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Members of the Bombyliidae, commonly called bee flies, are
most frequently found in arid and semiarid portions of the world,
with a few specialized forms occurring in moist tropical climates.
Adults range in size from 1 to 21 mm in length and are commonly
found on flowers, hovering in bright sunlight, or resting on warm,
bare ground. Many, if not all, are pollen and nectar feeders,
with some acting as primary pollinators of certain species of
desert wildflowers.
Species found in Australia include some of the most striking species
known; many have large, pictured wings and brightly colored cuticular
pigmentation or vestiture. There is a great deal of endemism exhibited
in the taxa from Australia and the Oceanic islands (Evenhuis 1982b).
The Oriental Region seems to be a source for many of the Oceanic
taxa, but some of the Australian fauna (viz., Comptosiini and
Dischistini) show possible Transantarctic relationships because
related genera of the 2 tribes are also found in central and southern
Chile. There are over 400 species known from the areas covered
in this catalog. As the fauna from these areas, especially the
Australian, becomes better known, this number could easily double
or even triple.
Immatures are infrequently encountered. Larvae of some genera
are predators of orthopteroid egg pods, most others are parasitoids
on immature stages of holometabolous orders of insects (DuMerle
1975) or are free-living in soil. Ectoparasitic representatives
from the regions covered here include Exhyalanthrax on
pupiparous higher Diptera, Systropus on saddleback moths
(Limacodidae), Geron on Psychidae, and the ubiquitous Anthrax
on a variety of hosts, primarily aculeate Hymenoptera. Hyperparasites
occurring in these areas include Exoprosopa and Ligyra
on Tiphiidae and Scoliidae (Hymenoptera), which are parasites
of grubs of Scarabaeoidea. Hardy (1921c) recorded Ligyra satyrus
as hyperparasitic on an unidentified asilid.
Classification of the family follows, for the most part, that
provided by Yeates (1994) with some exception, which follow that
of Evenhuis (1991). The Mythicomyiinae of Yeates has been raised
to family
level here following Zaitzev (1991) and Evenhuis (1993, 1994).
The subfamily Bombyliinae is represented in these regions only
by the tribe Dischistini. Species previously described in the
genus Bombylius should be transferred to other genera after
types have been examined. Due to the poor level of taxonomic knowledge
of these flies, previous synonymies for species in many genera
have been included here only when substantiated by examination
of type material.
The only comprehensive work on the bombyliid fauna from these
regions is the revisionary work of Roberts (1928a,b, 1929). The
late S.J. Paramonov contributed many papers on the Australian
Bombyliidae, and many of his unfinished manuscripts on various
genera are deposited in CSIRO, Canberra. David Yeates has begun
a series of studies at the subfamilial and generic level of the
Australian Bombyliidae.Chris Lambkin et al.(2003) revised the Australian Exoprosopini.
Refs.: Roberts (1928a,b, 1929, keys, rev. Australian spp.),
Edwards (1934, rev. Lomatiinae), Paramonov (1967. rev. Australian
Ligyra), Hull (1973, world gen.), DuMerle (1975, immat.
biol., bibliogr.), Evenhuis (1979b, rev. Australasian/Oceanian
Geron; 1982, Oceanian zoogeogr.; 1983b, bibliogr.; 1988a,
rev. Brachyanax; 1988b, phylog. Phthiriinae; 1991, classif.,
world gen.; 1994, cat. fossil Bombyliidae), Greathead (1988, phylog.
Ecliminae), Yeates (1988, rev. Oncodosia; 1994, fam. classif.),
Yeates & Hagen (1988; rev. Australian Systropus); Yeates
& Lambkin (1998, rev. Australian Anthracini); Evenhuis & Greathead (1999, world cat.); Lambkin et al. (2003, rev. Australian Exoprosopini).
SYSTROPUS Wiedemann, 1820a: 18. Type species: Systropus
macilentus Wiedemann, 1820, orig. des. (on plate).
SYSTROPUS Henning, 1832: 335. Type species: Systropus
macilentus Henning, 1832 [preocc., = Systropus macilentus
Wiedemann, 1820], mon. [Preocc. Wiedemann, 1820.]
CEPHENE Latreille, 1825: 496. Vernacular name without nomenclatural
status.
CEPHENUS Berthold, 1827: 506 (unnec. n. n. for Systropus).
XYSTROPHUS Agassiz, 1846b: 393 (unnec. emend. of Systropus).
CEPHENIUS Enderlein, 1926b: 70 (as gen.). Type species:
Systropus studyi Enderlein, 1926, orig. des.
SYSTROPHUS, error for Systropus.
SYSTROPHOPUS, error for Systropus.
doddi Roberts, 1929: 555. Australia (Qld).
flavoornatus Roberts, 1929: 554. Australia (Qld).
tipuloides Westwood, 1876c: 576. Indonesia (Maluku).
ZACLAVA Hull, 1973: 250. Type species: Systropus
clavifemoratus Hardy, 1921, orig. des.
clavifemorata Hardy, 1921c: 61 (Systropus). Australia
(NSW); Australia (Qld).
kraussi Evenhuis, 1979a: 87. New Caledonia (Loyalty Is);
New Caledonia (New Caledonia).
minima Roberts, 1929: 557 (Dolichomyia). Australia
(Qld); Australia (NSW).
occidentis Roberts, 1929: 557 (Dolichomyia). Australia
(WA).
TOXOPHORA Meigen, 1803: 270. Type species: Toxophora
maculata Meigen, 1804 [preocc., = ?Asilus fasciculatus
Villers, 1789], subs. mon. Meigen, 1804: 273.
ENICONEVRA Macquart, 1840: 110 [1841: 388]. Type species:
Eniconevra fuscipennis Macquart, 1840, mon.
ENICONEURA Agassiz, 1846a: 13 (unjust. emend. of Eniconevra).
HENICONEVRA Agassiz, 1846b: 138 (unjust. emend. of Eniconevra).
HENICONEURA Scudder, 1882: 159 (unjust. emend. of Eniconevra).
compta Roberts, 1929: 559. Australia (Qld).
GERON Meigen, 1820: 223. Type species: Geron gibbosus
Meigen, 1820 [= Bombylius gibbosus Olivier, 1789], des.
Duponchel in d'Orbigny, 1845b: 206.
GERON Meigen, 1820: 223 (as gen.). Type species: Geron
gibbosus Meigen, 1820 [= Bombylius gibbosus Olivier,
1789], des. Duponchel in d'Orbigny, 1845b: 206.
aaptis Evenhuis, 1979b: 19. New Caledonia (Loyalty Is).
aesion Evenhuis, 1979b: 20. New Caledonia (New Caledonia);
PNG (PNG).
australis Macquart, 1840: 118 [1841: 396]. Australia (NSW);
widesp. Australia.
hilaris White, 1916b: 188. Australia (Tas).
balpi Evenhuis, 1979b: 23. Australia (WA); Australia (NT).
cabon Evenhuis, 1979b: 24. Australia (WA).
chrysonotum Evenhuis, 1979b: 25. Australia (WA).
europacificus Evenhuis, 1979b: 27. French Polynesia (Society
Is).
flavocciput Evenhuis, 1979b: 24. Australia (ACT).
fumipennis Evenhuis, 1979b: 30. Australia (Qld); Australia
(ACT, NSW).
fusciscelis Evenhuis, 1979b: 30. French Polynesia (Society
Is).
inflatus Evenhuis, 1979b: 31. Australia (WA).
karakara Evenhuis, 1979b: 32. Australia (WA).
malekulanus Evenhuis, 1979b: 33. Vanuatu.
mononesensis Evenhuis, 1979b: 33. Indonesia (Maluku).
mysticus Evenhuis, 1979b: 33. Vanuatu.
neutralis Evenhuis, 1979b: 35. New Caledonia (New Caledonia);
New Caledonia (Loyalty Is).
paramonovi Evenhuis, 1979b: 36. Australia (NSW).
terminatus Evenhuis, 1979b: 37. Vanuatu.
validus Evenhuis, 1989: 362 (n. n. for robustus
Evenhuis). Australia (NT); Australia (Qld).
robustus Evenhuis, 1979b: 37. Australia (NT). [Preocc.
Cresson, 1919.]
