius). Australia (WA).
crassilabris Macquart, 1855: 97(77) (Bombylius). Australia (NSW).
pinguis Walker, 1849: 290 (Bombylius). Australia (WA).

Genus LAURELLA Hull

LAURELLA Hull, 1971: 3. Type species: Laurella auripila Hull, 1971 [= Dischistus formosus Roberts, 1928], orig. des.
formosa Roberts, 1928b: 451 (Dischistus). Australia (Qld).
     auripila Hull, 1971: 3. Australia (Qld).
pallidoventris Roberts, 1928b: 452 (Dischistus). Australia (Tas).

Genus MANDELLA Evenhuis

MANDELLA Evenhuis, 1983a: 207. Type species: Systoechus albohirtus Roberts, 1928, orig. des.
SYSTOECHUS, authors, not Loew, 1855, misid.
albohirta Roberts, 1928b: 422 (Systoechus). Australia (SA).
cinctiventris Roberts, 1928b: 424 (Systoechus). Australia (Qld).
flavovillosa Roberts, 1928b: 423 (Systoechus). Australia (Qld).
pallida Roberts, 1928b: 422 (Systoechus). Australia (Qld).
rubida Roberts, 1928b: 421 (Systoechus). Australia (Qld); Australia (NSW).

Genus MEOMYIA Evenhuis

MEOMYIA Evenhuis, 1983a: 210. Type species: Bombylius platyurus Walker, 1849 [= Bombylius fasciculatus Macquart, 1840], orig. des.
SYSTOECHUS, authors, not Loew, 1855, misid.
albiceps Macquart, 1848: 196(36) (Bombylius). "Nouvelle-Hollande"; Australia (NSW, Qld, Vic, WA).
callynthrophora Schiner, 1868: 137 (Systoechus). Australia (NSW); Australia (Qld).
fasciculata Macquart, 1840: 93 [1841: 371] (Bombylius). "Patrie inconnue" [= Australia (NSW)]; Australia (Qld, SA, Tas, Vic, WA).
     platyurus Walker, 1849: 286 (Bombylius). Australia (WA).
     crassus Walker, 1849: 287 (Bombylius). "New Holland.".
     notatipennis Macquart, 1855: 98(78) (Bombylius). Australia (NSW).
     platyrus, error for platyurus.
pausaria Jaennicke, 1867: 348 (Systoechus). "Australia."
penicillata Macquart, 1850: 422(118) (Bombylius). Australia, "côte orientale."
punctipennis Thomson, 1869: 487 (Bombylius). Australia (NSW).
sericans Macquart, 1850: 420(116) (Bombylius). Australia, "côte orientale"; Australia (Qld, WA).
tetratricha Walker, 1849: 291 (Bombylius). Australia (WA).
vetusta Walker, 1849: 286 (Bombylius). "New Holland."

Genus PILOSIA Hull

PILOSIA Hull, 1973: 137. Type species: Pilosia flavopilosa Hull, 1973, mon.
flavopilosa Hull, 1973: 137. Australia (WA).
immutata Walker, 1849: 292 (Bombylius). Australia (WA).

Genus SISYROMYIA White

SISYROMYIA White, 1916b: 192, 197. Type species: Bombylius auratus Walker, 1849, des. Bezzi, 1924a: 7.
BOMBYLIUS, authors, not Linnaeus, 1758, misid.
albavitta Macquart, 1850: 421(117) (Bombylius). Australia, "côte orientale"; Australia (NSW, Qld, WA).
     albovitta, albovittata, errors for albavitta.
aurata Walker, 1849: 289 (Bombylius). Australia (Tas) & (WA); Australia (NSW, SA).
binghi Evenhuis, 1989: 364. (n. n. for brevirostris Macquart). Australia (?NSW).
     brevirostris Macquart, 1850: 423(119) (Bombylius). "Tasmanie", error [prob. = Australia (NSW)]. [Preocc. Olivier, 1789.]
crassirostris Macquart, 1850: 423(117) (Bombylius). Australia, "côte orientale" [prob. = Australia (NSW)].
decorata Walker, 1849: 291 (Bombylius). Australia (WA).
eulabiata Bigot, 1892b: 366 (Systoechus). Australia (Tas).
limbata Bigot, 1892b: 369 (Sparnopolius). "Australie"; Australia (SA, WA).
lobalis Thomson, 1869: 487 (Bombylius). Australia (NSW).
loewii Jaennicke, 1867: 346 (Bombylius). "Australia."
pycnorrhyncha Thomson, 1869: 486 (Bombylius). Australia (NSW).
rutila Walker, 1849: 289 (Bombylius). Australia (NSW).
scutellaris Thomson, 1869: 480 (Bombylius). Australia (NSW).

Genus STAUROSTICHUS Hull

STAUROSTICHUS Hull, 1973: 76, 104. Type species: Staurostichus chiastoneura Hull, 1973, orig. des.
BOMBYLIUS, authors, not Linnaeus, 1758, misid.
albicinctus Macquart, 1847a: 54 [1847b: 70] (Bombylius). Australia (Tas).
alienus Hardy, 1942: 204 (Bombylius; n. n. for pulchellus Roberts). Australia (NSW).
     pulchellus Roberts, 1928b: 441 (Bombylius). Australia (NSW). [Preocc. Loew, 1863.]
chiastoneura Hull, 1973: 104. Australia (Qld).
chrysendetus White, 1916b: 195 (Bombylius). Australia (Tas).
consobrinus Macquart, 1847a: 54 [1847b: 70] (Bombylius). "Nouvelle-Hollande" & Australia (Tas).
dulcis Roberts, 1928b: 440 (Bombylius). Australia (Qld).
fuscanus Macquart, 1850: 423(119) (Bombylius). Australia (Tas); Australia (Vic).
limbipennis Macquart, 1840: 96 [1841: 374] (Bombylius). "Patrie inconnue" [= Australia]; Australia (NSW).
     australianus Bigot, 1892b: 364 (Bombylius). Australia (NSW).
matutinus Walker, 1849: 281 (Bombylius). "New Holland."
proprius Roberts, 1928b: 436 (Bombylius). Australia (NSW).
sciotus Bowden, 1971c: 296 (Bombylius). Australia (Tas).
tenuicornis Macquart, 1850: 423(116) (Bombylius). Australia (Tas).
viduus Walker, 1852: 199 (Bombylius). Australia (Tas).
     palliolatus White, 1916b: 194 (Bombylius). Australia (Tas).

