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26. Family CHIRONOMIDAE

[Originally] by Peter S. Cranston & Jon Martin

Chironomidae (nonbiting midges) are distributed from the Arctic to the Antarctic and from the seas to permanent montane snowfields. Larvae are important in biomass and species diversity in aquatic biotopes, from the most pristine to the most polluted. Several species are semiterrestrial, some fully terrestrial (even in cow dung); and are among the dominant insects in the intertidal zone of marine shores, with Pontomyia truly marine. Adult midges are short-lived, but large aerial mating swarms may occur (e.g., Chironomus duplex and Polypedilum nubifer in Australia). Elsewhere such nuisance midges can cause allergic reactions in exposed humans (Cranston et al. 1984), and this has recently been reported for "Chironomus" longilobus and Tanytarsus barbitarsis at Dampier in Western Australia.

The major taxonomic works on the Chironomidae of the regions covered here are those of Freeman (1959, New Zealand; 1961, Australia) and Tokunaga (1964a, Micronesia). These works consider only adults; however, current chironomid systematics depends upon integration of morphological features from all life history stages. The great accumulation of knowledge of the metamorphosis of Northern Hemisphere species has led to a modern holomorphological generic concept strongly biased towards Holarctic taxa. We have attempted to place all species in currently accepted genera by using reared material where possible or by reexamining adults using new characters. However, in many genera apparently restricted to Australasia, or at least to the austral continents, the immature stages are still unknown. The discovery of larvae and pupae belonging to species currently known only from the adults may result in reassessment of generic classifications.

An additional source of taxonomic instability is the undoubted existence of many undescribed species and genera in the regions covered here. Virtually all regional ecological studies of aquatic biotopes, especially lotic systems, reveal a substantial number of undescribed larval chironomids. This is particularly true of western Australia (D.H. Edward, pers. comm.), Queensland, and New Zealand, where rearing to pupae and adults confirms the novelty of the fauna.

The cataloged regional fauna contains 469 valid species in 115 genera and 7 subfamilies. Worldwide there are about 5,000 described valid species of Chironomidae in 330 genera and 10 subfamilies. Several genera are included in this catalog without named species; these are mostly based on recognition of larval characters. Subfamilies are listed in postulated phylogenetic sequence. Tribes (where recognized) are listed in phylogenetic sequence.
Refs.: Freeman (1959, 1961, keys, rev., adults), Tokunaga (1964a, keys, adults), Brundin (1966, rev., esp. Podonominae), Roback (1982a,b, rev., Tanypodinae), Wiederholm (1983, keys, larval gen.), Ashe (1983, cat. world gen.), Pinder (1986, biol. rev.).

Subfamily PODONOMINAE

Genus AUSTROCHLUS Cranston

AUSTROCHLUS Cranston in Cranston, Edwards & Cook, 2002: 362. Type species: Archaeochlus brundini Cranston, Edward & Colless, 1987, orig. des.
brundini Cranston, Edward & Colless, 1987: 323 (Archaeochlus). Australia (WA).
centralaustralicus Cranston, Edward & Cook, 2002: 365. Australia (NT).
parbrundini Cranston, Edward & Cook, 2002: 365. Australia (WA).

Genus PAROCHLUS Enderlein

PAROCHLUS Enderlein, 1912e: 109. Type species: Parochlus fuegianus Enderlein, 1912, orig. des.
ZELANDOCHLUS Brundin, 1966: 106. Type species: Zelandochlus latipalpis Brundin, 1966, mon.
annulatus Brundin, 1964: 427. Nomen nudum.
aotearoae
Brundin, 1966: 138. NZ (NZ).
bassianus Brundin, 1966: 144. Australia (Tas).
     kiefferi, Freeman, 1961, not Garrett, 1925, ?misid.
brevis Sublette & Wirth, 1980: 307. NZ (Campbell I); NZ (Auckland Is).
carinatus Brundin, 1966: 151. NZ (NZ).
conjugens Brundin, 1966: 128. NZ (NZ).
glacialis Brundin, 1966: 155. NZ (NZ).
gressitti Sublette & Wirth, 1980: 306. NZ (Auckland Is).
latipalpis Brundin, 1966: 107 (Zelandochlus). NZ (NZ).
longicornis Brundin, 1966: 155. NZ (NZ).
maorii Brundin, 1966: 144. NZ (NZ).
     maorii Brundin, 1964: 431. Nomen nudum.
novaezelandiae
Brundin, 1964: 429 (as “novae zelandiae”) [1966: 138]. NZ (NZ).
ohakunensis Freeman, 1959: 409 (Podonomus). NZ (NZ).
pauperatus Brundin, 1966: 153. NZ (NZ).
reductus Sublette & Wirth, 1980: 305. NZ (Campbell I).
rennelli Sublette & Wirth, 1980: 304. NZ (Campbell I); NZ (Antipodes Is, Auckland Is).
rieki Brundin, 1966: 182. Australia (Tas).
spinosus Brundin, 1966: 138. NZ (NZ).
tasmaniae Freeman, 1961: 632 (Podonomus). Australia (Tas).
tonnoiri Brundin, 1966: 181. Australia (NSW); Australia (Tas).

Genus PODOCHLUS Brundin

PODOCHLUS Brundin, 1966: 240. Type species: Podochlus grandis Brundin, 1966, orig. des.
australiensis Brundin, 1966: 258. Australia (NSW); Australia (?Tas).
cockaynei Brundin, 1966: 252. NZ (NZ).
grandis Brundin, 1966: 248. NZ (NZ).
knoxi Brundin, 1966: 253. NZ (NZ).
stouti Brundin, 1966: 250. NZ (NZ).
tasmaniensis Brundin, 1966: 258. Australia (Tas).

Genus PODONOMOPSIS Brundin

PODONOMOPSIS Brundin, 1966: 272. Type species: Podonomus muticus Edwards, 1931, orig. des.
discoceros Brundin, 1966: 285. Australia (NSW); Australia (Tas, Vic).
     muticus, Freeman, 1961, not Edwards, 1931, ?misid.
evansi Brundin, 1966: 272. Australia (Tas); Australia (NSW, Qld).

Genus PODONOMUS Philippi

PODONOMUS Philippi, 1865: 601. Type species: Podonomus stigmaticus Philippi, 1865, mon.
collessi Brundin, 1966: 214. Australia (NSW); Australia (Tas).
derwentensis Brundin, 1966: 215. Australia (Tas).
parochloides Brundin, 1966: 199. NZ (NZ).
pygmaeus Brundin, 1966: 200. NZ (NZ).
waikukupae Brundin, 1966: 200. NZ (NZ).

Genus RHEOCHLUS Brundin

RHEOCHLUS Brundin, 1966: 269. Type species: Rheochlus insignis Brundin, 1966, orig. des.
wirthi Freeman, 1961: 632 (Podonomus). Australia (NSW).

Subfamily APHROTENINAE

Genus APHROTENIA Brundin

APHROTENIA Brundin, 1966: 338. Type species: Aphrotenia tsitsikamae Brundin, 1966, orig. des.
australiensis Cranston & Edward, 1992: 50. Australia (SA).
     australiensis Hergstrom, 1974: 68. Nomen nudum.

Genus APHROTENIELLA Brundin

APHROTENIELLA Brundin, 1966: 348. Type species: Aphroteniella filicornis Brundin, 1966, orig. des.
ANAPHROTENIA Brundin, 1983: 423. Type species: Anaphrotenia lacustris Brundin, 1983, orig. des.
filicornis Brundin, 1966: 348. Australia (Qld); Australia (NSW, NT, Vic, WA).
     lacustris Brundin, 1983: 427 (Anaphrotenia). Australia (Qld).
tenuicornis Brundin, 1966: 352. Australia (NSW); Australia (ACT, Vic, WA).

Genus PARAPHROTENIA Brundin

PARAPHROTENIA Brundin, 1966: 352. Type species: Paraphrotenia excellens Brundin, 1966, orig. des.
fascipennis Brundin, 1966: 358. Australia (NSW); Australia (Vic).

Subfamily TANYPODINAE
Tribe TANYPODINI

Genus TANYPUS Meigen

TANYPUS Meigen, 1803: 261. Type species: Tipula cincta Fabricius, 1775 [misid., = Tanypus punctipennis Meigen, 1818], des. Latreille, 1810: 442.
Unidentified sp.--Australia (NT) (Roback, 1982b: 81).