PLICHTAMYIA Evenhuis, 1979b: 39. Type species: Geron
dispar Macquart, 1850, orig. des.
albihalteris Evenhuis, 1979b: 39. Solomon Is; PNG (Bismarck
Arch).
amboinensis Evenhuis, 1979b: 40. Indonesia (Maluku).
bowdeni Evenhuis, 1979b: 41. PNG (Bismarck Arch).
dispar Macquart, 1850: 426(122). Australia (Tas); Australia
(ACT, NSW, Qld, Vic).
cothurnatus Bigot, 1892b: 374. Australia (Tas).
erythroccipitalis Evenhuis, 1979b: 44. Solomon Is; PNG
(Bougainville I).
hemifuscis Evenhuis, 1979b: 45. PNG (PNG).
nigralis Roberts, 1929: 571. Australia (NSW); Australia
(Qld).
nigrocciput Evenhuis, 1979b: 48. Australia (Qld); Australia
(ACT, NSW, SA, Tas), PNG (PNG).
opacus Bowden, 1971d: 312. PNG (Bismarck Arch); Solomon
Is.
paraustralicus Evenhuis, 1979b: 51. Indonesia (Maluku).
robertsi Evenhuis, 1979b: 52. Australia (NSW); Australia
(Qld).
sparsus Bowden, 1971d: 312. PNG (Bismarck Arch).
westralicus Evenhuis, 1979b: 54. Australia (WA).
simplex Walker, 1858b: 90. Indonesia (Maluku); ?PNG
(PNG).
tenue Walker, 1857: 147. "Country unknown" [?
Australasian].
CHORISTUS Walker, 1852: 197. Type species: Choristus
bifrons Walker, 1852, mon.
ANASTOECHUS, authors, not Osten Sacken, 1877, misid.
annexus Roberts, 1928b: 434 (Anastoechus). Australia
(WA).
bifrons Walker, 1852: 198. Australia (NSW); Australia (WA).
perspicuus Roberts, 1928b: 433 (Anastoechus).
Australia (WA).
CRYOMYIA Hull, 1973: 131. Type species: Cryomyia
argyropila Hull, 1973, mon.
argyropila Hull, 1973: 131. Australia (WA).
perparva Roberts, 1928b: 453 (Anastoechus). Australia
(Qld).
DISSODESMA Bowden & Li in Li & Yeates, 2018: 328. Type species: Bombylius primogenitus Walker, 1849, orig. des.
flava Li & Yeates, 2018: 346. Ausrtalia (Qld); Australia (NSW).
immaculata Li & Yeates, 2018: 336. Australia (WA)
primogenita Walker, 1849: 292 (Bombylius). Australia (WA).
smarti Li & Yeates, 2018: 349. Australia (WA)
SYRPHOIDES Evenhuis, 1983a: 206. Type species: Bombylius
byrrhus Bowden, 1971, orig. des. [Preocc. Hippa, 1968.]
ERISTALOPSIS Evenhuis, 1985a: 289 (n. n. for Syrphoides).
APIFORMYIA Yeates, 2008: 32. Type species: Apiformyia australis Yeates, 2008, orig. des..
australis Yeates, 2008: 32 (Apiformyia). Australia (NT).
byrrha Bowden, 1971c: 297 (Bombylius). Australia
(WA).
robertsi Paramonov, 1934: 31 (Bombylius). Australia
(Qld).
rubriventris Bigot, 1892b: 365 (Bombylius). Australia
(NSW).
EUSURBUS Roberts, 1929: 580. Type species: Bombylius
crassilabris Macquart, 1855, orig. des.
BROMOGLYCIS Hull, 1971: 5. Type species: Bromoglycis
robustus Hull, 1971, mon.
altus Walker, 1849: 288 (Bombylius). Australia (WA).
pinguis Walker, 1849: 290 (Bombylius). Australia
(WA).
collaris Li & Yeates, 2016: 482. Australia (NSW); Australia (Qld).
crassilabris Macquart, 1855: 97(77) (Bombylius).
Australia (NSW); Australia (Qld).
robustus Hull, 1971: 7 (Bromoglycis). Australia (Qld).
danielsi Li & Yeates, 2018: 493. Australia (NSW).
glossus Li & Yeates, 2018: 497. Australia (NSW); Australia (Qld).
LAMBKINOMYIA Li & Yeates, 2018: 357. Type species: Bombylius cinctiventris Roberts, 1928, orig.
des.
cinctiventris Roberts, 1928b: 424 (Systoechus).
Australia (Qld).
flavovillosa Roberts, 1928b: 423 (Systoechus). Australia
(Qld).
pallida Roberts, 1928b: 422 (Systoechus). Australia
(Qld).
rubida Roberts, 1928b: 421 (Systoechus). Australia
(Qld); Australia (NSW).
LAURELLA Hull, 1971: 3. Type species: Laurella auripila
Hull, 1971 [= Dischistus formosus Roberts, 1928], orig.
des.
formosa Roberts, 1928b: 451 (Dischistus). Australia
(Qld); Australia (NSW).
auripila Hull, 1971: 3. Australia (Qld).
MANDELLA Evenhuis, 1983a: 207. Type species: Systoechus
albohirtus Roberts, 1928, orig. des.
SYSTOECHUS, authors, not Loew, 1855, misid.
albohirta Roberts, 1928b: 422 (Systoechus). Australia
(SA).
australis Guérin-Méneville, 1831: pl. 20,
fig. 4 [1838: 294] (Bombylius). Australia (NSW).
PARAMONOVELLA Hull, 1973: 61. Nomen nudum.
PARAMONOVELLA Zaitzev, 1982: 521. Nomen nudum.
MEOMYIA Evenhuis, 1983a: 210. Type species: Bombylius
platyurus Walker, 1849 [= Bombylius fasciculatus Macquart,
1840], orig. des.
SYSTOECHUS, authors, not Loew, 1855, misid.
albiceps Macquart, 1848: 196(36) (Bombylius). "Nouvelle-Hollande";
Australia (NSW, Qld, Vic, WA).
callynthrophora Schiner, 1868: 137 (Systoechus).
Australia (NSW); Australia (Qld).
fasciculata Macquart, 1840: 93 [1841: 371] (Bombylius).
"Patrie inconnue" [= Australia (NSW)]; Australia (Qld,
SA, Tas, Vic, WA).
platyurus Walker, 1849: 286 (Bombylius). Australia
(WA).
crassus Walker, 1849: 287 (Bombylius). "New
Holland.".
notatipennis Macquart, 1855: 98(78) (Bombylius).
Australia (NSW).
platyrus, error for platyurus.
pausaria Jaennicke, 1867: 348 (Systoechus). "Australia."
penicillata Macquart, 1850: 422(118) (Bombylius).
Australia, "côte orientale."
sericans Macquart, 1850: 420(116) (Bombylius). Australia,
"côte orientale"; Australia (Qld, WA).
tetratricha Walker, 1849: 291 (Bombylius). Australia
(WA).
vetusta Walker, 1849: 286 (Bombylius). "New
Holland."
yeatesi Evenhuis & Greathead, 1999: 160 (n.n. for Bombylius punctipennis Thomson, 1869). [Preocc. Loew, 1855].
punctipennis Thomson, 1869: 487 (Bombylius). Australia
(NSW).
NIGROMYIA Li & Yeates, 2018: 362. Type species: Nigromyia flavamana Li & Yeatres, 2018, orig. des.
collessi Li & Yeates, 2018: 364. Australia (NT).
crocea Li & Yeates, 2018: 369. Australia (Qld).
flavamana Li & Yeates, 2018: 374. Australia (NT).
longistriata Li & Yeates, 2018: 376. Australia (NT).
pantherina Li & Yeates, 2018: 379. Australia (WA).
parva Li & Yeates, 2018: 383. Australia (WA).
tomentosa Li & Yeates, 2018: 385. Australia (NT).
PILOSIA Hull, 1973: 137. Type species: Pilosia flavopilosa
Hull, 1973, mon.
flava Li & Yeates, 2018: 305. Australia (WA).
immutata Walker, 1849: 292 (Bombylius). Australia
(WA).
flavopilosa Hull, 1973: 137. Australia (WA).
incana Bowden & Li, 2018: 308. Australia (WA).
ROBERTSMYIA Li & Yeates, 2018: 386. Type species: Dischistus pallioventer Roberts, 1928, mon.
pallidoventris Roberts, 1928: 452 (Dischistus). Australia (Tas); Australia (NSW, Vic).
SISYROMYIA White, 1916b: 192, 197. Type species: Bombylius
auratus Walker, 1849, des. Bezzi, 1924a: 7.
BOMBYLIUS, authors, not Linnaeus, 1758, misid.
areolatus, error for aureolatum.
albavitta Macquart, 1850: 421(117) (Bombylius).