Unplaced species of Dischistini

aureolatus Walker, 1857: 145 (Bombylius). "Australia"; Australia (NSW, Qld).
australis Guérin-Méneville, 1831: pl. 20, fig. 4 [1838: 294] (Bombylius). Australia (NSW).
distinctus Walker, 1852: 201 (Bombylius). Australia (NSW & WA).
leucopygus Wulp, 1885c: 86 (Systoechus). Australia (SA).
nigracinctus Roberts, 1929: 581 (Eusurbus). Australia (WA).
     nigrivinctus, error for nigracinctus.
primogenitus Walker, 1849: 292 (Bombylius). Australia (WA).
probellus Hardy, 1942: 204 (Bombylius; n. n. for bellus Roberts). Australia (NSW).
     bellus Roberts, 1928b: 439 (Bombylius). Australia (NSW). [Preocc. Philippi, 1865.]
succandidus Roberts, 1928b: 438 (Bombylius). Australia (NSW).
tenuirostris Roberts, 1928b: 437 (Bombylius). Australia (Qld).
walkeri Evenhuis, 1978a: 101 (Bombylius; n. n. for nanus Walker). Australia (WA).
     nanus Walker, 1849: 278 (Bombylius). Australia (WA). [Preocc. Meigen, 1838.]

Tribe ECLIMINI

Genus MARMASOMA White

MARMASOMA White, 1916b: 184, 188. Type species: Marmasoma sumptuosa White, 1916, mon.
sumptuosum White, 1916b: 190. Australia (Tas) & (Vic).

Genus THEVENETIMYIA Bigot

THEVENEMYIA Bigot, 1875a: 196 [1875b: clxxiv]. Type species: Thevenemyia californica Bigot, 1875, mon.
THEVENETIMYIA Bigot, 1892b: 339 (emend. of Thevenemyia). Type species: Thevenemyia californica Bigot,
1875, aut.
ECLIMUS, authors, not Loew, 1844, misid.
australiensis Hall, 1969: 25. Australia (NSW).
furvicostata Roberts, 1929: 578 (Eclimus). Australia (Qld).
longipalpus Hardy, 1921c: 63 (Eclimus). Australia (NSW); Australia (WA).
mimula Hall, 1969: 61. Not given [Australia]; Australia (Qld).
nigrapicalis Roberts, 1929: 576 (Eclimus). Australia (Qld); Australia (ACT).
tenta Hall, 1969: 75. Australia (NSW); Australia (Qld).

Genus TILLYARDOMYIA Tonnoir

TILLYARDOMYIA Tonnoir, 1927: 101. Type species: Tillyardomyia gracilis Tonnoir, 1927, mon.
gracilis Tonnoir, 1927: 103. NZ (NZ).

Subfamily PHTHIRIINAE
Tribe PHTHIRIINI

Genus ACREOTRICHUS Macquart

ACREOTRICHUS Macquart, 1850: 425(121). Type species: Acreotrichus gibbicornis Macquart, 1850 [= Bombylius antecedens Walker, 1849], orig. des.
ACEOTRICHUS. Incorr. orig. spell. of Acreotrichus [Macquart, 1850: 315 (11)].
antecedens Walker, 1849: 293 (Bombylius). "Australia"; Australia (NSW, NT, WA).
     gibbicornis Macquart, 1850: 425(121). Australia, "côte orientale."
     fusicornis Macquart, 1850: 426(122). Australia, "côte orientale."
inappendiculatus Bigot, 1892b: 366. "Australie."
lineifer Walker, 1857: 146 (Phthiria). "Australia"; Australia (NSW).

Genus AUSTRALIPHTHIRIA Evenhuis

AUSTRALIPHTHIRIA Evenhuis, 1986: 8. Type species: Phthiria hilaris Walker, 1852, orig. des.
albocapita Roberts, 1929: 573 (Phthiria). Australia (WA); Australia (NT).
hilaris Walker, 1852: 194 (Phthiria). Australia (NSW); Australia (ACT).
pallipes Bigot, 1892b: 367 (Phthiria). "Australie."

Genus PYGOCONA Hull

PYGOCONA Hull, 1973: 195 (Phthiria subg.). Type species: Phthiria flavicincta Hull, 1973 [= Phthiria flava Hardy, 1933], orig. des.
flava Hardy, 1933b: 415 (Phthiria). Australia (Vic).
     flavicincta Hull, 1973: 195, 199 (Phthiria). Not given [Australia].
nigrina Hardy, 1933b: 415 (Phthiria). Australia (Vic); Australia (NSW).

Subfamily CYTHEREINAE

Genus NEOSARDUS Roberts

NEOSARDUS Roberts, 1929: 560. Type species: Neosardus principius Roberts, 1929, orig. des.
australis Yeates, 1996a: 53. Australia (SA); Australia (Qld).
circumdatus Roberts, 1929: 561. Australia (NSW); Australia (Qld).
cosmus Yeates, 1996a: 54. Australia (WA).
danielsi Yeates, 1996a: 56. Australia (Qld); Australia (NSW, NT, SA).
decus Yeates, 1996a: 57. Australia (NSW).
elegans Yeates, 1996a: 58. Australia (WA).
eremae Yeates, 1996a: 58. Australia (NT); Australia (WA).
fenestratus Yeates, 1996a: 59. Australia (WA)
lepidus Roberts, 1929: 561. Australia (NSW); Australia (Qld).
millstreamensis Yeates, 1996a: 62. Australia (WA).
nigratus Roberts, 1929: 563. Australia (WA); australia (SA, NT).
paramonovi Hull, 1973: 300. Australia (WA).
percultus Yeates, 1996a: 66. Australia (NT).
phaleratus Yeates, 1996a: 67. Australia (WA).
principius Roberts, 1929: 562. Australia (Qld); Australia (NSW, SA, WA).
rieki Yeates, 1996a: 69. Australia (WA).
sparsus Yeates, 1996a: 71. Australia (WA).
vicinus Yeates, 1996a: 72. Australia (NSW); Australia (Qld, WA).