Tribe COELOTANYPODINI

Genus COELOTANYPUS Kieffer

COELOTANYPUS Kieffer, 1913d: 154. Type species: Tanypus humeralis Loew, 1866, orig. des.
wirthi Freeman, 1961: 624. Australia (NSW).

Genus CLINOTANYPUS Kieffer

CLINOTANYPUS Kieffer, 1913d: 157. Type species: Procladius fuscosignatus Kieffer, 1910, des. Sublette & Sublette, 1965: 145.
crux Wiedemann, 1824: 10 (Tanypus). "India or."; Australia (NSW, NT, Qld); Oriental Reg.
     australiensis Kieffer, 1926: 104. Australia (Qld).
guamensis Tokunaga, 1964a: 492. Guam.

Tribe COELOPYNIINI

Genus COELOPYNIA Freeman

COELOPYNIA Freeman, 1961: 626. Type species: Coelopynia pruinosa Freeman, 1961, orig. des.
pruinosa Freeman, 1961: 626. Australia (Vic); widesp. Australia (excl. SA).

Tribe MACROPELOPINI

Genus APSECTROTANYPUS Fittkau

APSECTROTANYPUS Fittkau, 1962: 141. Type species: Tanypus trifascipennis Zetterstedt, 1838, orig. des.
maculosus Freeman, 1961: 622 (Anatopynia). Australia (Tas); Australia (NSW, Vic).

Genus FITTKAUIMYIA Karunakaran

FITTKAUIMYIA Karunakaran, 1969: 75. Type species: Fittkauimyia disparipes Karunakaran, 1969, orig. des.
disparipes Karunakaran, 1969: 75. Singapore; Australia (NT).

Genus GRESSITTIUS Sublette & Wirth

GRESSITTIUS Sublette & Wirth, 1980: 302. Type species: Corethra antarctica Hudson, 1892, orig. des.
antarcticus Hudson, 1892: 43 (Corethra). NZ (NZ); NZ (Antipodes Is, Auckland Is).
     hudsoni Kieffer, 1922a: 148 (Macropelopia). NZ (NZ).
umbrosus Freeman, 1959: 406 (Anatopynia). NZ (NZ).

Genus MACROPELOPIA Thienemann

MACROPELOPIA Thienemann, 1916: 497. Type species: Isoplastus bimaculata Kieffer, 1909 (as Isoplasus) [= Tanypus nebulosus Meigen, 1804], orig. des.

Subgenus ALOTANYPUS Roback

ALOTANYPUS Roback, 1971: 95 (as gen.). Type species: Tanypus venustus Coquillett, 1902, orig. des.
dalyupensis Freeman, 1961: 624 (Anatopynia). Australia (WA); Australia (SA).
vittiger Edwards, 1931: 242 (Anatopnyia). Argentina; Juan Fernandez Is; Chile.

Genus MONOPELOPIA Fittkau

MONOPELOPIA Fittkau, 1962: 34. Tanypus tenuicolor Kieffer, 1918, orig. des.
sp. — NZ(NZ) [Blakely, Cranson & Winterbourn, 2010: 39].


Unplaced species of Macropelopini

apicincta Freeman, 1959: 403 (Anatopynia). NZ (NZ).
apicinella Freeman, 1959: 406 (Anatopynia). NZ (NZ).
boninensis Tokunaga, 1964a: 501 (Anatopynia). NZ (NZ).
cana Freeman, 1959: 408 (Anatopynia). NZ (NZ).
debilis Hutton, 1902b: 186 (Tanypus). NZ (NZ).
     novaezelandiae Kieffer, 1922a: 147 (Macropelopia). NZ (NZ).
elongata Tokunaga, 1964a: 501 (Anatopynia). Northern Marianas; Micronesia.
flavipes Freeman, 1959: 405 (Anatopynia). NZ (NZ).
languidus Hutton, 1902b: 186 (Tanypus). NZ (NZ).
masteri Skuse, 1889e: 278 (Tanypus). Australia (NSW); Australia (ACT).
ogasasextdecimaSasa & Suzuki, 1997: 342. Bonin Is.
pallipes Freeman, 1961: 623 (Anatopynia). Australia (Tas); Australia (NSW).
pennipes Freeman, 1961: 623 (Anatopynia). NZ (NZ).
quadricincta Freeman, 1959: 407 (Anatopynia). NZ (NZ).
quinquepunctata Freeman, 1959: 405 (Anatopynia). NZ (NZ).

Tribe PROCLADIINI

Genus DJALMABATISTA Fittkau

DJALMABATISTA Fittkau, 1968: 328. Type species: Djalmabatista director Fittkau, 1968, orig. des.
DJALMABASTICA, error for Djalmabatista.
Undetermined sp.--Australia (NSW, NT).

Genus PROCLADIUS Skuse

PROCLADIUS Skuse, 1889e: 283. Type species: Procladius paludicola Skuse, 1889, des. Coquillett, 1910: 594.
goanna Roback, 1982b: 94. Australia (NT).
martini Roback, 1982b: 97. Australia (Vic): Australia (NSW).
paludicola Skuse, 1889e: 284. Australia (NSW); widesp. Australia.
     pentastictus Kieffer, 1917a: 201. Australia (NSW).
     pictipennis Skuse, 1889e: 284. Australia (NSW).
     subglaber Kieffer, 1917a: 202 (Trichotanypus). Australia (NSW).
villosimanus Kieffer, 1917a: 202. Australia (NSW); widesp. Australia (excl. Qld).
     australiensis Kieffer, 1917a: 200. Australia (NSW).

Unplaced species of Procladius Skuse

squamifer Freeman, 1961: 629. Australia (NSW); Australia (ACT, Tas, Vic).

Tribe PENTANEURINI

Genus ABLABESMYIA Johannsen

ABLABESMYIA Johannsen, 1905: 135. Type species: Tipula monilis Linnaeus, 1758, des. Johannsen, 1907: 400.
atromaculata Edwards, 1928c: 59. Western Samoa.
hilli Freeman, 1961: 620. Australia (Qld); Australia (Vic, WA), PNG (PNG), Western Samoa.
mala Hutton, 1902b: 187 (Tanypus). NZ (NZ); NZ (Campbell I).
monilis Linnaeus, 1758: 587 (Tipula). "Europa"; Bonin Is, Micronesia; Holarctic, Oriental Regs.
notabilis Skuse, 1889e: 280 (Isoplastus). Australia (NSW); widesp. Australia (excl. NT), Western Samoa.
     monilis, Kieffer, 1917, not Linnaeus, 1758, misid.


Genus AUSTRALOPELOPIA Cranston

AUSTRALOPELOPIA Cranston, 2000: 116. Type species: Australopelopia prionoptera Cranston, 2000, orig. des.
prionoptera Cranston, 2000: 118. Australia (Qld).

Genus LARSIA Fittkau

LARSIA Fittkau, 1962: 339. Type species: Ablabesmyia atrocincta Goetghebuer, 1942, orig. des.
planensis Johannsen, 1946: 284 (Pentaneura). Canada; Hawaiian Is; Nearctic Reg

Genus NILOTANYPUS Kieffer

NILOTANYPUS Kieffer, 1923: 191. Type species: Nilotanypus remotissimus Kieffer, 1923, mon.
Unnamed sp.--Australia (NSW, NT, Qld).

Genus PARAMERINA Fittkau

PARAMERINA Fittkau, 1962: 317. Type species: Paramerina cingulata Walker, 1856, orig. des.
dolosa Johannsen, 1932b: 500 (Pentaneura). Indonesia (Java); Micronesia.
ignobilis Johannsen, 1932b: 496 (Pentaneura). Indonesia (Sumatra); Micronesia.
levidensis Skuse, 1889e: 281 (Isoplastus). Australia (NSW); widesp. Australia (excl. NT).
parva Freeman, 1961: 617 (Pentaneura). Australia (Qld); Australia (NT).
taylori Roback, 1982b: 102. Australia (Qld).

Genus ZAVRELIMYIA Fittkau

ZAVRELIMYIA Fittkau, 1962: 285. Type species: Tanypus melanurus Meigen, 1804, orig. des.
harrisi Freeman, 1959: 400 (Pentaneura). NZ (NZ).