Australia, "côte orientale"; Australia (NSW, Qld,
WA).
albovitta, albovittata, errors for albavitta.
aurata Walker, 1849: 289 (Bombylius). Australia
(Tas) & (WA); Australia (NSW, SA).
binghi Evenhuis, 1989: 364. (Brychosoma; n. n. for brevirostris
Macquart). Australia (?NSW).
brevirostris Macquart, 1850: 423(119) (Bombylius).
"Tasmanie", error [prob. = Australia (NSW)]. [Preocc.
Olivier, 1789.]
crassirostris Macquart, 1850: 423(117) (Bombylius).
Australia, "côte orientale" [prob. = Australia
(NSW)].
decorata Walker, 1849: 291 (Bombylius). Australia
(WA).
eulabiata Bigot, 1892b: 366 (Systoechus). Australia
(Tas).
limbata Bigot, 1892b: 369 (Sparnopolius). "Australie";
Australia (SA, WA).
lobalis Thomson, 1869: 487 (Bombylius). Australia
(NSW).
loewii Jaennicke, 1867: 346 (Bombylius). "Australia."
pycnorrhyncha Thomson, 1869: 486 (Bombylius). Australia
(NSW).
rutila Walker, 1849: 289 (Bombylius). Australia
(NSW).
thomsoni Evenhuis & Greathead, 1999: 170 (n.n. for scutellaris Thomson, 1869). Australia (NSW).
scutellaris Thomson, 1869: 480 (Bombylius). Australia
(NSW).
STAUROSTICHUS Hull, 1973: 76, 104. Type species: Staurostichus
chiastoneura Hull, 1973, orig. des.
BOMBYLIUS, authors, not Linnaeus, 1758, misid.
BRYCHOSOMA Hull, 1973: 102. Type species: Brychosoma
pulchrum Hull, 1973 [= Bombylius pictipennis Macquart,
1850], mon.
albicinctus Macquart, 1847a: 54 [1847b: 70] (Bombylius).
Australia (Tas).
alienus Hardy, 1942: 204 (Bombylius; n. n. for pulchellus
Roberts). Australia (NSW).
pulchellus Roberts, 1928b: 441 (Bombylius).
Australia (NSW). [Preocc. Loew, 1863.]
aureolatus Walker, 1857: 145 (Bombylius). "Australia";
Australia (NSW, Qld).
chiastoneura Hull, 1973: 104. Australia (Qld).
chrysendetus White, 1916b: 195 (Bombylius). Australia
(Tas).
consobrinus Macquart, 1847a: 54 [1847b: 70] (Bombylius).
"Nouvelle-Hollande" & Australia (Tas).
dulcis Roberts, 1928b: 440 (Bombylius). Australia
(Qld).
fuscanus Macquart, 1850: 423(119) (Bombylius). Australia
(Tas); Australia (Vic).
hilaris Walker, 1849: 274 (Bombylius). Australia
(WA); Australia (NSW, Qld).
llimbipennis Macquart, 1840: 96 [1841: 374] (Bombylius).
"Patrie inconnue" [= Australia]; Australia (NSW).
australianus Bigot, 1892b: 364 (Bombylius).
Australia (NSW).
matutinus Walker, 1849: 281 (Bombylius). "New
Holland."
pictipennis Macquart, 1850: 422(118) (Bombylius).
Australia, "côte orientale"; Australia (NSW, Qld).
pulchrum Hull, 1973: 104. Australia (Qld).
proprius Roberts, 1928b: 436 (Bombylius). Australia
(NSW).
sciotus Bowden, 1971c: 296 (Bombylius). Australia
(Tas).
tenuicornis Macquart, 1850: 423(116) (Bombylius).
Australia (Tas).
viduus Walker, 1852: 199 (Bombylius). Australia
(Tas).
palliolatus White, 1916b: 194 (Bombylius). Australia
(Tas).
ZENTAMYIA Li & Yeates, 2018: 502. Type species: Eusurbus nigracinctus Roberts, 1929, orig. des.
nigracinctus Roberts, 1929: 581 (Eusurbus).
Australia (WA).
nigrivinctus, error for nigracinctus.
distinctus Walker, 1852: 201 (Bombylius). Australia
(NSW & WA).
leucopygus Wulp, 1885c: 86 (Systoechus). Australia
(SA).
primogenitus Walker, 1849: 292 (Bombylius). Australia
(WA).
probellus Hardy, 1942: 204 (Bombylius; n. n. for
bellus Roberts). Australia (NSW).
bellus Roberts, 1928b: 439 (Bombylius). Australia
(NSW). [Preocc. Philippi, 1865.]
succandidus Roberts, 1928b: 438 (Bombylius). Australia
(NSW).
tenuirostris Roberts, 1928b: 437 (Bombylius). Australia
(Qld).
walkeri Evenhuis, 1978a: 101 (Bombylius; n. n. for
nanus Walker). Australia (WA).
nanus Walker, 1849: 278 (Bombylius). Australia
(WA). [Preocc. Meigen, 1838.]
MARMASOMA White, 1916b: 184, 188. Type species: Marmasoma
sumptuosa White, 1916, mon.
hortorum Li & Yeates, 2018: 37. Australia (WA).
sumptuosum White, 1916b: 190. Australia (Tas) & (Vic); Australia ACT, SA, Vic).
THEVENEMYIA Bigot, 1875a: 196 [1875b: clxxiv]. Type
species: Thevenemyia californica Bigot, 1875, mon.
THEVENETIMYIA Bigot, 1892b: 339 (emend. of Thevenemyia).
Type species: Thevenemyia californica Bigot,
1875, aut.
ECLIMUS, authors, not Loew, 1844, misid.
australiensis Hall, 1969: 25. Australia (NSW); Australia (Qld, Vic).
fergusoni Li & Rodriguez in Li et al., 2018: 339. Australia (Qld); Australia (ACT, NSW).
furvicostata Roberts, 1929: 578 (Eclimus). Australia
(Qld).
infuscata Li & Yeates in Li et al., 2018: 346. Australia (ACT).
longipalpus Hardy, 1921c: 63 (Eclimus). Australia
(NSW); Australia (ACT, Qld, SA, Vic, WA).
major Li & Yeates in Li et al., 2018: 353. Australia (Qld); Australia (NT).
mimula Hall, 1969: 61. Not given [Australia]; Australia
(Qld).
nigrapicalis Roberts, 1929: 576 (Eclimus). Australia
(Qld); Australia (ACT, NSW, NT).
nuri Rodriguez & Lamas in Li et al., 2018: 362. Australia (Qld).
tenta Hall, 1969: 75. Australia (NSW); Australia (Qld).
TILLYARDOMYIA Tonnoir, 1927: 101. Type species: Tillyardomyia
gracilis Tonnoir, 1927, mon.
gracilis Tonnoir, 1927: 103. NZ (NZ).
ACREOTRICHUS Macquart, 1850: 425(121). Type species:
Acreotrichus gibbicornis Macquart, 1850 [= Bombylius
antecedens Walker, 1849], orig. des.
ACEOTRICHUS. Incorr. orig. spell. of Acreotrichus [Macquart,
1850: 315 (11)].
antecedens Walker, 1849: 293 (Bombylius). "Australia";
Australia (NSW, NT, WA).
gibbicornis Macquart, 1850: 425(121). Australia, "côte
orientale."
fusicornis Macquart, 1850: 426(122). Australia, "côte
orientale."
inappendiculatus Bigot, 1892b: 366. "Australie."
lineifer Walker, 1857: 146 (Phthiria). "Australia";
Australia (NSW).
AUSTRALIPHTHIRIA Evenhuis, 1986: 8. Type species: Phthiria
hilaris Walker, 1852, orig. des.
albocapita Roberts, 1929: 573 (Phthiria). Australia
(WA); Australia (NT).
hilaris Walker, 1852: 194 (Phthiria). Australia
(NSW); Australia (ACT).
pallipes Bigot, 1892b: 367 (Phthiria). "Australie."
PHTHIRAXIA Li & Yeates, 2016: 3. Type
species: Phthiraxia bowdeni Li & Yeates, 2016, orig. des.
bowdeni Li & Yeates, 2016: 3. Australia(WA).
PYGOCONA Hull, 1973: 195 (Phthiria subg.). Type
species: Phthiria flavicincta Hull, 1973 [= Phthiria
flava Hardy, 1933], orig. des.
flava Hardy, 1933b: 415 (Phthiria). Australia (Vic).
flavicincta Hull, 1973: 195, 199 (Phthiria).
Not given [Australia].
nigrina Hardy, 1933b: 415 (Phthiria). Australia
(Vic); Australia (NSW).