Subfamily LOMATIINAE
Tribe COMPTOSIINI

Genus ALEUCOSIA Edwards

ALEUCOSIA Edwards, 1934: 95. Type species: Neuria corculum Newman, 1841, orig. des.
CHELINA Hull, 1973: 306 (Comptosia subg.). Type species: Chelina norrisi Hull, 1973, orig. des.
PARAMONOVOMYIA Evenhuis, 1980: 329. Type species: Paramonovomyia halli Evenhuis, 1980, orig. des.
abdita Yeates, 1991a: 171. Australia (WA).
adusta Yeates, 1991a: 203. Australia (WA).
affinis Yeates, 1991a: 178. Australia (WA).
albovittata Yeates, 1991a: 147. Australia (WA).
angusta Edwards, 1934: 100 (Comptosia). Australia (WA).
aquila Yeates, 1991a: 180. Australia (WA).
arguta Yeates, 1991a: 165. Australia (WA).
atherix Newman, 1841: 222 (Neuria). Australia (SA); Australia (ACT, NSW, SA, Tas, Vic).
     tricellata Macquart, 1847a: 53 [1847b: 69] (Comptosia). Australia (Tas).
     geometrica Macquart, 1847a: 53 [1847b: 69] (Comptosia). Australia (Tas).
     obscura Walker, 1852: 176 (Anthrax). Australia (Tas). [Preocc. Weber, 1801.]
     tripunctata Edwards, 1934: 98 (Comptosia). Australia (Tas).
australensis Evenhuis, 1980: 332 (Paramonovomyia). Australia (WA).
     australica Evenhuis, 1989: 366 (Alyosia; n.n. for australensis Evenhuis). Australia (WA).
calophthalma Thomson, 1869: 129 (Comptosia). Australia (NSW); Australia (ACT, Qld, SA, Vic, Tas).
cincta Edwards, 1934: 99 (Comptosia). Australia (WA).
colona Yeates, 1991a: 187. Australia (SA).
corculum Newman, 1841: 221 (Neuria). Australia (WA); Australia (SA).
costalis Edwards, 1934: 99 (Comptosia). Australia (Qld); Australia (ACT, NSW).
cuneata Edwards, 1934: 100 (Comptosia). Australia"; Australia (WA).
danielsorum Yeates, 1991a: 202. Australia (NSW); Australia (ACT, Vic).
directa Yeates, 1991a: 161. Australia (NSW).
dorsalis Walker, 1849: 269 (Anthrax). "New Holland"; Australia (WA).
fulvipes Bigot, 1892b: 359 (Comptosia). "Australie"; Australia (NSW, SA, Vic, WA).
guttata Yeates, 1991a: 189. Australia (WA).
hemiteles Schiner, 1868: 132 (Neuria). Australia (NSW); Australia (Qld).
hyalina Yeates, 1991a: 154. Australia (WA).
illota Yeates, 1991a: 197. Australia (WA).
macfarlandi Evenhuis, 1980: 332 (Paramonovomyia). Australia (WA).
maculosa Newman, 1841: 221 (Neuria). Australia (WA); Australia (SA, Vic).
marginata Yeates, 1991a: 169. Australia (NSW); Australia (Qld).
multipunctata Yeates, 1991a: 185. Australia (WA).
norrisi Hull, 1973: 306. Australia (WA).
     halli Evenhuis, 1980: 329 (Paramonovomyia). Australia (WA).
obtusa Yeates, 1991a: 193. Australia (WA).
pallida Yeates, 1991a: 192. Australia (NSW); Australia (Vic).
plena Walker, 1849: 270 (Anthrax). Australia (WA).
preclara Yeates, 1991: 170. Australia (WA).
saltuaria Yeates, 1991a: 155. Australia (WA).
serpentigera Walker, 1849: 270 (Anthrax). Australia (WA).
sugillata Yeates, 1991a: 163. Australia (WA).
tridentata Yeates, 1991a: 175. Australia (WA).