Unplaced species of Pentaneurini

carolinensis Tokunaga, 1964a: 498 (Pentaneura). Guam; Micronesia.
formulosus Skuse, 1889e: 282 (Isoplastus). Australia (NSW).
ponapensis Tokunaga, 1964a: 495 (Pentaneura). Micronesia.
punctipennis Freeman, 1961: 618 (Pentaneura). Australia (ACT); Australia (NSW, Tas, Vic).

Subfamily DIAMESINAE
Tribe HEPTAGYINI

Genus PARAHEPTAGYIA Brundin

PARAHEPTAGYIA Brundin, 1966: 379. Type species: Heptagyia cinerascens Edwards, 1931, orig. des.
tasmaniae Freeman, 1961: 635 (Heptagyia). Australia (Tas).
tonnoiri Freeman, 1961: 636 (Heptagyia). Australia (ACT); Australia (NSW).

Tribe LOBODIAMESINI

Genus LOBODIAMESA Pagast

LOBODIAMESA Pagast, 1947: 446. Type species: Lobodiamesa campbelli Pagast, 1947, mon.
campbelli Pagast, 1947: 446. NZ (NZ).

Genus MAORIDIAMESA Pagast

MAORIDIAMESA Pagast, 1947: 448. Type species: Maoridiamesa harrisi Pagast, 1947, mon.
glacialis Brundin, 1966: 414. NZ (NZ).
harrisi Pagast, 1947: 448. NZ (NZ).
insularis Brundin, 1966: 414. NZ (Campbell I).
intermedia Brundin, 1966: 411. NZ (NZ).
stouti Brundin, 1966: 412. NZ (NZ).

Subfamily TELMATOGETONINAE

Genus TELMATOGETON Schiner

TELMATOGETON Schiner, 1866c: 931 [1868: 25]. Type species: Telmatogeton sanctipauli Schiner, 1866 (as St. Pauli), mon.
PARACLUNIO Kieffer, 1911f: 103. Type species: Paraclunio trilobatus Kieffer, 1911, orig. des.
CHARADROMYIA Terry, 1913: 292. Type species: Charadromyia torrenticola Terry, 1913, orig. des.
abnormis Terry, 1913: 295 (Charadromyia). Hawaiian Is.
antipodensis Sublette & Wirth, 1980: 310. NZ (Antipodes Is).
australicus Womersley, 1936: 437. Australia (SA); Australia (Vic).
fluviatilis Wirth, 1947b: 166. Hawaiian Is.
hirtus Wirth, 1947b: 158. Hawaiian Is.
japonicus Tokunaga, 1933a: 95. Japan; Australia (NSW, SA, Vic, WA), Hawaiian Is; Holarctic, Oriental Regs.
macquariensis Brundin, 1962: 945 (Halirytus). Macquarie I.
mortoni Leader, 1975: 209. NZ (NZ).
pacificus Tokunaga, 1935: 15. Japan; Hawaiian Is.
pusillum Edwards, 1933b: 88. French Polynesia (Marquesas); Northern Marianas.
torrenticola Terry, 1913: 292 (Charadromyia). Hawaiian Is.
williamsi Wirth, 1947b: 162. Hawaiian Is.

Genus THALASSOMYA Schiner

THALASSOMYA Schiner, 1856: 219. Type species: Thalassomya frauenfeldi Schiner, 1856, mon.
THALASSOMYIA, error for Thalassomya.
africana Edwards, 1926c: 787. Tanzania; French Polynesia (Marquesas).
maritima Wirth, 1947a: 131. Hong Kong; Belau, Guam, Marshall Is, New Caledonia (New Caledonia), Northern Marianas, Western Samoa.
pilipes Edwards, 1933b: 110. American Samoa; French Polynesia (Marquesas), Vanuatu; Hong Kong.
sabroskyi Tokunaga, 1964a: 542. Belau; Micronesia.
setosipennis Wirth, 1947a: 121. Hawaiian Is.

Subfamily ORTHOCLADIINAE

Genus ALLOMETRIOCNEMUS Freeman

ALLOMETRIOCNEMUS Freeman, 1961: 657. Type species: Allometriocnemus coloensis Freeman, 1961, orig. des.
coloensis Freeman, 1961: 659. Australia (NSW).
pictus Freeman, 1961: 659. Australia (NSW).

Genus ALLOTRISSOCLADIUS Freeman

ALLOTRISSOCLADIUS Freeman, 1964: 148. Type species: Allotrissocladius amphibius Freeman, 1964, orig. des.
amphibius Freeman, 1964: 149. Australia (WA).


Genus ANZACLADIUS Cranston

ANZACLADIUS Cranston, 2009: 131. type species: Anzacladius numbat Cranston, 2009, orig. des.
kangaroo Cranston, 2009: 136. Australia (SA).
kiwi Cranston, 2009: 136. NZ (NZ).
numbat Cranston, 2009: 136. Australia (WA).

Genus AUSTROBRILLIA Freeman

AUSTROBRILLIA Freeman, 1961: 640. Type species: Austrobrillia longipes Freeman, 1961, mon. [Freeman, 1961: 640 designated Austrobrillia nudipennis as "type and only known species." A. nudipennis is a nomen nudum and error for longipes, the only included species.]
longipes Freeman, 1961: 640. Australia (Tas); Australia (Vic).
     nudipennis, error for longipes.

Genus AUSTROCLADIUS Freeman

AUSTROCLADIUS Freeman, 1961: 647. Type species: Camptocladius terjugus Skuse, 1889, orig. des.
harrisi Freeman, 1959: 418 (Chaetocladius). NZ (NZ).
numerosus Skuse, 1889e: 256 (Orthocladius). Australia (NSW); Australia (ACT, Tas, Vic).
terjugus Skuse, 1889e: 262 (Camptocladius). Australia (NSW); Australia (ACT, Vic).
trichiatus Freeman, 1961: 649. Australia (Vic).


Genus BOTRYOCLADIUS Cranston & Edward

BOTRYOCLADIUS Cranston & Edward, 1999: 307. Type species: Botryocladius grapeth Cranston & Edward, 1999, orig. des.
australoalpinus Cranston & Edward, 1999: 324. Australia (Tas).
bibulmun Cranston & Edward, 1999: 325. Australia (WA).
brindabella Cranston & Edward, 1999: 323. Australia (ACT).
collessi Cranston & Edward, 1999: 324. Australia (NSW).
freemani Cranston & Edward, 1999: 326. Australia (WA); Australia (ACT, NSW).
grapeth Cranston & Edward, 1999: 321. Australia (NSW); Australia (ACT, Qld, SA, Tas, Vic, WA).
mdfrc Cranston & Edward, 1999: 323. Australia (Vic).
petrophilus Cranston & Edward, 1999: 325. Australia (NT).
tasmania Cranston & Edward, 1999: 324. Australia (Tas).


Genus BRILLIA Kieffer

BRILLIA Kieffer, 1913: 34. Type species: Metriocnemus bifidus Kieffer, 1913 [= Chironomus modesta Meiogen, 1830], orig. des.
ogasaquinta Sasa & Suzuki, 1997: 324. Bonin Is.

Genus CAMPTOCLADIUS Wulp

CAMPTOCLADIUS Wulp, 1874: 133. Type species: Tipula byssina Schrank, 1803 [= Tipula stercoraria De Geer, 1776], des. Coquillett, 1910: 518.
stercorarius De Geer, 1776: 388 (Tipula). Sweden; Australia (NSW), NZ (NZ); Holarctic Reg.

Genus CARDIOCLADIUS Kieffer

CARDIOCLADIUS Kieffer, 1912a: 22. Type species: Cardiocladius ceylanicus Kieffer, 1912, mon.
australiensis Freeman, 1961: 639. Australia (Tas); Australia (ACT, NSW).

Genus CLUNIO Haliday

CLUNIO Haliday, 1855: 62. Type species: Clunio marinus Haliday, 1855, mon.
CLUNEO, error for Clunio.
brevis Stone & Wirth, 1947: 212. Hawaiian Is.
littoralis Stone & Wirth, 1947: 203. Hawaiian Is.
martini Hashimoto, 1973: 1. Australia (Vic).
pacificus Edwards, 1926c: 790. Western Samoa; American Samoa, Australia (NSW), Belau, Northern Marianas, Vanuatu; Ryukyus.
tsushimensis Tokunaga, 1933a: 92. Japan; Hawaiian Is.
tuthilli Tokunaga, 1964a: 537. Marshall Is.
vagans Stone & Wirth, 1947: 206. Hawaiian Is.