NEOSARDUS Roberts, 1929: 560. Type species: Neosardus
principius Roberts, 1929, orig. des.
australis Yeates, 1996a: 53. Australia (SA); Australia
(Qld).
circumdatus Roberts, 1929: 561. Australia (NSW); Australia
(Qld).
cosmus Yeates, 1996a: 54. Australia (WA).
danielsi Yeates, 1996a: 56. Australia (Qld); Australia
(NSW, NT, SA).
decus Yeates, 1996a: 57. Australia (NSW).
elegans Yeates, 1996a: 58. Australia (WA).
eremae Yeates, 1996a: 58. Australia (NT); Australia (WA).
fenestratus Yeates, 1996a: 59. Australia (WA)
lepidus Roberts, 1929: 561. Australia (NSW); Australia
(Qld).
millstreamensis Yeates, 1996a: 62. Australia (WA).
nigratus Roberts, 1929: 563. Australia (WA); australia
(SA, NT).
paramonovi Hull, 1973: 300. Australia (WA).
percultus Yeates, 1996a: 66. Australia (NT).
phaleratus Yeates, 1996a: 67. Australia (WA).
principius Roberts, 1929: 562. Australia (Qld); Australia
(NSW, SA, WA).
rieki Yeates, 1996a: 69. Australia (WA).
sparsus Yeates, 1996a: 71. Australia (WA).
vicinus Yeates, 1996a: 72. Australia (NSW); Australia (Qld,
WA).
ALEUCOSIA Edwards, 1934: 95. Type species: Neuria
corculum Newman, 1841, orig. des.
CHELINA Hull, 1973: 306 (Comptosia subg.). Type
species: Chelina norrisi Hull, 1973, orig. des.
PARAMONOVOMYIA Evenhuis, 1980: 329. Type species: Paramonovomyia
halli Evenhuis, 1980, orig. des.
abdita Yeates, 1991a: 171. Australia (WA).
adusta Yeates, 1991a: 203. Australia (WA).
affinis Yeates, 1991a: 178. Australia (WA).
albovittata Yeates, 1991a: 147. Australia (WA).
angusta Edwards, 1934: 100 (Comptosia). Australia
(WA).
aquila Yeates, 1991a: 180. Australia (WA).
arguta Yeates, 1991a: 165. Australia (WA).
atherix Newman, 1841: 222 (Neuria). Australia (SA);
Australia (ACT, NSW, SA, Tas, Vic).
tricellata Macquart, 1847a: 53 [1847b: 69] (Comptosia).
Australia (Tas).
geometrica Macquart, 1847a: 53 [1847b: 69] (Comptosia).
Australia (Tas).
obscura Walker, 1852: 176 (Anthrax). Australia
(Tas). [Preocc. Weber, 1801.]
tripunctata Edwards, 1934: 98 (Comptosia). Australia
(Tas).
australensis Evenhuis, 1980: 332 (Paramonovomyia).
Australia (WA).
australica Evenhuis, 1989: 366 (Alyosia; n.n.
for australensis Evenhuis). Australia (WA).
calophthalma Thomson, 1869: 129 (Comptosia). Australia
(NSW); Australia (ACT, Qld, SA, Vic, Tas).
cincta Edwards, 1934: 99 (Comptosia). Australia
(WA).
colona Yeates, 1991a: 187. Australia (SA).
corculum Newman, 1841: 221 (Neuria). Australia (WA);
Australia (SA).
costalis Edwards, 1934: 99 (Comptosia). Australia
(Qld); Australia (ACT, NSW).
cuneata Edwards, 1934: 100 (Comptosia). Australia";
Australia (WA).
danielsorum Yeates, 1991a: 202. Australia (NSW); Australia
(ACT, Vic).
directa Yeates, 1991a: 161. Australia (NSW).
dorsalis Walker, 1849: 269 (Anthrax). "New
Holland"; Australia (WA).
fulvipes Bigot, 1892b: 359 (Comptosia). "Australie";
Australia (NSW, SA, Vic, WA).
guttata Yeates, 1991a: 189. Australia (WA).
hemiteles Schiner, 1868: 132 (Neuria). Australia
(NSW); Australia (Qld).
hyalina Yeates, 1991a: 154. Australia (WA).
illota Yeates, 1991a: 197. Australia (WA).
macfarlandi Evenhuis, 1980: 332 (Paramonovomyia).
Australia (WA).
maculosa Newman, 1841: 221 (Neuria). Australia (WA);
Australia (SA, Vic).
marginata Yeates, 1991a: 169. Australia (NSW); Australia
(Qld).
multipunctata Yeates, 1991a: 185. Australia (WA).
norrisi Hull, 1973: 306. Australia (WA).
halli Evenhuis, 1980: 329 (Paramonovomyia).
Australia (WA).
obtusa Yeates, 1991a: 193. Australia (WA).
pallida Yeates, 1991a: 192. Australia (NSW); Australia
(Vic).
plena Walker, 1849: 270 (Anthrax). Australia (WA).
preclara Yeates, 1991: 170. Australia (WA).
saltuaria Yeates, 1991a: 155. Australia (WA).
serpentigera Walker, 1849: 270 (Anthrax). Australia
(WA).
sugillata Yeates, 1991a: 163. Australia (WA).
tridentata Yeates, 1991a: 175. Australia (WA).
COMPTOSIA Macquart, 1840: 80 [1841: 358]. Type species:
Comptosia fascipennis Macquart, 1840, mon.
NEURIA Newman, 1841: 220. Type species: Neuria lateralis
Newman, 1841, des. Hardy, 1921c: 54.
ALYOSIA Rondani, 1863: 54 [1864: 54]. Type species: Comptosia
maculipennis Macquart, 1846 [= Neuria ocellata Newman,
1841], des. Becker, 1913a: 465.
EPIDOSIA Hull, 1973: 306 (Comptosia subg.). Type
species: Comptosia vittata Edwards, 1934, orig. des.
OPSONIA Hull, 1973: 306 (Comptosia subg.). Type
species: Comptosia fenestrata Hull, 1973, orig. des.
ANTHOCOLON Hull, 1973: 307 (Comptosia subg.). Type
species: Comptosia casimira Hull, 1973, orig. des.
PARADOSIA Hull, 1973: 306 (Comptosia subg.). Type
species: Comptosia duofasciata Hull, 1973, orig. des.
LIGYRA, authors, not Newman, 1841, misid.
EPIDOSINA, error for Epidosia.
acantha Yeates, 1991b: 1151. Australia (WA); Australia
(Vic).
apicalis Macquart, 1848: 195(35). "Nouvelle-Hollande";
Australia (ACT, NSW, Qld, Vic).
aurescens Yeates, 1991b: 1087. Australia (WA).
aurifrons Macquart, 1850: 417(113). Australia, "côte
orientale."
australensis Schiner, 1868: 129 (Lomatia). Australia
(NSW); Australia (Qld).
bancrofti Edwards, 1934: 92. Australia (Qld).
barrowensis Yeates & Oberprieler, 2013: 351. Australia (WA).
basalis Walker, 1849: 267 (Anthrax). "New Holland"
[lectotype locality]; Australia (WA).
biguttata Edwards, 1934: 93. Australia (Qld).
brunnea Edwards, 1934: 90. Australia (SA); Australia (ACT,
NSW, SA, Vic).
caesariata Yeates, 1991b: 1059. Australia (Qld); Australia
(ACT, NSW).
calignea Yeates, 1991b: 1123. Australia (Qld).
capillata Yeates, 1991b: 1061. Australia (Qld); Australia
(NSW, Vic).
casimira Hull, 1973: 307. Not given [= Australia (WA)];
Australia (ACT).
decedens Walker, 1849: 271 (Anthrax). Australia
(WA); Australia (Vic).
extensa Walker, 1835a: 473 (Anthrax). Australia
(WA); Australia (ACT).
fascipennis Macquart, 1840: 81 [1841: 359]. "Monte
Video," error [Australia]; Australia (ACT, NSW, Qld, Vic).
flava Yeates, 1991b: 1079. Australia (WA); Australia (NSW).
flavipenna Yeates, 1991b: 1106. Australia (NT); Australia
(WA).
flexuosa Yeates, 1991b: 1096. Australia (WA); Australia
(SA).
gemina Hardy, 1941: 228 (n. n. for cognata Walker).