Genus COMPTOSIA Macquart

COMPTOSIA Macquart, 1840: 80 [1841: 358]. Type species: Comptosia fascipennis Macquart, 1840, mon.
NEURIA Newman, 1841: 220. Type species: Neuria lateralis Newman, 1841, des. Hardy, 1921c: 54.
ALYOSIA Rondani, 1863: 54 [1864: 54]. Type species: Comptosia maculipennis Macquart, 1846 [= Neuria ocellata Newman, 1841], des. Becker, 1913a: 465.
EPIDOSIA Hull, 1973: 306 (Comptosia subg.). Type species: Comptosia vittata Edwards, 1934, orig. des.
OPSONIA Hull, 1973: 306 (Comptosia subg.). Type species: Comptosia fenestrata Hull, 1973, orig. des.
ANTHOCOLON Hull, 1973: 307 (Comptosia subg.). Type species: Comptosia casimira Hull, 1973, orig. des.
PARADOSIA Hull, 1973: 306 (Comptosia subg.). Type species: Comptosia duofasciata Hull, 1973, orig. des.
LIGYRA, authors, not Newman, 1841, misid.
EPIDOSINA, error for Epidosia.
acantha Yeates, 1991b: 1151. Australia (WA); Australia (Vic).
apicalis Macquart, 1848: 195(35). "Nouvelle-Hollande"; Australia (ACT, NSW, Qld, Vic).
aurescens Yeates, 1991b: 1087. Australia (WA).
aurifrons Macquart, 1850: 417(113). Australia, "côte orientale."
australensis Schiner, 1868: 129 (Lomatia). Australia (NSW); Australia (Qld).
bancrofti Edwards, 1934: 92. Australia (Qld).
barrowensis Yeates & Oberprieler, 2013: 351. Australia (WA).
basalis Walker, 1849: 267 (Anthrax). "New Holland" [lectotype locality]; Australia (WA).
biguttata Edwards, 1934: 93. Australia (Qld).
brunnea Edwards, 1934: 90. Australia (SA); Australia (ACT, NSW, SA, Vic).
caesariata Yeates, 1991b: 1059. Australia (Qld); Australia (ACT, NSW).
calignea Yeates, 1991b: 1123. Australia (Qld).
capillata Yeates, 1991b: 1061. Australia (Qld); Australia (NSW, Vic).
casimira Hull, 1973: 307. Not given [= Australia (WA)]; Australia (ACT).
decedens Walker, 1849: 271 (Anthrax). Australia (WA); Australia (Vic).
extensa Walker, 1835a: 473 (Anthrax). Australia (WA); Australia (ACT).
fascipennis Macquart, 1840: 81 [1841: 359]. "Monte Video," error [Australia]; Australia (ACT, NSW, Qld, Vic).
flava Yeates, 1991b: 1079. Australia (WA); Australia (NSW).
flavipenna Yeates, 1991b: 1106. Australia (NT); Australia (WA).
flexuosa Yeates, 1991b: 1096. Australia (WA); Australia (SA).
gemina Hardy, 1941: 228 (n. n. for cognata Walker). Australia (WA).
     cognata Walker, 1852: 177 (Anthrax). Australia (WA). [Preocc. Walker, 1849.]
     fenestrata Hull, 1973: 306. Not given [Australia].
heliophila Yeates, 1991b: 1075. Australia (Qld); Australia (NSW).
inclusa Walker, 1849: 268 (Anthrax). Australia (Tas); Australia (ACT, NSW, Vic).
insignis Walker, 1849: 266 (Anthrax). "New Holland"; Australia (NSW); Australia (ACT).
     ducens Walker, 1852: 176 (Anthrax). Australia (NSW).
     rufoscutellata Jaennicke, 1867: 345. "Australia".
     grandis Schiner, 1868: 132 (Neuria). Australia (NSW).
     albofasciata Thomson, 1869: 484. Australia (NSW).
     nigroscutellata, error for rufoscutellata.
insula Yeates, 1991b: 1108. Australia (Qld).
karijinii Yeates & Oberprieler, 2013: 352. Australia (WA).
kuranda
Yeates, 1991b: 1079. Australia (Qld).
lactea Yeates, 1991b: 1142. Australia (WA).
lateralis Newman, 1841: 220 (Neuria). Australia (NSW).
mackerrasi Yeates, 1991b: 1114. Australia (Qld).
magna Yeates, 1991b: 1063. Australia (WA); Australia (SA).
mallota Yeates, 1991b: 1146. Australia (Qld).
microrhynchus Yeates, 1991b: 1154. Australia (SA); Australia (WA).
moretonii Macquart, 1855: 95(75). Australia (NSW); Australia (Qld); NZ (NZ) [introduced; not established; see Evenhuis, 1986b: 7.]
murina Newman, 1841: 221 (Neuria). Australia (SA).
neoapicalis Yeates, 1991b: 1117. Australia (NSW); Australia (Qld).
neobiguttata Yeates, 1991b: 1112. Australia (NSW); Australia (ACT, Qld).
neosobria Yeates, 1991b: 1078. Australia (Qld).
nigrescens Newman, 1841: 221 (Neuria). Australia (NSW).
nitella Yeates, 1991b: 1091. Australia (WA).
ocellata Newman, 1841: 221 (Neuria). Australia (Tas).
     maculipennis Macquart, 1846: 244(116). Australia (Tas).
paramonovi Yeates, 1991b: 1089. Australia (SA); Australia (Vic).
partita Newman, 1841: 221 (Neuria). Australia (WA).
paucispina Yeates, 1991b: 1042. Australia WA).
pilosa Yeates, 1991b: 1056. Australia (WA); Australia (NSW, SA, Vic).
pitereka Yeates, 1991b: 1048. Australia (SA).
praeargentata Macleay, 1826: 468 (Anthrax). Not given [Australia]; Australia (ACT, NSW, Qld).
     edwardsi Hardy, 1941: 227. Australia (Qld).
quadripennis Walker, 1849: 268 (Anthrax). "Australia"; Australia (ACT, NSW, Qld, SA, Vic).
rubrifera Bigot, 1881c: 23 (Lygira). "Australia"; Australia (NSW, Qld).
sandaraca Yeates, 1991b: 1081. Australia (WA).
scitula Yeates, 1991b: 1148. Australia (Qld).
sobria Walker, 1849: 269 (Anthrax). "Australia"; Australia (NSW); PNG (PNG).
     bicolor Macquart, 1850: 418 (114). Australia, "côte orientale."
subsenex Walker, 1857: 144 (Anthrax). "Australia."
sobricula Walker, 1857: 144 (Anthrax). "Australia"; Australia (ACT, NSW, Qld, Tas, Vic).
     simplex Macquart, 1847a: 52 [1847b: 68] (Anthrax). Australia (Tas). [Preocc. Wiedemann, 1819.]
     anthracina Thomson, 1869: 485. Australia (NSW).
     prosimplex Hardy, 1942: 203 (Villa; n. n. for simplex Macquart). Australia (Tas).
     sorbicula, error for sobricula.
soror Yeates, 1991b: 1115. Australia (Qld).
speciosa Yeates, 1991b: 1066. Australia (WA).
stria Walker, 1849: 267 (Anthrax). "Australia"; Australia (NSW, Tas).
     fasciata Fabricius, 1805: 118 (Anthrax). "Insulis oceani pacifica" [= Australia (Tas); see Evenhuis 1988]. [Preocc. Meigen, 1804.]
tendens Walker, 1849: 271 (Anthrax). Australia (WA).
duofasciata Hull, 1973: 306. Not given [= Australia (WA)].
thyris Yeates, 1991b: 1121. Australia (WA).
tutela Yeates, 1991b: 1045. Australia (Qld); Australia (NSW).
vittata Edwards, 1934: 89. Australia (Vic); Australia (NSW, SA).
walkeri Edwards, 1934: 94. Not given [Australia]; Australia (NSW, Qld).
wilkinsi Edwards, 1934: 94. Australia (NT).
xanthobasis Yeates, 1991b: 1084. Australia (WA).
zona Yeates, 1991b: 1119. Australia (WA).

Genus DODDOSIA Edwards

DODDOSIA Edwards, 1934: 103. Type species: Doddosia picta Edwards, 1934, orig. des.
collessi Yeates, 1990: 78. Australia (NT).
lurida Yeates, 1990: 77. Austrlia (Qld); Australia (SA).
picta Edwards, 1934: 103. Australia (Qld); Australia (NT).
procax Yeates, 1990: 78. Australia (qld).
torresiana Yeates, 1990: 79. Australia (Qld).

Genus ONCODOSIA Edwards

ONCODOSIA Edwards, 1934: 101. Type species: Anthrax patula Walker, 1849, orig. des.
ampla Walker, 1852: 185 (Anthrax). Australia (WA).
patula Walker, 1849: 273 (Anthrax). Australia (WA); Australia (SA, Vic).
plana Walker, 1849: 272 (Anthrax). Australia (WA).
triangularis Yeates, 1988: 516. Australia (Vic); Australia (SA, WA).

Subfamily TOMOMYZINAE

Genus DOCIDOMYIA White

DOCIDOMYIA White, 1916b: 200, 203. Type species: Docidomyia puellaris White, 1916, mon.
PARACOSMUS, Evenhuis, 1989, not Osten Sacken, 1877, misid.
albifrons Yeates, 1996b: 410. Australia (NT); Australia (Qld, WA).
angusta Yeates, 1996b: 413. Australia (WA).
brevipilata Yeates, 1996b: 414. Australia (Qld); Australia (NSW).
danielsi Yeates, 1996b: 415. Australia (NSW); Australia (NT, WA).
froggatti Paramonov, 1951a: 748. Australia (NSW).
genualia Yeates, 1996b: 417. Australia (WA); Australia (NT).
hespera Yeates, 1996b: 420. Australia (WA).
isla Yeates, 1996b: 421. Australia (Qld).
nicholsoni Yeates, 1996b: 422. Australia (WA).
nigra Yeates, 1996b: 423. Australia (WA).
norrisi Paramonov, 1951a: 748. Australia (WA).
parva Yeates, 1996b: 425. Australia (Qld); Australia (NSW).
puellaris White, 1916b: 204. Australia (Tas) & (Vic); Australia (ACT, NSW, Qld, SA).
swani Paramonov, 1951a: 750. Australia (SA); Australia (NSW).
victoriana Paramonov, 1951a: 749. Australia (Vic); Australia (NSW).

Subfamily ANTONIINAE

Genus ANTONIAUSTRALIA Becker

ANTONIAUSTRALIA Becker, 1913a: 458. Type species: Antoniaustralia hermanni Becker, 1913, orig. des.
arida Evenhuis, 1993: 3. Australia (NT).
hermanni Becker, 1913a: 458. "Australien" [= Australia (NT)].