Genus COMPTEROSMITTIA Saether

COMPTEROSMITTIA Saether, 1981: 20. Type species: Compterosmittia dentispina Saether, 1981, orig. des.
claggi Tokunaga, 1964a: 506 (Metriocnemus). Bonin Is; Belau, Micronesia.
longipalpis Ueno, Satake & Yamano, 2015: 71. Tuvalu.
pectinata Freeman, 1961: 662 (Gymnometriocnemus). Australia (NSW).
tuberculifera Tokunaga, 1964a: 530 (Smittia). Micronesia; Belau, Bonin Is, Guam.

Genus CORYNONEURA Winnertz

CORYNONEURA Winnertz, 1846: 12. Type species: Corynoneura scutellata Winnertz, 1846, des. Coquillett, 1910: 528.
australiensis Freeman, 1961: 673. Australia (ACT).
imperfecta Skuse, 1889e: 307 (Ceratopogon). Australia (NSW).
scutellata Winnertz, 1846: 13. Germany; Australia (Vic, WA), ?NZ (NZ); widesp. Holarctic, Neotrop. Regs.
     ?donovani Forsyth, 1971: 121. NZ (NZ).

Genus CRICOTOPUS Wulp

CRICOTOPUS Wulp, 1874: 132. Type species: Chironomus tibialis Meigen, 1804, des. Coquillett, 1910: 528.

Subgenus CRICOTOPUS Wulp

CRICOTOPUS Wulp, 1874: 132 (as gen.). Type species: Chironomus tibialis Meigen, 1804, des. Coquillett, 1910: 528.
acornis Cranston & Krosch, 2015: 298. Austrlaia (NSW).
albitarsis Drayson, Cranston & Krosch, 2015: 6. Australia (SA); Australia (ACT, NSW, Qld, NT, Vic, WA).
albitibia Walker--not Australasian/Oceanian.
annuliventris Skuse, 1889e: 255 (Orthocladius). Australia (NSW); Australia (ACT, SA, Tas, Vic).
aucklandensis Sublette & Wirth, 1980: 313. NZ (Auckland Is); NZ (NZ).
bicinctus Meigen, 1818: 41 (Chironomus). Germany; Hawaiian Is; widesp. Afrotrop., Holarctic (incl. Japan) Regs.
brevicornis Drayson & Cranston in Drayson, Cranston & Krosch, 2015: 10. Australia (NSW); Australia (NT, Qld).
cingulatus Hutton, 1902b: 184 (Orthocladius). NZ (NZ).
conicornis Drayson & Cranston in Drayson, Cranston & Krosch, 2015: 11. Australia (ACT); Australia (NSW, Qld).
gressitti
Tokunaga, 1964a: 510. Micronesia; PNG (PNG).
hillmani Drayson & Cranston in Drayson, Cranston & Krosch, 2015: 13. Australia (NSW); Australia (ACT, Qld, Tas).
howensis
Cranston & Krosch, 2015: 298. Lord Howe I.
planus
Boothroyd, 1990: 427. NZ (NZ).
ogasaseptimus Sasa & Suzuki, 1997: 326. Bonin Is.
ogasasextus Sasa & Suzuki, 1997: 325. Bonin Is.
parbicinctus Drayson, Cranston & Krosch, 2015: 16. Australia (SA).; Australia (ACT, NSW, Qld, SA, WA).
quadrizonatus
Tokunaga, 1964a: 512. Bonin Is.
sabroskyi Tokunaga, 1964a: 511. Belau.
tasmania Drayson & Cranston in Drayson, Cranston & Krosch, 2015: 18. Australia (Tas).
varicornis Cranston & Krosch, 2015: 299. Austraia (ACT).
vicenti Boothroyd, 1990: 421. NZ (NZ).
wangi Cranston & Krosch, 2015: 299. Austraia (NT).
zealandicus Freeman, 1959: 413. NZ (NZ).

Subgenus ISOCLADIUS Kieffer

ISOCLADIUS Kieffer, 1909: 44. Type species: Isocladius albipes Kieffer, 1909 [= Tipula silvestris Fabricius, 1794], orig. des.
Unidentified sp.—Australia (Vic).

Genus DOLOPLASTUS Skuse

DOLOPLASTUS Skuse, 1889e: 260. Type species: Doloplastus monticola Skuse, 1889, orig. des.
monticola Skuse, 1889e: 260. Australia (Vic).


Genus ECHINOCLADIUS Cranston

ECHINOCLADIUS Cranston, 2000: 109. Type species: Echinocladius martini Cranston, 2000, orig. des.
martini Cranston, 2000: 113. Australia (ACT).

Genus EUKIEFFERIELLA Thienemann

EUKIEFFERIELLA Thienemann, 1926: 325. Type species: Dactylocladius longicalcar Kieffer, 1911 [misid., = Spaniotoma gracei Edwards, 1929], orig. des.
heveli Sublette & Wirth, 1980: 312. NZ (Campbell I); NZ (Auckland Is).
insolida Skuse, 1889e: 258 (Orthocladius). Australia (NSW).
latifurca Freeman, 1961: 670 (Pseudosmittia). Australia (NSW).
ogasaoctava Sasa & Suzuki, 1997: 329. Bonin Is.

Genus GYMNOMETRIOCNEMUS Goetghebuer

GYMNOMETRIOCNEMUS Goetghebuer, 1932: 23 (Metriocnemus subg.). Type species: Metriocnemus subnudus Edwards, 1929, des. Edwards, 1932a: 141. [Unavailable name; genus-group name proposed after 1930 without type-species designation; authors prefer to retain current authorship and date pending ruling by I.C.Z.N.]
lobifer Freeman, 1959: 412 (Metriocnemus). NZ (NZ).
nitidulus Skuse, 1889e: 276 (Metriocnemus). Australia (NSW).
wilsoni Freeman, 1961: 661. Australia (Vic).

Genus GYNNIDOCLADIUS Sublette & Wirth

GYNNIDOCLADIUS Sublette & Wirth, 1980: 322. Type species: Gynnidocladius pilulus Sublette & Wirth, 1980, orig. des.
pilulus Sublette & Wirth, 1980: 322. NZ (Campbell I).


Genus HAHAYUSURIKA Sasa & Suzuki

HAHAYUSURIKA Sasa & Suzuki, 1997: 332. Type species: Hahayusurika ogasaundecima Sasa & Suzuki, 1997, orig. des.
ogasaundecima Sasa & Suzuki, 1997: 332. Bonin Is.

Genus HEVELIUS Sublette & Wirth

HEVELIUS Sublette & Wirth, 1980: 320. Type species: Hevelius carinatus Sublette & Wirth, 1980, orig. des.
carinatus Sublette & Wirth, 1980: 321. NZ (Campbell I); NZ (Auckland Is).


Genus KANIWHANIWHANUS Boothroyd

KANIWHANIWHANUS Boothroyd, 1999: 342. Type species: Kaniwhaniwhanus chapmani Boothroyd, 1999, orig. des.
chapmani Boothroyd, 1999: 345. NZ (NZ).

Genus KIEFFEROPHYES Freeman

KIEFFEROPHYES Freeman, 1961: 663. Type species: Camptocladius invenustulus Skuse, 1889, orig. des.
invenustulus Skuse, 1889e: 265 (Camptocladius). Australia (NSW); Australia (Tas, Vic).
leei Freeman, 1961: 665. Australia (NSW).
lobifer Freeman, 1961: 664. Australia (Vic).

Genus KUSCHELIUS Sublette & Wirth

KUSCHELIUS Sublette & Wirth, 1980: 314. Type species: Kuschelius dentifer Sublette & Wirth, 1980, orig. des.
dentifer Sublette & Wirth, 1980: 315. NZ (Auckland Is).

Genus LIMNOPHYES Eaton

LIMNOPHYES Eaton, 1875: 60. Type species: Limnophyes pusillus Eaton, 1875 [= Chironomus minimus Meigen, 1818], mon.
vestitus Skuse, 1889e: 263 (Camptocladius). Australia (NSW); Australia (WA).

Genus LJUNGNERIA Brundin

LJUNGNERIA Brundin, 1964: 430. Nomen nudum; genus-group name proposed after 1930 without designation of type species from two included species.
maorii Brundin, 1964: 431. NZ (NZ).