Australia (WA).
cognata Walker, 1852: 177 (Anthrax). Australia
(WA). [Preocc. Walker, 1849.]
fenestrata Hull, 1973: 306. Not given [Australia].
heliophila Yeates, 1991b: 1075. Australia (Qld); Australia
(NSW).
inclusa Walker, 1849: 268 (Anthrax). Australia (Tas);
Australia (ACT, NSW, Vic).
insignis Walker, 1849: 266 (Anthrax). "New
Holland"; Australia (NSW); Australia (ACT).
ducens Walker, 1852: 176 (Anthrax). Australia
(NSW).
rufoscutellata Jaennicke, 1867: 345. "Australia".
grandis Schiner, 1868: 132 (Neuria). Australia
(NSW).
albofasciata Thomson, 1869: 484. Australia (NSW).
nigroscutellata, error for rufoscutellata.
insula Yeates, 1991b: 1108. Australia (Qld).
karijinii Yeates & Oberprieler, 2013: 352. Australia (WA).
kuranda Yeates, 1991b: 1079. Australia (Qld).
lactea Yeates, 1991b: 1142. Australia (WA).
lateralis Newman, 1841: 220 (Neuria). Australia
(NSW).
mackerrasi Yeates, 1991b: 1114. Australia (Qld).
magna Yeates, 1991b: 1063. Australia (WA); Australia (SA).
mallota Yeates, 1991b: 1146. Australia (Qld).
microrhynchus Yeates, 1991b: 1154. Australia (SA); Australia
(WA).
moretonii Macquart, 1855: 95(75). Australia (NSW); Australia
(Qld); NZ (NZ) [introduced; not established; see Evenhuis, 1986b:
7.]
murina Newman, 1841: 221 (Neuria). Australia (SA).
neoapicalis Yeates, 1991b: 1117. Australia (NSW); Australia
(Qld).
neobiguttata Yeates, 1991b: 1112. Australia (NSW); Australia
(ACT, Qld).
neosobria Yeates, 1991b: 1078. Australia (Qld).
nigrescens Newman, 1841: 221 (Neuria). Australia
(NSW).
nitella Yeates, 1991b: 1091. Australia (WA).
ocellata Newman, 1841: 221 (Neuria). Australia (Tas).
maculipennis Macquart, 1846: 244(116). Australia (Tas).
paramonovi Yeates, 1991b: 1089. Australia (SA); Australia
(Vic).
partita Newman, 1841: 221 (Neuria). Australia (WA).
paucispina Yeates, 1991b: 1042. Australia WA).
pilosa Yeates, 1991b: 1056. Australia (WA); Australia (NSW,
SA, Vic).
pitereka Yeates, 1991b: 1048. Australia (SA).
praeargentata Macleay, 1826: 468 (Anthrax). Not
given [Australia]; Australia (ACT, NSW, Qld).
edwardsi Hardy, 1941: 227. Australia (Qld).
quadripennis Walker, 1849: 268 (Anthrax). "Australia";
Australia (ACT, NSW, Qld, SA, Vic).
rubrifera Bigot, 1881c: 23 (Lygira). "Australia";
Australia (NSW, Qld).
sandaraca Yeates, 1991b: 1081. Australia (WA).
scitula Yeates, 1991b: 1148. Australia (Qld).
sobria Walker, 1849: 269 (Anthrax). "Australia";
Australia (NSW); PNG (PNG).
bicolor Macquart, 1850: 418 (114). Australia, "côte
orientale."
subsenex Walker, 1857: 144 (Anthrax). "Australia."
sobricula Walker, 1857: 144 (Anthrax). "Australia";
Australia (ACT, NSW, Qld, Tas, Vic).
simplex Macquart, 1847a: 52 [1847b: 68] (Anthrax).
Australia (Tas). [Preocc. Wiedemann, 1819.]
anthracina Thomson, 1869: 485. Australia (NSW).
prosimplex Hardy, 1942: 203 (Villa; n. n. for
simplex Macquart). Australia (Tas).
sorbicula, error for sobricula.
soror Yeates, 1991b: 1115. Australia (Qld).
speciosa Yeates, 1991b: 1066. Australia (WA).
stria Walker, 1849: 267 (Anthrax). "Australia";
Australia (NSW, Tas).
fasciata Fabricius, 1805: 118 (Anthrax). "Insulis
oceani pacifica" [= Australia (Tas); see Evenhuis 1988].
[Preocc. Meigen, 1804.]
tendens Walker, 1849: 271 (Anthrax). Australia (WA).
duofasciata Hull, 1973: 306. Not given [= Australia (WA)].
thyris Yeates, 1991b: 1121. Australia (WA).
tutela Yeates, 1991b: 1045. Australia (Qld); Australia
(NSW).
vittata Edwards, 1934: 89. Australia (Vic); Australia (NSW,
SA).
walkeri Edwards, 1934: 94. Not given [Australia]; Australia
(NSW, Qld).
wilkinsi Edwards, 1934: 94. Australia (NT).
xanthobasis Yeates, 1991b: 1084. Australia (WA).
zona Yeates, 1991b: 1119. Australia (WA).
DODDOSIA Edwards, 1934: 103. Type species: Doddosia
picta Edwards, 1934, orig. des.
collessi Yeates, 1990: 78. Australia (NT).
lurida Yeates, 1990: 77. Austrlia (Qld); Australia (SA).
picta Edwards, 1934: 103. Australia (Qld); Australia (NT).
procax Yeates, 1990: 78. Australia (qld).
torresiana Yeates, 1990: 79. Australia (Qld).
ONCODOSIA Edwards, 1934: 101. Type species: Anthrax
patula Walker, 1849, orig. des.
ampla Walker, 1852: 185 (Anthrax). Australia (WA).
patula Walker, 1849: 273 (Anthrax). Australia (WA);
Australia (SA, Vic).
plana Walker, 1849: 272 (Anthrax). Australia (WA).
triangularis Yeates, 1988: 516. Australia (Vic); Australia
(SA, WA).
DOCIDOMYIA White, 1916b: 200, 203. Type species: Docidomyia
puellaris White, 1916, mon.
PARACOSMUS, Evenhuis, 1989, not Osten Sacken, 1877, misid.
albifrons Yeates, 1996b: 410. Australia (NT); Australia
(Qld, WA).
angusta Yeates, 1996b: 413. Australia (WA).
brevipilata Yeates, 1996b: 414. Australia (Qld); Australia
(NSW).
danielsi Yeates, 1996b: 415. Australia (NSW); Australia
(NT, WA).
froggatti Paramonov, 1951a: 748. Australia (NSW).
genualia Yeates, 1996b: 417. Australia (WA); Australia
(NT).
hespera Yeates, 1996b: 420. Australia (WA).
isla Yeates, 1996b: 421. Australia (Qld).
nicholsoni Yeates, 1996b: 422. Australia (WA).
nigra Yeates, 1996b: 423. Australia (WA).
norrisi Paramonov, 1951a: 748. Australia (WA).
parva Yeates, 1996b: 425. Australia (Qld); Australia (NSW).
puellaris White, 1916b: 204. Australia (Tas) & (Vic);
Australia (ACT, NSW, Qld, SA).
swani Paramonov, 1951a: 750. Australia (SA); Australia
(NSW).
victoriana Paramonov, 1951a: 749. Australia (Vic); Australia
(NSW).
ANTONIAUSTRALIA Becker, 1913a: 458. Type species: Antoniaustralia
hermanni Becker, 1913, orig. des.
arida Evenhuis, 1993: 3. Australia (NT).
hermanni Becker, 1913a: 458. "Australien" [=
Australia (NT)].
CYX Evenhuis, 1993: 7. Type species: Antonia rieki
Paramonov, 1953, orig. des.
decorata Paramonov, 1950d: 532 (Antonia). Not given
[Australia]; Australia (Qld).
papuana Evenhuis, 1993: 10. PNG (PNG).
rieki Paramonov, 1953c: 204 (Antonia). Australia
(NSW).
roddi Paramonov, 1950d: 530 (Antonia). Australia
(NSW).
MYONEMA Roberts, 1929: 563. Type species: Myonema
humile Roberts, 1929, orig. des.
humile Roberts, 1929: 564. Australia (Qld).
ANTHRAX Scopoli, 1763: 358. Type species: Musca morio
Linnaeus, 1758 [misid., = Musca anthrax Schrank, 1781],
mon.
arcus de Meijere, 1913b: 322 (Argyramoeba). Indonesia
(Irian Jaya).
argentius Yeates & Lambkin, 1998: 991 (as argentia).