Genus CYX Evenhuis

CYX Evenhuis, 1993: 7. Type species: Antonia rieki Paramonov, 1953, orig. des.
decorata Paramonov, 1950d: 532 (Antonia). Not given [Australia]; Australia (Qld).
papuana Evenhuis, 1993: 10. PNG (PNG).
rieki Paramonov, 1953c: 204 (Antonia). Australia (NSW).
roddi Paramonov, 1950d: 530 (Antonia). Australia (NSW).

Genus MYONEMA Roberts

MYONEMA Roberts, 1929: 563. Type species: Myonema humile Roberts, 1929, orig. des.
humile Roberts, 1929: 564. Australia (Qld).

Subfamily ANTHRACINAE
Tribe ANTHRACINI

Genus ANTHRAX Scopoli

ANTHRAX Scopoli, 1763: 358. Type species: Musca morio Linnaeus, 1758 [misid., = Musca anthrax Schrank, 1781], mon.
arcus de Meijere, 1913b: 322 (Argyramoeba). Indonesia (Irian Jaya).
argentius Yeates & Lambkin, 1998: 991 (as argentia). Australia (WA); Australia (SA).
angularis Thomson, 1869: 482. Australia (NSW); Australia (NT, Qld, SA, WA).
asciculus Yeates & Lambkin, 1998: 1010. Australia (Qld); Australia (NSW, SA, Vic).
clinatus Yeates & Lambkin, 1998: 1013. Australia (WA); Australia (NSW, NT, Qld).
coeruleopennis Doleschall, 1857: 400. Indonesia (Maluku); ?PNG (PNG).
     caeruleopennis, error for coeruleopennis.
confluensis Roberts, 1928: 139. Australia (Qld); Australia (ACT, NSW, Nt, SA, Vic).
crenatus Yeates & Lambkin, 1998: 1018. Australia (NT); Australia (ACT, Qld, SA, Vic, WA).
distigma Wiedemann, 1828: 309. Indonesia (Java); Indonesia (Maluku), PNG (PNG).
doleschalli Evenhuis & Greathead, 1999: 299 (n.n. for argyropyga Doleschall, 1857). Indonesia (Maluku).
     argyropyga Doleschall, 1857: 401. Indonesia (Maluku). [Preocc. Wiedemann, 1828.]
dolobratus Yeates & Lambkin, 1998: 1022. Australia (NSW); Australia (ACT, Qld, Vic).
emissa Walker, 1864: 233. Indonesia (Maluku); PNG (PNG).
emittens Walker, 1860b: 148. Indonesia (Maluku).
incisus Macquart, 1847a: 52 [1847b: 68]. Australia (Qld).
incomptus Walker, 1849: 253. Australia (WA); Australia (ACT, NSW, NT, Qld, SA, Vic), ?PNG (Bismarck Arch), Vanuatu; Christmas I (Indian Ocean).
     semimacula Walker, 1849: 254. "New Holland."
koshunensis Matsumura, 1916: 282. Taiwan; Bonin Is, Guam, Hawaiian Is [introduced]; Japan.
     distigma, authors, not Wiedemann, misid.
lepidiotus Roberts, 1928a: 138. Australia (Qld); Australia (NT).
maculatus Macquart, 1846: 240(112). Australia (Tas); Australia (ACT, NSW, NT, Qld, SA, Vic, WA).
     diana Walker, 1849: 252. Not given [= Australia (WA)].
     australis Walker, 1852: 193. "Van Diemen's Land" [= Australia (Tas)] & Australia (WA).
matilei Evenhuis, 1991: 281. New Caledonia (New Caledonia).
melanius Wulp, 1885c: 84. Indonesia (Java); PNG (PNG).
opacus Yeates & Lambkin, 1998: 1018. Australia (Qld); Australia (NSW, SA).
proconcisus Hardy, 1942: 203 (Villa; n. n. for concisa Macquart, 1850). Australia, "côte orientale" [= eastern Australia]; Australia (ACT, NSW, Qld, SA, Vic).
     concisa Macquart, 1850: 415(111). Australia "côte orientale" [= eastern Australia] [Preocc. Macquart, 1840.]
semiscitus Walker, 1857: 118. Malaysia (Sarawak); ?French Polynesia (Society Is), Indonesia (Irian Jaya), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia (Sulawesi).
     tripunctata Wulp, 1869: 109. Indonesia (Irian Jaya). [Preocc. Wiedemann, 1820.]
siccus Yeates & Lambkin in Evenhuis & Greathead, 1999: 314 (n.n. for funestus Yeates & Lambkin, 1998). Australia (WA); Australia (Vic).
     funestus Yeates & Lambkin, 1998: 1024. Australia (WA). [Preocc. Walker, 1849.]
succedens Walker, 1852: 192. Not given [unknown].
torulus Yeates & Lambkin, 1998: 1001. Australia (Qld); Australia (NT, WA).

Genus BRACHYANAX Evenhuis

BRACHYANAX Evenhuis, 1981: 190. Type species: Anthrax satellita Walker, 1857 [= Brachyanax thelestrephones Evenhuis, 1981], orig. des.
HEMIPENTHES, authors, not Loew, 1869, misid.
ater Roberts, 1928a: 140 (Anthrax). Australia (Qld); Indonesia (Nusa Tenggara).
     australianus Evenhuis, 1981: 197. Australia (Qld).
aterrimus Doleschall, 1858: 93 (Anthrax). Indonesia (Maluku); PNG (Bismarck Arch, PNG), Solomon Is; Indonesia (Sulawesi), Japan, Philippines.
     proferens Walker, 1859c: 113 (Anthrax). Indonesia (Sulawesi).
     oceanicus Evenhuis, 1981: 197. Solomon Is.
chichijimensis Evenhuis, 1981: 195. Bonin Is.
leucostigma Wulp, 1898a: 419 (Anthrax; n. n. for terminalis Wulp). Indonesia (Maluku); PNG (PNG).
     terminalis Wulp, 1869: 108 (Anthrax). Indonesia (Maluku). [Preocc. Wiedemann, 1830.]
magnipennis Evenhuis, 1981: 195. Solomon Is.
papuanus Evenhuis, 1981: 198. PNG (PNG).
satellitius Walker, 1857: 119 (Anthrax). Malaysia (Sarawak); Indonesia (Maluku); Indonesia (Kalimantan), Malaysia (Pen, Sabah).
     satellita, error for satellitius.