Genus MARYELLA Sublette & Wirth

MARYELLA Sublette & Wirth, 1980: 325. Type species: Maryella reducta Sublette & Wirth, 1980, orig. des.
reducta Sublette & Wirth, 1980: 126. NZ (Auckland Is).

Genus MECAORUS Sublette & Wirth

MECAORUS Sublette & Wirth, 1980: 316. Type species: Mecaorus elongatus Sublette & Wirth, 1980, orig. des.
elongatus Sublette & Wirth, 1980: 316. NZ (Auckland Is).

Genus METRIOCNEMUS Wulp

METRIOCNEMUS Wulp, 1874: 136. Type species: Chironomus albolineatus Meigen, 1818, des. Coquillett, 1910: 569.
hornsbyensis Freeman, 1961: 657. Australia (NSW).
ogasadecimus Sasa & Suzuki, 1997: 331. Bonin Is.

Genus NAKATAIA Sublette & Wirth

NAKATAIA Sublette & Wirth, 1980: 324. Type species: Nakataia cisdentifer Sublette & Wirth, 1980, orig. des.
cisdentifer Sublette & Wirth, 1980: 324. NZ (Auckland Is).

Genus NANOCLADIUS Kieffer

NANOCLADIUS Kieffer, 1913b: 31. Type species: Nanocladius vitellinus Kieffer, 1913, orig. des.
Undescribed spp.--widesp. Australia, Micronesia.


Genus NAONELLA Boothroyd

NAONELLA Boothroyd, 1994: 309. Type species: Naonella forsythi Boothroyd, 1994, orig. des.
forsythi Boothroyd, 1994: 311. NZ (NZ).
kimihia Boothroyd, 2005: 12. NZ (NZ).

Genus NASUTICLADIUS Freeman

NASUTICLADIUS Freeman, 1961: 650. Type species: Nasuticladius tonnoiri Freeman, 1961, orig. des.
ater Freeman, 1961: 651. Australia (Tas).
niger Freeman, 1961: 651. Australia (Tas).
tonnoiri Freeman, 1961: 650. Australia (Tas).
wilsoni Freeman, 1961: 654. Australia (Vic).
wirthi Freeman, 1961: 651. Australia (NSW).

Genus NESIOCLADIUS Sublette & Wirth

NESIOCLADIUS Sublette & Wirth, 1980: 323. Type species: Nesiocladius gressitti Sublette & Wirth, 1980, orig. des.
gressitti Sublette & Wirth, 1980: 323. NZ (Auckland Is).

Genus ORTHOCLADIUS Wulp

ORTHOCLADIUS Wulp, 1874: 132. Type species: Tipula stercoraria De Geer, 1776 [misid., = Chironomus oblidens Walker, 1856], des. Coquillett, 1910: 581.
pictipennis Freeman, 1959: 419. NZ (NZ).

Genus PARAKIEFFERIELLA Thienemann

PARAKIEFFERIELLA Thienemann, 1936: 195. Type species: Spaniotoma coronata Edwards, 1929, orig. des.
Unidentified spp.--Australia (ACT, SA, Vic), Hawaiian Is.

Genus PARALIMNOPHYES Brundin

PARALIMNOPHYES Brundin, 1966: 129. Type species: Camptocladius hydrophilus Goetghebuer, 1921, orig. des.
albibasis Freeman, 1961: 656 (Limnophyes). Australia (NSW); Australia (ACT, Tas).
pullulus Skuse, 1889e: 259 (Orthocladius). Australia (NSW); Australia (Vic, WA).

Genus PARAMETRIOCNEMUS Goetghebuer

PARAMETRIOCNEMUS Goetghebuer, 1932: 22 (Metriocnemus subg.). Type species: Metriocnemus stylatus Kieffer, 1924, mon.
claggi Tokunaga, 1964a: 563 (Chironomus). Bonin Is; Guam, Micronesia, Wake I.
flavellus Tokunaga, 1964a: 502 (Metriocnemus). Micronesia.
ornaticornis Kieffer, 1917a: 225 (Metriocnemus). Australia (NSW); Australia (ACT, WA).

Genus PARATRICHOCLADIUS Santos Abreu

PARATRICHOCLADIUS Santos Abreu, 1918: 204. Type species: Orthocladius tenuipes Becker, 1908 [= Chironomus rufiventris Meigen, 1830], des. Hirvenoja, 1973: 88.
SYNCRICOTOPUS Brundin, 1956: 106. Type species: Chironomus rufiventris Meigen, 1830, orig. des.
TRICHOCLADIUS, Freeman, 1959, not Kieffer, 1906, ?misid.
pluriserialis Freeman, 1959: 414 (Trichocladius). NZ (NZ); Australia (ACT, NSW, Vic).

Genus PARATRISSOCLADIUS Zavrel

PARATRISSOCLADIUS Zavrel, 1937: 10. Type species: Trissocladius fluviatilis Goetghebeur, 1937 [= Chironomus exerptus Walker, 1856], orig. des.
ogasaduodecimus Sasa & Suzuki, 1997: 334. Bonin Is.


Genus PIRARA Boothroyd & Cranston

PIRARA Boothroyd & Cranston, 1995: 391. Type species: Pirara matakiri Boothroyd & Cranston, 1995, orig. des.
australiensis Boothroyd & Cranston, 1995: 397. NZ (NZ).
matakiri Boothroyd & Cranston, 1995: 396. Australia (NSW).

Genus PSEUDOSMITTIA Goetghebuer

PSEUDOSMITTIA Goetghebuer, 1932: 126 (Smittia subg.). Type species: Spaniotoma angusta Edwards, 1929, des. Edwards, 1932a: 141. [Unavailable name; genus-group name proposed after 1930 without type-species designation; current authorship and date retained here pending ruling by I.C.Z.N.]
amplexivirga Ueno, Satake & Yamano, 2015: 75.Tuvalu.
bicinctura
Tokunaga, 1964a: 533 (Smittia). Belau.
bifurcata Tokunaga, 1936: 310 (Spaniotoma). Japan; Bonin Is, Kiribati.
     ogasatridecima Sasa & Suzuki, 1997: 335. Bonin Is.
brachydicrana
Edwards, 1927b: 243 (Orthocladius). French Polynesia (Society Is); Belau, Marshall is, Micronesia.
     micronesiana Tokunaga, 1964a: 526 (Smittia). Micronesia.
     setiforceps Tokunaga, 1964a: 519 (Smittia). Belau.
     micronesia, error for micronesiana (Cranston & Martin, 1989: 262).
     brachidicranus, error for brachydicrana.
christmasensis Ferrington & Saether, 2011: 203. Kiribati.
dupla Tokunaga, 1964a: 522 (Smittia). Marshall Is; Belau.
fusivenosa Tokunaga, 1964a: 534 (Smittia). Micronesia.
insulsa Johannsen, 1946: 191 (Smittia). Guam; Belau, Northern Marianas.
kauaiensis Hardy, 1960: 139 (Orthocladius). Hawaiian Is.
kraussi Tokunaga, 1964a: 523 (Smittia). Belau; Micronesia.
licina Ferrington & Saether, 2011: 201. Kiribati.
macrobrachia Edwards, 1928c: 62 (Orthocladius). Western Samoa.
mauiensis Hardy, 1960: 141 (Orthocladius). Hawaiian Is.
oahuensis Hardy, 1960: 142 (Orthocladius). Hawaiian Is.
palauensis Tokunaga, 1964a: 520 (Smittia). Belau.
paraconjuncta Hardy, 1960: 143 (Orthocladius). Hawaiian Is.
tericristata Ferrington & Saether, 2011: 171. Australia (WA).
tokunagi Ferrington & Saether, 2011: 232. Belau.
triangula Tokunaga, 1964a: 524 (Smittia). Belau; Marshall Is.
tuvaluensis Ueno, Satake & Yamano, 2015: 73.Tuvalu.
yapensis Tokunaga, 1964a: 525 (Smittia). Micronesia; Marshall Is.
zonata Tokunaga, 1964a: 528 (Smittia). Belau; Guam, Northern Marianas, Micronesia.
     guamensis Tokunaga, 1964a: 532 (Smittia). Guam.