Australia (WA); Australia (SA).
angularis Thomson, 1869: 482. Australia (NSW); Australia
(NT, Qld, SA, WA).
asciculus Yeates & Lambkin, 1998: 1010. Australia (Qld);
Australia (NSW, SA, Vic).
clinatus Yeates & Lambkin, 1998: 1013. Australia (WA);
Australia (NSW, NT, Qld).
coeruleopennis Doleschall, 1857: 400. Indonesia (Maluku);
?PNG (PNG).
caeruleopennis, error for coeruleopennis.
confluensis Roberts, 1928: 139. Australia (Qld); Australia
(ACT, NSW, Nt, SA, Vic).
crenatus Yeates & Lambkin, 1998: 1018. Australia (NT);
Australia (ACT, Qld, SA, Vic, WA).
distigma Wiedemann, 1828: 309. Indonesia (Java); Indonesia
(Maluku), PNG (PNG).
doleschalli Evenhuis & Greathead, 1999: 299 (n.n. for argyropyga
Doleschall, 1857). Indonesia (Maluku).
argyropyga Doleschall, 1857: 401. Indonesia (Maluku).
[Preocc. Wiedemann, 1828.]
dolobratus Yeates & Lambkin, 1998: 1022. Australia
(NSW); Australia (ACT, Qld, Vic).
emissa Walker, 1864: 233. Indonesia (Maluku); PNG (PNG).
emittens Walker, 1860b: 148. Indonesia (Maluku).
incisus Macquart, 1847a: 52 [1847b: 68]. Australia (Qld).
incomptus Walker, 1849: 253. Australia (WA); Australia
(ACT, NSW, NT, Qld, SA, Vic), ?PNG (Bismarck Arch), Vanuatu; Christmas
I (Indian Ocean).
semimacula Walker, 1849: 254. "New Holland."
koshunensis Matsumura, 1916: 282. Taiwan; Bonin Is, Guam,
Hawaiian Is [introduced]; Japan.
distigma, authors, not Wiedemann, misid.
lepidiotus Roberts, 1928a: 138. Australia (Qld); Australia
(NT).
maculatus Macquart, 1846: 240(112). Australia (Tas); Australia
(ACT, NSW, NT, Qld, SA, Vic, WA).
diana Walker, 1849: 252. Not given [= Australia (WA)].
australis Walker, 1852: 193. "Van Diemen's Land"
[= Australia (Tas)] & Australia (WA).
matilei Evenhuis, 1991: 281. New Caledonia (New Caledonia).
melanius Wulp, 1885c: 84. Indonesia (Java); PNG (PNG).
opacus Yeates & Lambkin, 1998: 1018. Australia (Qld);
Australia (NSW, SA).
proconcisus Hardy, 1942: 203 (Villa; n. n. for concisa
Macquart, 1850). Australia, "côte orientale" [=
eastern Australia]; Australia (ACT, NSW, Qld, SA, Vic).
concisa Macquart, 1850: 415(111). Australia "côte
orientale" [= eastern Australia] [Preocc. Macquart, 1840.]
semiscitus Walker, 1857: 118. Malaysia (Sarawak); ?French
Polynesia (Society Is), Indonesia (Irian Jaya), PNG (Bismarck
Arch, Bougainville I, PNG), Solomon Is; Indonesia (Sulawesi).
tripunctata Wulp, 1869: 109. Indonesia (Irian Jaya).
[Preocc. Wiedemann, 1820.]
siccus Yeates & Lambkin in Evenhuis & Greathead,
1999: 314 (n.n. for funestus Yeates & Lambkin, 1998).
Australia (WA); Australia (Vic).
funestus Yeates & Lambkin, 1998: 1024. Australia
(WA). [Preocc. Walker, 1849.]
succedens Walker, 1852: 192. Not given [unknown].
torulus Yeates & Lambkin, 1998: 1001. Australia (Qld);
Australia (NT, WA).
BRACHYANAX Evenhuis, 1981: 190. Type species: Anthrax
satellita Walker, 1857 [= Brachyanax thelestrephones
Evenhuis, 1981], orig. des.
HEMIPENTHES, authors, not Loew, 1869, misid.
ater Roberts, 1928a: 140 (Anthrax). Australia (Qld);
Indonesia (Nusa Tenggara).
australianus Evenhuis, 1981: 197. Australia (Qld).
aterrimus Doleschall, 1858: 93 (Anthrax). Indonesia
(Maluku); PNG (Bismarck Arch, PNG), Solomon Is; Indonesia (Sulawesi),
Japan, Philippines.
proferens Walker, 1859c: 113 (Anthrax). Indonesia
(Sulawesi).
oceanicus Evenhuis, 1981: 197. Solomon Is.
chichijimensis Evenhuis, 1981: 195. Bonin Is.
leucostigma Wulp, 1898a: 419 (Anthrax; n. n. for
terminalis Wulp). Indonesia (Maluku); PNG (PNG).
terminalis Wulp, 1869: 108 (Anthrax). Indonesia
(Maluku). [Preocc. Wiedemann, 1830.]
magnipennis Evenhuis, 1981: 195. Solomon Is.
papuanus Evenhuis, 1981: 198. PNG (PNG).
satellitius Walker, 1857: 119 (Anthrax). Malaysia
(Sarawak); Indonesia (Maluku); Indonesia (Kalimantan), Malaysia
(Pen, Sabah).
satellita, error for satellitius.
THRAXAN Yeates & Lambkin, 1998: 1027. Type species:
Thraxan simulans Yeates & Lambkin, 1998, orig. des.
abditus Yeates & Lambkin, 1998: 1030. Australia (Qld);
Australia (NSW).
acutus Yeates & Lambkin, 1998: 1031. Australia (Qld);
Australia (NSW).
caligneus Yeates & Lambkin, 1998: 1034. Australia (Qld).
cinctus Yeates & Lambkin, 1998: 1037. Australia (Qld);
Australia (NSW, NT, WA).
cornuatus Yeates & Lambkin, 1998: 1039. Australia (WA);
Australia (NT, Qld).
depressus Yeates & Lambkin, 1998: 1041. Australia (NSW);
Australia (NT, Qld).
ebeneus Yeates & Lambkin, 1998: 1042. Australia (ACT);
Australia (Qld).
echinatus Yeates & Lambkin, 1998: 1030. Australia (Qld);
Australia (NT, WA).
emicatus Yeates & Lambkin, 1998: 1045. Australia (NT);
Australia (WA).
hamulus Yeates & Lambkin, 1998: 1046. Australia (Qld).
luteus Yeates & Lambkin, 1998: 1047. Australia (NT);
Australia (Qld, WA).
misatulus Yeates & Lambkin, 1998: 1049. Australia (WA);
Australia (NSW, NT, Qld, Vic).
nodus Yeates & Lambkin, 1998: 1051. Australia (Qld).
norrisi Yeates & Lambkin, 1998: 1052. Australia (WA).
obstipus Yeates & Lambkin, 1998: 1054. Australia (NT);
Australia (NSW, Qld).
patielus Yeates & Lambkin, 1998: 1055. Australia (NSW);
Australia (ACT, Qld).
planus Yeates & Lambkin, 1998: 1056. Australia (Qld);
Australia (NT, WA).
prolatus Yeates & Lambkin, 1998: 1057. Australia (Qld);
Australia (NSW).
simulatus Yeates & Lambkin, 1998: 1028. Australia (Qld).
spiculus Yeates & Lambkin, 1998: 1059. Australia (Qld).
boninensis Matsumura, 1916: 284 (Agryramoeba).
Bonin Is.
Tribe EXOPROSOPINI
Genus ATRICHOCHIRA Hesse
ATRICHOCHIRA Hesse, 1956; 911. type species: Exoprosopa pediformis Bezzi, 1921, orig. des.
commoni Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 855. Australia (Qld).
paramonovi Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 857. Australia (NSW); Australia (Qld).
Genus BALAANA Lambkin & Yeates
BALAANA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 757. Type species: Exoprosopa latelimbata Bigot, 1892, orig. des.
abscondita Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 761. Australia (NW); Australia (ACT, NT, Qld, SA, Vic, WA).
biscuspis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 764. Australia (NT); Australia (Qld, WA).
centrosa Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 766. Australia (WA); Australia (NT, SA, Vic, WA).
gigantea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 768. Australia (Qld); Australia (NSW, Vic, WA).
kingcascadensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 770. Australia (WA); Australia (NT).
latelimbata Bigot, 1892b: 346 (Exoprosopa). "Australie"; Australia (ACT, NSW, Qld, SA).
obliquebifasciata Macquart, 1850: 411(107) (Exoprosopa). Australia (Tas).
obliquefasciata, error for obliquebifasciata.