Genus THRAXAN Yeates & Lambkin

THRAXAN Yeates & Lambkin, 1998: 1027. Type species: Thraxan simulans Yeates & Lambkin, 1998, orig. des.

abditus Yeates & Lambkin, 1998: 1030. Australia (Qld); Australia (NSW).
acutus Yeates & Lambkin, 1998: 1031. Australia (Qld); Australia (NSW).
caligneus Yeates & Lambkin, 1998: 1034. Australia (Qld).
cinctus Yeates & Lambkin, 1998: 1037. Australia (Qld); Australia (NSW, NT, WA).
cornuatus Yeates & Lambkin, 1998: 1039. Australia (WA); Australia (NT, Qld).
depressus Yeates & Lambkin, 1998: 1041. Australia (NSW); Australia (NT, Qld).
ebeneus Yeates & Lambkin, 1998: 1042. Australia (ACT); Australia (Qld).
echinatus Yeates & Lambkin, 1998: 1030. Australia (Qld); Australia (NT, WA).
emicatus Yeates & Lambkin, 1998: 1045. Australia (NT); Australia (WA).
hamulus Yeates & Lambkin, 1998: 1046. Australia (Qld).
luteus Yeates & Lambkin, 1998: 1047. Australia (NT); Australia (Qld, WA).
misatulus Yeates & Lambkin, 1998: 1049. Australia (WA); Australia (NSW, NT, Qld, Vic).
nodus Yeates & Lambkin, 1998: 1051. Australia (Qld).
norrisi Yeates & Lambkin, 1998: 1052. Australia (WA).
obstipus Yeates & Lambkin, 1998: 1054. Australia (NT); Australia (NSW, Qld).
patielus Yeates & Lambkin, 1998: 1055. Australia (NSW); Australia (ACT, Qld).
planus Yeates & Lambkin, 1998: 1056. Australia (Qld); Australia (NT, WA).
prolatus Yeates & Lambkin, 1998: 1057. Australia (Qld); Australia (NSW).
simulatus Yeates & Lambkin, 1998: 1028. Australia (Qld).
spiculus Yeates & Lambkin, 1998: 1059. Australia (Qld).

Unplaced species of Anthracini

boninensis Matsumura, 1916: 284 (Agryramoeba). Bonin Is.

Tribe EXOPROSOPINI

Genus ATRICHOCHIRA Hesse

ATRICHOCHIRA Hesse, 1956; 911. type species: Exoprosopa pediformis Bezzi, 1921, orig. des.
commoni Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 855. Australia (Qld).
paramonovi Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 857. Australia (NSW); Australia (Qld).

Genus BALAANA Lambkin & Yeates

BALAANA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 757. Type species: Exoprosopa latelimbata Bigot, 1892, orig. des.
abscondita Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 761. Australia (NW); Australia (ACT, NT, Qld, SA, Vic, WA).
biscuspis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 764. Australia (NT); Australia (Qld, WA).
centrosa Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 766. Australia (WA); Australia (NT, SA, Vic, WA).
gigantea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 768. Australia (Qld); Australia (NSW, Vic, WA).
kingcascadensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 770. Australia (WA); Australia (NT).
latelimbata Bigot, 1892b: 346 (Exoprosopa). "Australie"; Australia (ACT, NSW, Qld, SA).
obliquebifasciata Macquart, 1850: 411(107) (Exoprosopa). Australia (Tas).
     obliquefasciata, error for obliquebifasciata.



Genus EXOPROSOPA Macquart

EXOPROSOPA Macquart, 1840: 35 [1841: 313]. Type species: Anthrax pandora Fabricius, 1805, des. Coquillett (1910: 544) [Designation conserved by I.C.Z.N. 1991: 82.].
bicellata Macquart, 1847a: 51 [1847b: 61]. Australia (Tas).
sylvana Fabricius, 1775: 758 (Bibio). "nova Hollandia" [= Australia (Qld); see Musgrave, 1932: 54].


Genus KAPU Lambkin & Yeates

KAPUA Lambkin & Yeates, 2003: 776. Type species: Exoprosopa adelaidica Macquart, 1855, orig. des. [Preocc. Marwick, 1931.]
KAPU Lambkin & Yeates, 2006: 161 (n.n. for Kapua Lambkin & Yeates).
adelaidica Macquart, 1855: 90(70) (Exoprosopa). Australia (SA); Australia (NSW, NT, Qld, Tas, WA).
corusca Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 780. Australia (NSW); Australia (ACT, NT, Qld, WA).
irwini Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 783. Australia (Qld); Australia (NSW, SA).
westralica Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 785. Australia (WA).


Genus LARRPANA Lambkin & Yeates

LARRPANA Lambkin & Yeates, 2003: 787. Type species: Exoprosopa dimidiatipennis Bowden, 1971, orig. des.
bushblitz Lambkin & Bartlett, 2011: 248. Australia (Qld) .
collessiLambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 783. Australia (NT); Australia (Qld).
dimidiatipennis
Bowden, 1971b: 64 (Exoprosopa; n. n. for dimidiata Roberts). Australia (NT).
     dimidiata Roberts, 1928a: 109 (Exoprosopa). Australia (NT). [Preocc. Macquart, 1846.]
zwicki Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 793. Australia (Qld).