Genus PTEROSIS Sublette & Wirth

PTEROSIS Sublette & Wirth, 1980: 317. Type species: Pterosis wisei Sublette & Wirth, 1980, orig. des.
wisei Sublette & Wirth, 1980: 317. NZ (Auckland Is); NZ (Campbell I).


Genus RHAGOSMITTIA Ferrington & Saether

RHAGOSMITTIA Ferrington & Saether, 2006: 244. Type species: Spaniotoma maculiventris Edwards, 1935, org. des.
maculiventris Edwards, 1933b: 89 (Spaniotoma). French Polynesia (Marquesas); Belau, Hawaiian Is, Western Samoa.
     postcinctura Tokunaga, 1964a: 533 (Smittia). Belau.


Genus RHEOCRICOTOPUS Thienemann & Harnish

RHEOCRICOTOPUS Thienemann & Harnish, 1932: 135. Type species: Chironomus effusus Walker, 1856, des. Brundin, 1956: 118. [Unavailable name; genus-group name proposed after 1930 without type-species designation; authors prefer to retain current authorship and date pending ruling by I.C.Z.N.]
Undetermined sp.--Australia (NSW, NT, Vic).

Genus RHINOCLADIUS Edwards

RHINOCLADIUS Edwards, 1931c: 268. Type species: Rhinocladius culicinus Edwards, 1931, orig. des.
tonnoiri Freeman, 1961: 662. Australia (Tas); Australia (Vic).

Genus SEMIOCLADIUS Sublette & Wirth

SEMIOCLADIUS Sublette & Wirth, 1980: 318. Type species: Camptocladius crassipennis Skuse, 1889, orig. des.
brevicornis Tokunaga, 1964a: 518 (Smittia). Belau; Micronesia.
crassipennis Skuse, 1889e: 264 (Camptocladius). Australia (NSW).
kuscheli Sublette & Wirth, 1980: 319. NZ (Auckland Is); NZ (Antipodes Is, Campbell I).
reinga Leader, 1975: 211 (Smittia). NZ (NZ).
whangaroa Leader, 1975: 213 (Smittia). NZ (NZ).

Genus SMITTIA Holmgren

SMITTIA Holmgren, 1869: 47. Type species: Chironomus brevipennis Boheman, 1866, mon.
ORTHOSMITTIA Goetghebuer, 1943: 110 (Smittia subg.). Type species: Orthocladius brevifurcatus Edwards, 1926, orig. des.
aterrima Meigen, 1818: 47 (Chironomus). Germany; Australia (ACT, NSW, Qld, Tas, WA); Holarctic Reg. (incl. Japan).
     macleayi Skuse, 1889e: 266 (Camptocladius). Australia (NSW).
retracta Freeman, 1961: 667. Australia (NSW).
reyei Freeman, 1961: 671 (Orthosmittia). Australia (Qld).
verna Hutton, 1902b: 185 (Camptocladius). NZ (NZ).
wirthi Freeman, 1961: 666. Australia (NSW).

Genus STICTOCLADIUS Edwards

STICTOCLADIUS Edwards, 1931c: 279 (Spaniotoma subg.). Type species: Spaniotoma pulchripennis Edwards, 1931, orig. des.
lacuniferus Freeman, 1959: 416 (Diplocladius). NZ (NZ).
multiserialis Freeman, 1961: 641. Australia (NSW); Australia (Tas).
occidentalis Cranston & Saether, 2010: 139. Australia (WA).
pictus Freeman, 1959: 417 (Diplocladius). NZ (NZ).
sofour Cranston & Saether, 2010: 139. Australia (ACT).
uniserialis Freeman, 1961: 643. Australia (Tas); Australia (ACT, NSW, Vic, WA).
victoriensis Freeman, 1961: 643. Australia (Vic); Australia (ACT, NSW, Vic).

Genus SYMBIOCLADIUS Kieffer

SYMBIOCLADIUS Kieffer, 1925: 565. Type species: Phaenocladius rhitrogenae Zavrel, 1924, orig. des.

Subgenus ACLETIUS Roback

ACLETIUS Roback, 1965: 115. Type species: Symbiocladius wygodzinskyi Roback, 1965, orig. des.
aurifodinae Hynes, 1976: 49. Australia (Vic).

Genus SYNORTHOCLADIUS Thienemann

SYNORTHOCLADIUS Thienemann, 1935: 211. Type species: Dactylocladius semivirens Kieffer, 1909, orig. des.
Unidentified sp.--Australia (ACT, NSW, SA).

Genus THIENEMANNIELLA Kieffer

THIENEMANNIELLA Kieffer, 1911a: 201 (Corynoneura subg.). Type species: Corynoneura clavicornis Kieffer, 1911, des. Goetghebuer, 1939: 7.
ogasaquardecima Sasa & Suzuki, 1997: 337. Bonin Is.
ogasaquindecima Sasa & Suzuki, 1997: 338. Bonin Is.
trivittata Goetghebuer, 1935: 377. Zaire; Australia (NSW); Afrotrop. Reg.


Genus TRONDIA Ferrington & Saether

TRONDIA Ferrington & Saether, 2006: 247. Type species: Trondia anderseni Ferrington & Saether, 2006, orig. des.
anderseni Ferrington & Saether, 2006: 248. Australia (WA).

Unplaced species of Orthocladiinae

adjectus Tokunaga, 1964a: 508 (Metriocnemus). Micronesia.
aequalis Kieffer, 1917a: 227 (Cricotopus). PNG (PNG).
campestris Hardy, 1960: 138 (Orthocladius). Hawaiian Is.
commensalis Tonnoir, 1923d: 284 (Dactylocladius). NZ (NZ).
davisi Hardy, 1960: 131 (Orthocladius). Hawaiian Is.
grimshawi Hardy, 1960: 132 (Orthocladius). Hawaiian Is.
herbicola Hardy, 1960: 147 (Metriocnemus). Hawaiian Is.
membranisensoria Hardy, 1960: 134 (Orthocladius). Hawaiian Is.
publicus Hutton, 1902b: 184 (Orthocladius). NZ (NZ). Nomen dubium.
williamsi Hardy, 1960: 135 (Orthocladius). Hawaiian Is.
wirthi Hardy, 1960: 145 (Orthocladius). Hawaiian Is.


Subfamily CHIRONOMINAE
Tribe PSEUDOCHIRONOMINI

Genus MEGACENTRON Freeman

MEGACENTRON Freeman, 1961: 676. Type species: Chironomus erebeus Skuse, 1889, orig. des.
erebeum Skuse, 1889e: 243 (Chironomus). Australia (NSW); Australia (Vic).

Genus RIETHIA Kieffer

RIETHIA Kieffer, 1917a: 203. Type species: Riethia stictoptera Kieffer, 1917, orig. des.
cinctipes Freeman, 1961: 679. Australia (Qld); Australia (ACT, Tas, Vic).
plumosa Freeman, 1961: 679. Australia (Tas).
stictoptera Kieffer, 1917a: 203. Australia (NSW); Australia (ACT, SA, Tas, Vic), PNG (PNG).
     astictica Kieffer, 1917a: 205. Australia (NSW).
zeylandica Freeman, 1959: 422. NZ (NZ); Australia (ACT, NSW, Tas).

Tribe TANYTARSINI

Genus CLADOTANYTARSUS Kieffer

CLADOTANYTARSUS Kieffer, 1921c: 277. Type species: Cladotanytarsus pallidus Kieffer, 1922, subs. mon. Kieffer, 1922d: 100.
australomancus Glover, 1973: 429. Australia (SA); Australia (ACT, NSW, Qld, Vic, WA).
     mancus, Freeman, 1961, not Walker, 1856, misid.
bilinearis Glover, 1973: 434. Australia (SA); Australia (Qld).
furcatus Freeman, 1961: 728. Australia (NSW); Australia (Qld).
tasmanicus Glover, 1973: 434. Australia (Tas).
unilinearis Glover, 1973: 434. Australia (SA); Australia (NT).

Genus CORYNOCERA Zetterstedt

CORYNOCERA Zetterstedt, 1838: 856. Type species: Corynocera ambigua Zetterstedt, 1838, mon.
Undescribed sp.--NZ (NZ) (Stark, 1981: 64).