Genus EXOPROSOPA Macquart
EXOPROSOPA Macquart, 1840: 35 [1841: 313]. Type species:
Anthrax pandora Fabricius, 1805, des. Coquillett (1910:
544) [Designation conserved by I.C.Z.N. 1991: 82.].
bicellata Macquart, 1847a: 51 [1847b: 61]. Australia (Tas).
sylvana Fabricius, 1775: 758 (Bibio). "nova
Hollandia" [= Australia (Qld); see Musgrave, 1932: 54].
Genus KAPU Lambkin & Yeates
KAPUA Lambkin & Yeates, 2003: 776. Type species: Exoprosopa adelaidica Macquart, 1855, orig. des. [Preocc. Marwick, 1931.]
KAPU Lambkin & Yeates, 2006: 161 (n.n. for Kapua Lambkin & Yeates).
adelaidica Macquart, 1855: 90(70) (Exoprosopa). Australia (SA); Australia
(NSW, NT, Qld, Tas, WA).
corusca Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 780. Australia (NSW); Australia (ACT, NT, Qld, WA).
irwini Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 783. Australia (Qld); Australia (NSW, SA).
westralica Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 785. Australia (WA).
Genus LARRPANA Lambkin & Yeates
LARRPANA Lambkin & Yeates, 2003: 787. Type species: Exoprosopa dimidiatipennis Bowden, 1971, orig. des.
bushblitz Lambkin & Bartlett, 2011: 248. Australia (Qld) .
collessiLambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 783. Australia (NT); Australia (Qld).
dimidiatipennis Bowden, 1971b: 64 (Exoprosopa; n. n. for dimidiata Roberts). Australia (NT).
dimidiata Roberts, 1928a: 109 (Exoprosopa). Australia (NT). [Preocc.
Macquart, 1846.]
zwicki Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 793. Australia (Qld).
LIGYRA Newman, 1841: 220. Type species: Anthrax bombyliformis
Macleay, 1826, mon.
LYGIRA, error for Ligyra.
HYPERALONIA, authors, not Rondani, 1863, misid.
alea Bowden, 1971d: 321. PNG (Bismarck Arch).
audouinii Macquart, 1840: 36 [1841: 314] (Exoprosopa).
"Indes orientales"; Indonesia (Irian Jaya, Maluku);
China, Indonesia (Java, Sulawesi), Philippines, Taiwan.
albicincta Macquart, 1840: 48 [1841: 316] (Exoprosopa).
China.
formosana Paramonov, 1931: 195 (Hyperalonia).
Taiwan.
macassarensis Paramonov, 1931: 198 (Hyperalonia).
Indonesia (Sulawesi).
bombyliformis Macleay, 1826: 468 (Anthrax). Australia
(NSW); Australia (NT, Qld, WA).
albiventris Thomson, 1869: 480 (Exoprosopa).
Australia (NSW).
bombiliformis, bombyliiformis, errors for bombyliformis.
burnsi Paramonov, 1967: 143. Australia (NSW).
calabyana Paramonov, 1967: 136. Australia (WA); Australia
(NT).
campbelli Paramonov, 1967: 139. Australia (NT).
chinnicki Paramonov, 1967: 138. Australia (SA); Australia
(NSW, Vic).
cingulata Wulp, 1885b: 62 (Exoprosopa). Australia
(SA); Australia (NSW, NT, Qld, WA).
incompleta Paramonov, 1967: 141 (cingulata var.).
[Unavailable name; varietal name proposed after 1960.]
commonsi Paramonov, 1967: 134. Australia (Qld).
cruciata Bowden, 1971a: 11. PNG (PNG).
dentata Roberts, 1928a: 102 (Hyperalonia). Australia
(Vic).
devecta Walker, 1860b: 148 (Anthrax). Indonesia
(Maluku).
delecta, error for devecta.
doryca Boisduval, 1835: 665 (Anthrax). Indonesia
(Maluku); Australia (NSW, Qld), Indonesia (Irian Jaya, Maluku),
PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia
(Java, Timor).
pelops Walker, 1858b: 90 (Anthrax). Indonesia
(Maluku).
eyreana Paramonov, 1967: 136. Australia (SA).
fasciata Paramonov, 1967: 135. Australia (Qld).
flaviventris Doleschall, 1857: 400 (Anthrax). Indonesia
(Maluku).
flavotomentosa de Meijere, 1929a: 386 [1929b: 416] (Hyperalonia).
Indonesia (Maluku).
fuscipennis Macquart, 1848: 193(33) (Exoprosopa).
Indonesia (Java); "Australia," PNG (PNG); "Borneo,"
Cambodia, China, ?India, Indonesia (Java, Sulawesi, Sumatra),
Malaysia (Pen), ?Nepal.
confirmata Walker, 1861d: 283 (Anthrax). Indonesia
(Maluku).
chrysolampis Jaennicke, 1867: 344 (Hyperalonia).
Indonesia (Java).
tantalus, authors, not Fabricius, 1794, misid.
hemifusca Roberts, 1928a: 97 (Hyperalonia). Australia
(Qld); Australia (NT, WA).
inquinita Roberts, 1928a: 103 (Hyperalonia). Australia
(Qld); Australia (NSW).
leptyna Bowden, 1971a: 12. PNG (Bougainville I).
leuconoe Jaennicke, 1867: 345 (Exoprosopa). Indonesia
(Maluku).
nigrocostalis Guérin-Méneville, 1838: 295
(Anthrax). Indonesia (Maluku)
ochracea Bowden, 1971a: 10. PNG (PNG).
orest Paramonov, 1967: 134. Australia (Qld); Australia
(NT).
pilad Paramonov, 1967: 143. Australia (WA).
punctipennis Macquart, 1850: 410(106) (Exoprosopa).
Australia, "côte orientale"; widesp. Australia.
contrasta Paramonov, 1967: 133. Australia (NSW).
robertsi Paramonov, 1967: 132. Australia (WA); Australia
(NSW, NT, Qld).
satyrus Fabricius, 1775: 758 (Bibio). "nova
Hollandia" [= Australia (Qld); see Musgrave, 1932: 54]; widesp.
Australia, Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch,
Bougainville I, PNG); ?China, Indonesia (Timor).
funesta Walker, 1849: 242 (Anthrax). Australia
(NT).
insignis Macquart, 1855: 93(73) (Exoprosopa).
Australia (Qld).
septentrionis Roberts, 1928a: 103 (Hyperalonia).
Australia (Qld); Australia (NT, WA).
sinuatifascia Macquart, 1855: 92(72) (Exoprosopa).
Australia (NSW); Australia (ACT, SA, Vic, WA).
macraspis Thomson, 1869: 479 (Exoprosopa). Australia
(NSW).
argenticincta Bigot, 1892b: 343 (Hyperalonia).
Australia (NSW).
tantalus Fabricius--not Australasian/Oceanian.
tenebrosa Paramonov, 1967: 132. Australia (Qld).
unicincta Guérin-Méneville, 1831: pl. 20,
fig. 9 [1838: 294] (Anthrax). Indonesia (Maluku).
ventrimacula Doleschall, 1857: 399 (Anthrax). Indonesia
(Maluku).
LITOMYZA Hull, 1973: 426. Nomen nudum.
LITORHINA Bowden, 1975: 314. Type species: Litorhynchus
corticeus Bezzi, 1924, orig. des.
LITORHYNCHUS, authors, not Macquart, 1840, misid.
collaris Wiedemann--not Australasian/Oceanian.
lar Fabricius, 1782: 414 (Bibio). India; ?Australia (WA);
Sri Lanka.
Genus MUNJUA Lambkin & Yeates
MUNJUA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 795. Type species: Munjua erugata Yeates & Lambkin, 2003, orig. des.
erugata Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 795. Australia (WA); Australia (NSW, NT, SA, Vic).
lepidokingi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 799. Australia (WA).
paralutea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 801. Australia (NSW); Australia (NT, Qld, SA, WA).
Genus MUWARNA Lambkin & Yeates
MUWARNA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 807. Type species: Anthrax stellifer Walker, 1849, orig. des.
stellifera Walker, 1849: 244 (Anthrax). "New
Holland" & Australia (WA); Australia (SA, Vic, WA).
vitreilinearis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 810. Australia (Vic); Australia (WA).
Genus NGALKI Lambkin
NGALKI Lambkin in Lambkin & Bartlett, 2011: 254. Type species: Munjua trigona Lambkin & Yeates, 2003, orig. des.
trigonum Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 804 (Munjua). Australia (Vic); Australia (NSW, SA, WA).