Genus LIGYRA Newman

LIGYRA Newman, 1841: 220. Type species: Anthrax bombyliformis Macleay, 1826, mon.
LYGIRA, error for Ligyra.
HYPERALONIA, authors, not Rondani, 1863, misid.
alea Bowden, 1971d: 321. PNG (Bismarck Arch).
audouinii Macquart, 1840: 36 [1841: 314] (Exoprosopa). "Indes orientales"; Indonesia (Irian Jaya, Maluku); China, Indonesia (Java, Sulawesi), Philippines, Taiwan.
     albicincta Macquart, 1840: 48 [1841: 316] (Exoprosopa). China.
     formosana Paramonov, 1931: 195 (Hyperalonia). Taiwan.
     macassarensis Paramonov, 1931: 198 (Hyperalonia). Indonesia (Sulawesi).
bombyliformis Macleay, 1826: 468 (Anthrax). Australia (NSW); Australia (NT, Qld, WA).
     albiventris Thomson, 1869: 480 (Exoprosopa). Australia (NSW).
     bombiliformis, bombyliiformis, errors for bombyliformis.
burnsi Paramonov, 1967: 143. Australia (NSW).
calabyana Paramonov, 1967: 136. Australia (WA); Australia (NT).
campbelli Paramonov, 1967: 139. Australia (NT).
chinnicki Paramonov, 1967: 138. Australia (SA); Australia (NSW, Vic).
cingulata Wulp, 1885b: 62 (Exoprosopa). Australia (SA); Australia (NSW, NT, Qld, WA).
     incompleta Paramonov, 1967: 141 (cingulata var.). [Unavailable name; varietal name proposed after 1960.]
commonsi Paramonov, 1967: 134. Australia (Qld).
cruciata Bowden, 1971a: 11. PNG (PNG).
dentata Roberts, 1928a: 102 (Hyperalonia). Australia (Vic).
devecta Walker, 1860b: 148 (Anthrax). Indonesia (Maluku).
     delecta, error for devecta.
doryca Boisduval, 1835: 665 (Anthrax). Indonesia (Maluku); Australia (NSW, Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia (Java, Timor).
     pelops Walker, 1858b: 90 (Anthrax). Indonesia (Maluku).
eyreana Paramonov, 1967: 136. Australia (SA).
fasciata Paramonov, 1967: 135. Australia (Qld).
flaviventris Doleschall, 1857: 400 (Anthrax). Indonesia (Maluku).
flavotomentosa de Meijere, 1929a: 386 [1929b: 416] (Hyperalonia). Indonesia (Maluku).
fuscipennis Macquart, 1848: 193(33) (Exoprosopa). Indonesia (Java); "Australia," PNG (PNG); "Borneo," Cambodia, China, ?India, Indonesia (Java, Sulawesi, Sumatra), Malaysia (Pen), ?Nepal.
     confirmata Walker, 1861d: 283 (Anthrax). Indonesia (Maluku).
     chrysolampis Jaennicke, 1867: 344 (Hyperalonia). Indonesia (Java).
     tantalus, authors, not Fabricius, 1794, misid.
hemifusca Roberts, 1928a: 97 (Hyperalonia). Australia (Qld); Australia (NT, WA).
inquinita Roberts, 1928a: 103 (Hyperalonia). Australia (Qld); Australia (NSW).
leptyna Bowden, 1971a: 12. PNG (Bougainville I).
leuconoe Jaennicke, 1867: 345 (Exoprosopa). Indonesia (Maluku).
nigrocostalis Guérin-Méneville, 1838: 295 (Anthrax). Indonesia (Maluku)
ochracea Bowden, 1971a: 10. PNG (PNG).
orest Paramonov, 1967: 134. Australia (Qld); Australia (NT).
pilad Paramonov, 1967: 143. Australia (WA).
punctipennis Macquart, 1850: 410(106) (Exoprosopa). Australia, "côte orientale"; widesp. Australia.
     contrasta Paramonov, 1967: 133. Australia (NSW).
robertsi Paramonov, 1967: 132. Australia (WA); Australia (NSW, NT, Qld).
satyrus Fabricius, 1775: 758 (Bibio). "nova Hollandia" [= Australia (Qld); see Musgrave, 1932: 54]; widesp. Australia, Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG); ?China, Indonesia (Timor).
     funesta Walker, 1849: 242 (Anthrax). Australia (NT).
     insignis Macquart, 1855: 93(73) (Exoprosopa). Australia (Qld).
septentrionis Roberts, 1928a: 103 (Hyperalonia). Australia (Qld); Australia (NT, WA).
sinuatifascia Macquart, 1855: 92(72) (Exoprosopa). Australia (NSW); Australia (ACT, SA, Vic, WA).
     macraspis Thomson, 1869: 479 (Exoprosopa). Australia (NSW).
     argenticincta Bigot, 1892b: 343 (Hyperalonia). Australia (NSW).
tantalus Fabricius--not Australasian/Oceanian.
tenebrosa Paramonov, 1967: 132. Australia (Qld).
unicincta Guérin-Méneville, 1831: pl. 20, fig. 9 [1838: 294] (Anthrax). Indonesia (Maluku).
ventrimacula Doleschall, 1857: 399 (Anthrax). Indonesia (Maluku).

Genus LITORHINA Bowden

LITOMYZA Hull, 1973: 426. Nomen nudum.
LITORHINA Bowden, 1975: 314. Type species: Litorhynchus corticeus Bezzi, 1924, orig. des.
LITORHYNCHUS, authors, not Macquart, 1840, misid.
collaris Wiedemann--not Australasian/Oceanian.
lar Fabricius, 1782: 414 (Bibio). India; ?Australia (WA); Sri Lanka.


Genus MUNJUA Lambkin & Yeates

MUNJUA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 795. Type species: Munjua erugata Yeates & Lambkin, 2003, orig. des.
erugata Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 795. Australia (WA); Australia (NSW, NT, SA, Vic).
lepidokingi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 799. Australia (WA).
paralutea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 801. Australia (NSW); Australia (NT, Qld, SA, WA).


Genus MUWARNA Lambkin & Yeates

MUWARNA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 807. Type species: Anthrax stellifer Walker, 1849, orig. des.
stellifera Walker, 1849: 244 (Anthrax). "New Holland" & Australia (WA); Australia (SA, Vic, WA).
vitreilinearis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 810. Australia (Vic); Australia (WA).


Genus NGALKI Lambkin

NGALKI Lambkin in Lambkin & Bartlett, 2011: 254. Type species: Munjua trigona Lambkin & Yeates, 2003, orig. des.
trigonum Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 804 (Munjua). Australia (Vic); Australia (NSW, SA, WA).

Genus PALRIKA Yeates & Lambkin

PALRIKA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 811. Type species: Palrika decora Lambkin & Yeates, 2003, orig. des.
anaxios Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 814. Australia (Qld).
basilikos Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 817. Australia (WA).
blackdownensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 820. Australia (Qld).
bouchardi Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 821. Australia (Qld); Australia (NSW, NT, WA).
culgoafloodplainensis Lambkin & Bartlett, 2011: 242. Australia (Qld) .
cyanea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 823. Australia (Qld); Australia (NT, WA).
danielsi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 824. Australia (NSW); Australia (Qld).
decora Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 812. Australia (Qld); Australia (ACT).
mackenziei Lambkin & Bartlett, 2011: 245. Australia (Qld) .
marginicollis Gray in Griffith & Pidgeon, 1832: 780 (Anthrax). Not given [Australia]; Australia (NSW, NT, Qld, WA).
viridula Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 828. Australia (Qld); Australia (NSW).
whyalla Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 831. Australia (SA); Australia (WA).

Genus PSEUDOPENTHES Roberts

PSEUDOPENTHES Roberts, 1928a: 132. Type species: Pseudopenthes fenestrata Roberts, 1928, orig. des.
fenestrata Roberts, 1928a: 133. Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
hesperis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 863. Australia (WA).

Genus PTEROBATES Bezzi

PTEROBATES Bezzi, 1921b: 130 (Exoprosopa subg.). Type species: Anthrax apicalis Wiedemann, 1821, mon.
pennipes Wiedemann, 1821b: 129 (Anthrax). Indonesia (Java); Indonesia (Maluku); China, Hong Kong, India, Macau, Malaysia (Sabah), Philippines.

Genus WURDA Lambkin & Yeates

WURDA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 833. Type species: Wurda emu Lamkin & Yeates, 2003, orig. des.
emu Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 833. Australia (NSW); Australia (NT, Qld, SA, WA).
impatientis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 838. Australia (WA); Australia (SA).
montebelloensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 840. Australia (WA); Australia (NT, Qld, SA).
norrisi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 843. Australia (WA); Australia (SA, Vic).
patrellia Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 845. Australia (NSW).
skevingtoni Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 847. Australia (WA).
windorah Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 848. Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
wyperfeldensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 851. Australia (Vic); Australia (NSW, Qld, SA, WA).