Genus MICROPSECTRA Kieffer

MICROPSECTRA Kieffer, 1909: 50 (Tanytarsus subg.). Type species: Tanytarsus inermipes Kieffer, 1909 [= Chironomus notescens Walker, 1856], des. Kieffer, 1921g: 35.
bryanti Hardy, 1960: 102 (Calopsectra). Hawaiian Is.
hawaiiensis Hardy, 1960: 104 (Calopsectra). Hawaiian Is.
     ssp. albifasciata Hardy, 1960: 106 (Calopsectra). Hawaiian Is.
     ssp. hawaiiensis s. str. Hawaiian Is.
kaalae Hardy, 1960: 107 (Calopsectra). Hawaiian Is.
     ssp. kaalae s. str. Hawaiian Is.
     ssp. monticola Hardy, 1960: 109 (Calopsectra). Hawaiian Is.
lacteiclava Grimshaw, 1901: 5 (Tanytarsus). Hawaiian Is.
     ssp. lacteiclava s. str. Hawaiian Is.
     ssp. latifasciata Hardy, 1960: 110 (Calopsectra). Hawaiian Is.


Genus NEOZAVRELIA Goetghebeur

NEOZAVRELIA Goetghebeur in Goetghebeur & Thienemann, 1941: 106. Type species: Neozavrelia luteola Goetghebeur, 1941, mon.
bowmani Cranston, 1998: 109. Australia (NT).
optoputealis Cranston, 1998: 108. Australia (NSW).


Genus PARATANYTARSUS Thienemann & Bause

PARATANYTARSUS Thienemann & Bause in Bause, 1913: 120 [1914: 120]. Type species: Tanytarsus lauterborni Kieffer, 1909, des. Reiss & Säwedel, 1981: 74.
LUNDSTROEMIA, Freeman, 1961, not Kieffer, 1921, misid.
furvus Glover, 1973: 411. Australia (NSW); Australia (Vic, WA).
grimmii Schneider, 1885: 301 (Chironomus). ?USSR; widesp. Australia (excl. NT, Tas), NZ (NZ); virtually cosmopol.
     parthenogeneticus Freeman, 1961: 721 (Lundstroemia). Australia (WA).
     agameta Forsyth, 1971: 137 (Lundstroemia). NZ (NZ).
jefferyi Glover, 1973: 413. Australia (NSW); Australia (ACT, Qld, Vic, WA).
kathleenae Glover, 1973: 414. Australia (SA); Australia (NSW).
modicus Skuse, 1889e: 274 (Tanytarsus). Australia (NSW); Australia (ACT).

Genus PONTOMYIA Edwards

PONTOMYIA Edwards, 1926c: 796. Type species: Pontomyia natans Edwards, 1926, mon.
cottoni Womersley, 1937a: 102. Australia (SA); Australia (NSW, Qld, Vic, WA).
natans Edwards, 1926c: 796. Fiji. Australia (Qld); Marshall Is, Western Samoa; Palearctic Reg (Sinai).
oceana Tokunaga, 1964a: 626. Belau; Australia (Qld).
pacifica Tokunaga, 1932: 44. Japan; Australia (Qld); Singapore.

Genus RHEOTANYTARSUS Thienemann & Bause

RHEOTANYTARSUS Thienemann & Bause in Bause, 1913: 120 [1914: 20]. Type species: Tanytarsus pentapoda Kieffer, 1909, des. Goetghebuer, 1954: 132.
barrengarryensis Cranston, 1997: 711. Australia (NSW).
christinae Glover, 1973: 420. Australia (WA); Australia (NT, Qld).
flabellatus Glover, 1973: 420. Australia (ACT); Australia (NSW, Qld, SA, Tas, Vic).
gloveri Cranston, 1997: 718. Australia (NT): australia (Qld, WA).
juliae Glover, 1973: 424. Australia (SA); Australia (ACT, NSW, Qld, SA, Tas, Vic), Lord Howe I.
oss Cranston, 1997: 722. Australia (Qld); Australia (NT).
petropholeter Cranston, 1997: 722. Australia (NT).
trivittatus
Johannsen, 1932: 546 (Tanytarsus). Indonesia (Sumatra); Australia (NSW, NT, Qld, SA,Vic).
     johnstoni Glover, 1973: 422. Australia (SA).
underwoodi
Glover, 1973: 425. Australia (WA).

Genus STEMPELLINA Thienemann & Bause

STEMPELLINA Thienemann & Bause in Bause, 1913: 120 [1914: 120]. Type species: Tanytarsus bausei Kieffer, 1911, mon.
australiensis Freeman, 1961: 729. Australia (NSW).
emarginata Tokunaga, 1964a: 624 (Tanytarsus). Belau.
gressitti Tokunaga, 1964a: 625 (Tanytarsus). Belau.
johni Glover, 1973: 428. Australia (SA); Australia (NSW, Vic).


Genus STEMPELLINELLA Brundin

STEMPELLINELLA Brundin, 1947: 87. Type species: Tanytarsus saltuum Goetghebeur, 1921, orig. des.
Undetermined sp. - Hawaiian Is (Wolff et al., 2002: 32).

Genus TANYTARSUS Wulp

TANYTARSUS Wulp, 1874: 134. Type species: Chironomus signatus Wulp, 1874, des. I.C.Z.N., 1961: 361.
CALOPSECTRA Kieffer, 1909: 50 (Tanytarsus subg.). Type species: Tanytarsus gregarius Kieffer, 1909, des. Kieffer, 1921g: 36.
albanyensis Forsyth, 1971: 137. NZ (NZ).
annulatus Glover, 1973: 419 (Rheotanytarsus). Australia (NSW).
aviungulatus Glover, 1973: 472. Australia (NSW).
barbitarsis Freeman, 1961: 722. Australia (WA); Australia (Qld, SA, Vic).
     barbitarsus, error for barbitarsis
belairensis Glover, 1973: 468. Australia (SA); Australia (ACT, NSW).
belmorensis Glover, 1973: 474. Australia (NSW).
bispinosus Freeman, 1961: 727. Australia (NSW); Australia (ACT, NT, Qld, SA, Tas, Vic).
boninensis Tokunaga, 1964a: 606. Bonin Is; Guam.
brachyurus Tokunaga, 1964a: 621. Bonin Is.
collessi Glover, 1973: 457. Australia (WA).
commoni Glover, 1973: 458. Australia (ACT); Australia (NSW).
dostinei Cranston, 2000: 149. Australia (NT); Australia (WA).
dybasi Tokunaga, 1964a: 609. Belau; Northern Marianas, Micronesia.
edwardi Glover, 1973: 466. Australia (NSW); Australia (Tas, WA).
esakii Tokunaga, 1940a: 224. Micronesia; Marshall Is.
formosanus Kieffer, 1912: 42. Taiwan; Australia (Qld).
     dycei Glover, 1973: 444. Australia (Qld).
funebris
Freeman, 1959: 436. NZ (NZ); NZ (Auckland Is).
fuscithorax Skuse, 1889e: 272. Australia (NSW); widesp. Australia.
graniticus Glover, 1973: 460. Australia (WA).
gulungul Cranston, 2000: 141. Australia (NT); Australia (Qld, WA).
halophilae Edwards, 1926c: 791. Fiji; Belau, Marshall Is, Micronesia, Northern Marianas, Western Samoa.
hardwicki Cranston, 2000: 144. Australia (NT).
haugheyi
Glover, 1973: 467. Australia (WA).
hopkinsi Edwards, 1928c: 62. Western Samoa.
humphreyi Cranston, 2000: 152. Australia (Qld); Australia (NT).
inextentus Skuse, 1889e: 269. Australia (NSW); Australia (ACT, Tas, Vic, WA).
     communis Skuse, 1889e: 271. Australia (NSW).
insulicola Tokunaga, 1964a: 616. Belau; Guam.
lamnicaudus Tokunaga, 1964a: 608. Micronesia; Belau.
latiforceps Tokunaga, 1964a: 619. Belau.
leipae Glover, 1973: 461. Australia (ACT); Australia (NSW).
magnihamatus Tokunaga, 1933b: 362. Japan; Belau, Guam, Marshall Is, Micronesia.
manleyensis Glover, 1973: 462. Australia (NSW); Australia (Vic).
mendumae Glover, 1973: 462. Australia (ACT); Australia (NT).
micksmithi Cranston, 2000: 145. Australia (NT).
moruyaensis Glover, 1973: 450. Australia (NSW); Australia (WA).
neosydneyensis Glover, 1973: 475. Australia (NSW); Australia (SA), Lord Howe I.
     sydneyensis, Freeman, 1961, not Kieffer, 1917, ?misid.
nichollsi Glover, 1973: 450. Australia (WA); Australia (NT).
ogasaquartus Sasa & Suzuki, 1997: 322. Bonin Is.
ogasatertius Sasa & Suzuki, 1997: 322. Bonin Is.
ogilbyi Skuse, 1889e: 273 (Tanytarsus). Australia (NSW).
ovatus Johannsen, 1932c: 545. Indonesia (Java); Belau.
palmatus Freeman, 1961: 726. Australia (Vic); Australia (ACT, SA).
paskevillensis Glover, 1973: 452. Australia (SA).
pelagicus Tokunaga, 1933b: 364. Japan; Belau.
ponapensis Tokunaga, 1964a: 612. Micronesia; Belau.
richardsi Glover, 1973: 463. Australia (WA); Australia (NT, Qld).
riedi Cranston, 2000: 153. Australia (Qld); Australia (NT, WA).
rieki Glover, 1973: 471. Australia (NSW); Australia (Tas).
rosario Cranston, 2000: 151. Australia (NT).
semibarbitarsus Glover, 1973: 454. Australia (SA); Australia (NSW, NT, WA).
simplex Tokunaga, 1964a: 614. Belau; Micronesia.
spinosus Freeman, 1961: 726. Australia (Vic); Australia (ACT, NSW).
subinextentus Glover, 1973: 456. Australia (SA); Australia (NSW, Qld, Tas, Vic).
tricuspis Tokunaga, 1964a: 622. Belau.
upoluensis Ashe, 1985: 231 (n. n. for maritimus Edwards, 1926). Fiji; Belau, Micronesia, Western Samoa.
     maritimus Edwards, 1926c: 794. Fiji. [Preocc. Meunier, 1904.]
vespertinus Hutton, 1902b: 185. NZ (NZ).
wendylee Cranston, 2000: 149. Australia (NT).