Genus PALRIKA Yeates & Lambkin
PALRIKA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 811. Type species: Palrika decora Lambkin & Yeates, 2003, orig. des.
anaxios Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 814. Australia (Qld).
basilikos Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 817. Australia (WA).
blackdownensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 820. Australia (Qld).
bouchardi Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 821. Australia (Qld); Australia (NSW, NT, WA).
culgoafloodplainensis Lambkin & Bartlett, 2011: 242. Australia (Qld) .
cyanea Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 823. Australia (Qld); Australia (NT, WA).
danielsi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 824. Australia (NSW); Australia (Qld).
decora Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 812. Australia (Qld); Australia (ACT).
mackenziei Lambkin & Bartlett, 2011: 245. Australia (Qld) .
marginicollis Gray in Griffith & Pidgeon, 1832:
780 (Anthrax). Not given [Australia]; Australia (NSW, NT, Qld, WA).
viridula Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 828. Australia (Qld); Australia (NSW).
whyalla Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 831. Australia (SA); Australia (WA).
PSEUDOPENTHES Roberts, 1928a: 132. Type species: Pseudopenthes
fenestrata Roberts, 1928, orig. des.
fenestrata Roberts, 1928a: 133. Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
hesperis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 863. Australia (WA).
PTEROBATES Bezzi, 1921b: 130 (Exoprosopa subg.).
Type species: Anthrax apicalis Wiedemann, 1821, mon.
pennipes Wiedemann, 1821b: 129 (Anthrax). Indonesia
(Java); Indonesia (Maluku); China, Hong Kong, India, Macau, Malaysia
(Sabah), Philippines.
Genus WURDA Lambkin & Yeates
WURDA Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 833. Type species: Wurda emu Lamkin & Yeates, 2003, orig. des.
emu Lambkin & Yeates in Lambkin in Lambkin, Yeates & Greathead, 2003: 833. Australia (NSW); Australia (NT, Qld, SA, WA).
impatientis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 838. Australia (WA); Australia (SA).
montebelloensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 840. Australia (WA); Australia (NT, Qld, SA).
norrisi Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 843. Australia (WA); Australia (SA, Vic).
patrellia Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 845. Australia (NSW).
skevingtoni Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 847. Australia (WA).
windorah Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 848. Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
wyperfeldensis Lambkin & Yeates in Lambkin, Yeates & Greathead, 2003: 851. Australia (Vic); Australia (NSW, Qld, SA, WA).
albirufus Walker, 1857: 143 (Anthrax). "Australia."
alternans Macquart, 1850: 414(110) (Anthrax). Australia
(Tas).
alternus Walker, 1849: 261 (Anthrax). "New
Holland"; Australia (Tas).
CHRYSANTHRAX Osten Sacken, 1886: 121 (as Anthrax
"group"). Type species: Anthrax fulvohirta Wiedemann,
1821 [= Anthrax cypris Meigen, 1820], des. Coquillett,
1910: 523.
cypris Meigen, 1820: 158 (Anthrax). North America [see Meigen, 1830: 325]; Hawaiian Is [introduced into quarantine; not established]; widesp. Nearctic Reg.
fulvohirta Wiedemann, 1821a: 99 [1821b: 149] (Anthrax). USA.
Undetermined sp.—Marshall Is.
EXECHOHYPOPION Evenhuis, 1991: 283. Type species: Exechohypopion
greatheadi Evenhuis, 1991, orig. des.
albatum Roberts, 1928a: 124 (Villa). Australia (WA).
bruneum Roberts, 1928a: 118 (Villa). Australia (NSW).
greatheadi Evenhuis, 1991: 286. New Caledonia (New Caledonia).
minus Macquart, 1850: 415(111) (Anthrax). Australia
(Tas); ?New Caledonia (New Caledonia).
nigricostatum Macquart, 1850: 415(111) (Anthrax).
Australia (Tas); Australia (NSW, SA, Vic).
nigricosta, error for nigricostata.
subobscurum Hardy, 1942: 203 (Anthrax; n. n. for
obscura Macquart). Australia (Tas).
obscura Macquart, 1846: 240(112) (Anthrax).
Australia (Tas). [Preocc. Weber, 1801.]
therephaga Yeates in Irwin & Yeates, 1995: 1319.
Australia (NSW).
velox White, 1916b: 211 (Anthrax). Australia (Tas).
EXHYALANTHRAX Becker, 1916a: 44 (Villa subg.). Type
species: Anthrax vagans Loew, 1862, des. Bezzi, 1924a:
7.
afer Fabricius--not Australasian/Oceanian.
ogasawarensis Matsumura, 1916: 281 (Anthrax). Bonin
Is.
LEPIDANTHRAX Osten Sacken, 1886: 107. Type species:
Anthrax disjunctus Wiedemann, 1830, des. Coquillett, 1910:
559.
albifrons Roberts, 1928a: 131. Australia (SA).
linguatus Roberts, 1928a: 130. Australia (Qld).
VILLA Lioy, 1864a: 732. Type species: Anthrax concinnus
Meigen, 1820 [= Anthrax abbadon Fabricius, 1794], des.
Coquillett, 1910: 619.
ANTHRAX, authors, not Scopoli, 1763, misid.
abeilla Hull, 1973: 374 (n. n. for antecedens Walker).
Indonesia (Sulawesi); Indonesia (Maluku); Philippines.
antecedens Walker, 1859c: 111 (Anthrax). Indonesia
(Sulawesi). [Preocc. Walker, 1852.]
apicifera Walker, 1865b: 111 (Anthrax). "New
Guinea" [= Indonesia (Irian Jaya)].
argentipennis White, 1916b: 212 (Anthrax). Australia
(Tas).
commista Macquart, 1850: 413(109) (Anthrax). Australia,
"côte orientale" [prob. = Australia (NSW)].
consimilis Thomson, 1869: 481 (Anthrax). Australia
(NSW).
flaveola Macquart, 1850: 413(109) (Anthrax). Australia,
"côte orientale" [prob. = Australia (NSW)].
fumea Roberts, 1928a: 127. Australia (NSW).
fuscicostata Macquart, 1846: 239(111) (Anthrax).
Australia (Tas); Australia (NSW, Vic).
marginata Walker, 1852: 178 (Anthrax). Australia
(Tas).
lipposa Bigot, 1892b: 353 (Glossista). Australia
(NSW).
macquarti Evenhuis, 1989: 373 (n. n. for basalis
Macquart). Australia (NSW); ?New Caledonia (New Caledonia).
basalis Macquart, 1855: 95(75) (Anthrax). Australia
(NSW). [Preocc. Walker, 1849.]
pellucida Walker, 1852: 182 (Anthrax). Australia
(Tas).
pretendens Walker, 1859c: 111 (Anthrax). Indonesia
(Sulawesi); Indonesia (Irian Jaya, Maluku), PNG (PNG).
praetendens, error for pretendens.
quinqueguttata Roberts, 1928a: 126. Australia (NSW).
rava Roberts, 1928a: 125. Australia (Qld).
resurgens Walker, 1849: 259 (Anthrax). Australia
(WA).
subannula Walker, 1849: 261 (Anthrax). Not given
[unknown].
trivincula Roberts, 1928a: 119. Australia (Qld); Australia
(NSW), Lord Howe I.
troglodyta Fabricius, 1794: 759 (Bibio). Indonesia
(Maluku); PNG (PNG).
vitrea Walker, 1852: 181 (Anthrax). Australia (WA).
zonipennis Evenhuis, 1978b: 247 (n. n. for varipennis
Roberts). Australia (NSW).
varipennis Roberts, 1928a: 121. Australia (NSW). [Preocc.
Macquart, 1850.]
tasmanica Hardy, 1924a: 76 (Anthrax).
DESMATONEURA Williston, 1895: 267. Type species: Desmatoneura
argentifrons Williston, 1895, mon.
Undetermined sp.--Australia (WA).
PETROROSSIA Bezzi, 1908a: 35. Type species: Bibio
hesperus Rossi, 1790, orig. des.
PETROSSIA, error for Petrorossia.
angustata Doleschall, 1858: 93 (Anthrax). Indonesia
(Maluku); Indonesia (Sulawesi).
fulvula Wiedemann, 1821b: 148 (Anthrax). Indonesia
(Java); ?Australia, Indonesia (Maluku), ?PNG (PNG); Indonesia
(Kalimantan, Sulawesi), Malaysia (Pen, Sabah, Sarawak), Singapore.
degenera Walker, 1856a: 15 (Anthrax). Singapore.
fulunla, error for fulvula.
oceanica Bowden, 1971d: 313. PNG (Bismarck Arch).
This page last revised 21 June 2018