Unplaced species of Exoprosopini

albirufus Walker, 1857: 143 (Anthrax). "Australia."
alternans Macquart, 1850: 414(110) (Anthrax). Australia (Tas).
alternus Walker, 1849: 261 (Anthrax). "New Holland"; Australia (Tas).

 

Tribe VILLINI
Genus CHRYSANTHRAX Osten Sacken

CHRYSANTHRAX Osten Sacken, 1886: 121 (as Anthrax "group"). Type species: Anthrax fulvohirta Wiedemann, 1821 [= Anthrax cypris Meigen, 1820], des. Coquillett, 1910: 523.
cypris Meigen, 1820: 158 (Anthrax). North America [see Meigen, 1830: 325]; Hawaiian Is [introduced into quarantine; not established]; widesp. Nearctic Reg.
     fulvohirta Wiedemann, 1821a: 99 [1821b: 149] (Anthrax). USA.
Undetermined sp.—Marshall Is.

Genus EXECHOHYPOPION Evenhuis

EXECHOHYPOPION Evenhuis, 1991: 283. Type species: Exechohypopion greatheadi Evenhuis, 1991, orig. des.
albatum Roberts, 1928a: 124 (Villa). Australia (WA).
bruneum Roberts, 1928a: 118 (Villa). Australia (NSW).
greatheadi Evenhuis, 1991: 286. New Caledonia (New Caledonia).
minus Macquart, 1850: 415(111) (Anthrax). Australia (Tas); ?New Caledonia (New Caledonia).
nigricostatum Macquart, 1850: 415(111) (Anthrax). Australia (Tas); Australia (NSW, SA, Vic).
     nigricosta, error for nigricostata.
subobscurum Hardy, 1942: 203 (Anthrax; n. n. for obscura Macquart). Australia (Tas).
     obscura Macquart, 1846: 240(112) (Anthrax). Australia (Tas). [Preocc. Weber, 1801.]
therephaga Yeates in Irwin & Yeates, 1995: 1319. Australia (NSW).
velox White, 1916b: 211 (Anthrax). Australia (Tas).

Genus EXHYALANTHRAX Becker

EXHYALANTHRAX Becker, 1916a: 44 (Villa subg.). Type species: Anthrax vagans Loew, 1862, des. Bezzi, 1924a: 7.
afer Fabricius--not Australasian/Oceanian.
ogasawarensis Matsumura, 1916: 281 (Anthrax). Bonin Is.

Genus LEPIDANTHRAX Osten Sacken

LEPIDANTHRAX Osten Sacken, 1886: 107. Type species: Anthrax disjunctus Wiedemann, 1830, des. Coquillett, 1910: 559.
albifrons Roberts, 1928a: 131. Australia (SA).
linguatus Roberts, 1928a: 130. Australia (Qld).

Genus VILLA Lioy

VILLA Lioy, 1864a: 732. Type species: Anthrax concinnus Meigen, 1820 [= Anthrax abbadon Fabricius, 1794], des. Coquillett, 1910: 619.
ANTHRAX, authors, not Scopoli, 1763, misid.
abeilla Hull, 1973: 374 (n. n. for antecedens Walker). Indonesia (Sulawesi); Indonesia (Maluku); Philippines.
     antecedens Walker, 1859c: 111 (Anthrax). Indonesia (Sulawesi). [Preocc. Walker, 1852.]
apicifera Walker, 1865b: 111 (Anthrax). "New Guinea" [= Indonesia (Irian Jaya)].
argentipennis White, 1916b: 212 (Anthrax). Australia (Tas).
commista Macquart, 1850: 413(109) (Anthrax). Australia, "côte orientale" [prob. = Australia (NSW)].
consimilis Thomson, 1869: 481 (Anthrax). Australia (NSW).
flaveola Macquart, 1850: 413(109) (Anthrax). Australia, "côte orientale" [prob. = Australia (NSW)].
fumea Roberts, 1928a: 127. Australia (NSW).
fuscicostata Macquart, 1846: 239(111) (Anthrax). Australia (Tas); Australia (NSW, Vic).
     marginata Walker, 1852: 178 (Anthrax). Australia (Tas).
lipposa Bigot, 1892b: 353 (Glossista). Australia (NSW).
macquarti Evenhuis, 1989: 373 (n. n. for basalis Macquart). Australia (NSW); ?New Caledonia (New Caledonia).
     basalis Macquart, 1855: 95(75) (Anthrax). Australia (NSW). [Preocc. Walker, 1849.]
pellucida Walker, 1852: 182 (Anthrax). Australia (Tas).
pretendens Walker, 1859c: 111 (Anthrax). Indonesia (Sulawesi); Indonesia (Irian Jaya, Maluku), PNG (PNG).
     praetendens, error for pretendens.
quinqueguttata Roberts, 1928a: 126. Australia (NSW).
rava Roberts, 1928a: 125. Australia (Qld).
resurgens Walker, 1849: 259 (Anthrax). Australia (WA).
subannula Walker, 1849: 261 (Anthrax). Not given [unknown].
trivincula Roberts, 1928a: 119. Australia (Qld); Australia (NSW), Lord Howe I.
troglodyta Fabricius, 1794: 759 (Bibio). Indonesia (Maluku); PNG (PNG).
vitrea Walker, 1852: 181 (Anthrax). Australia (WA).
zonipennis Evenhuis, 1978b: 247 (n. n. for varipennis Roberts). Australia (NSW).
     varipennis Roberts, 1928a: 121. Australia (NSW). [Preocc. Macquart, 1850.]

Nomen nudum in Villa Lioy

tasmanica Hardy, 1924a: 76 (Anthrax).

Tribe XERAMOEBINI

Genus DESMATONEURA Williston

DESMATONEURA Williston, 1895: 267. Type species: Desmatoneura argentifrons Williston, 1895, mon.
Undetermined sp.--Australia (WA).

Genus PETROROSSIA Bezzi

PETROROSSIA Bezzi, 1908a: 35. Type species: Bibio hesperus Rossi, 1790, orig. des.
PETROSSIA, error for Petrorossia.
angustata Doleschall, 1858: 93 (Anthrax). Indonesia (Maluku); Indonesia (Sulawesi).
fulvula Wiedemann, 1821b: 148 (Anthrax). Indonesia (Java); ?Australia, Indonesia (Maluku), ?PNG (PNG); Indonesia (Kalimantan, Sulawesi), Malaysia (Pen, Sabah, Sarawak), Singapore.
     degenera Walker, 1856a: 15 (Anthrax). Singapore.
     fulunla, error for fulvula.
oceanica Bowden, 1971d: 313. PNG (Bismarck Arch).


This page last revised 14 April 2016