Unplaced species of Tanytarsini

pauciplumatus Hardy, 1960: 115 (Chironomus). Hawaiian Is.
sydneyensis Kieffer, 1917a: 224 (Tanytarsus). Australia (NSW). Nomen dubium.

Tribe CHIRONOMINI

Genus ANUNCOTENDIPES Cranston

ANUNCOTENDIPES Cranston, 1999: 292. Type species: Anuncotendipes australotropicus Cranston, 1999, orig. des.
australotropicus Cranston, 1999: 294. Australia (Qld).
kakadu Cranston, 1999: 295. Australia (Qld).


Genus APEDILUM Townes

APEDILUM Townes, 1945: 32. Type species: Apedilum subcinctum Townes, 1945, orig. des.
Undetermined sp. — Hawaiian Is (Wolff et al., 2002: 32).


Genus AXARUS Roback

ANCEUS Roback, 1963: 237 (Xenochironomus subg.). Type species: Chironomus festivus Say, 1823, orig. des. [Preocc. Risso, 1816.]
AXARUS Roback, 1980: 32 (n. n. for Anceus).
Undescribed spp.--Australia (NT), PNG (PNG).

Genus CHIRONOMUS Meigen

CHIRONOMUS Meigen, 1803: 260. Type species: Tipula plumosa Linnaeus, 1758, des. Latreille, 1810: 442.

Subgenus CAMPTOCHIRONOMUS Kieffer

CAMPTOCHIRONOMUS Kieffer, 1918a: 45. Type species: Chironomus aprilinus Meigen, 1830 [misid., = Chironomus tentans Fabricius, 1805], des. Goetghebuer, 1937: 21. [I.C.Z.N. ruling required on conservation of misidentified type species.]
crassiforceps Kieffer, 1916: 111 (Tendipes). Taiwan; widesp. Pacific; Oriental Reg.
     esakii Tokunaga, 1940a: 221. Northern Marianas.
     insolens Johannsen, 1946: 192. Guam.

Subgenus CHIRONOMUS Meigen

CHIRONOMUS Meigen, 1803: 260 (as gen.). Type species: Tipula plumosa Linnaeus, 1758, des. Latreille, 1810: 442.
analis Freeman, 1959: 425. NZ (NZ).
antipodensis Sublette & Wirth, 1980: 326. NZ (Antipodes Is).
australis Macquart, 1847a: 9 [1847b: 25]. Australia (Tas); Australia (ACT, NSW, SA, Vic).
     applicatus Walker, 1856d: 424. Australia (Tas).
     imitans Walker, 1856d: 425. Australia (Tas).
bicoloris Tokunaga, 1964a: 557. Micronesia; Australia (Qld), Belau.
calipterus Kieffer, 1908: 158. Namibia; Belau, Australia (Qld); widesp. Afrotrop., Oriental, Palearctic Regs.
     pallidinubeculosus Tokunaga, 1964a: 549. Belau.
carolinensis Tokunaga, 1964a: 564. Micronesia; Belau.
cloacalis Atchley & Martin, 1971: 322. Australia (Vic); Australia (NSW, Qld, SA, WA).
     cloacalis Martin, 1967: 259 [1969a: 273]. Nomen nudum.
crassicaudus Tokunaga, 1964a: 556. Belau; Guam, Micronesia, Northern Marianas.
duplex Walker, 1856d: 424. Australia (Tas); Australia (NSW, SA, Vic).
forsythi Martin, 1999: 248. NZ (NZ).
hawaiiensis Grimshaw, 1901: 4. Hawaiian Is; ?Fiji.
javanus Kieffer, 1924a: 263. Indonesia (Java); Belau, Marshall Is, Micronesia.
magnivalva Kieffer, 1917a: 219. Australia (Qld); Fiji, French Polynesia (Society Is).
nepeanensis Skuse, 1889e: 231. Australia (NSW); Australia (NT, Qld, SA, Vic, WA).
nigerilateralis Tokunaga, 1964a: 565. Micronesia.
occidentalis Skuse, 1889e: 230. Australia (WA); Australia (NT).
oppositus Walker, 1856d: 425. Australia (Tas); Australia (NSW, Qld, SA, Vic).
     ssp. connori Martin & Lee, 1984: 191. Australia (NSW & Tas & Vic).
     ssp. oppositus s. str. Australia (Tas); Australia (NSW, SA, Vic).
     egregius Skuse, 1889e: 232. Australia (NSW & Qld).
     ssp. tyleri Martin & Lee, 1984: 191. Australia (NSW & SA & Tas & Vic).
     ssp. whitei Martin & Lee, 1984: 191. Australia (NSW & SA & Tas & Vic).
plumatisetigerus Tokunaga, 1964a: 560. Belau; Australia (NT), Guam, Micronesia, PNG (PNG); India.
praeapicalis Tokunaga, 1964a: 562. Belau; Micronesia.
samoensis Edwards, 1928c: 67. Western Samoa & American Samoa & Tonga; Belau, ?French Polynesia (Marquesas), Guam, Marshall Is, Micronesia.
     eximius Johannsen, 1946: 193. Guam.
     dorsalis, Tokunaga, 1940, not Meigen, 1818, misid.
sexipunctatus Tokunaga, 1964a: 569. Guam.
subantarcticus Sublette & Wirth, 1980: 327. NZ (Campbell I); NZ (Auckland Is).
tepperi Skuse, 1889e: 244. Australia (SA); widesp. Australia (excl. Tas).
trifasciatus Tokunaga, 1964a: 561. Micronesia.
zealandicus Hudson, 1892: 43. NZ (NZ).
     novaezelandiae Kieffer, 1922a: 146. NZ (NZ).

Nomina dubia in Chironomus Meigen

alternans Walker, 1856d: 423. Australia (NSW); Australia (ACT, Qld, SA, Vic, WA).
arcuatus Kieffer, 1917a: 219. Australia (NSW).
imberbis Kieffer, 1917a: 210. Australia (NSW).
januarius Skuse, 1889e: 239. Australia (NSW).
pervagatus Skuse, 1889e: 233. Australia (NSW).
subdolus Skuse, 1889e: 236. Australia (NSW).
tenuiventris Kieffer, 1917a: 217. Australia (NSW).

Nomina nuda in Chironomus Meigen

februarius Martin, 1967: 259 [1969a: 273].
pseudoppositus Martin, 1969a: 273.

Genus CLADOPELMA Kieffer

CLADOPELMA Kieffer, 1921c: 274 [1921e: 63; 1921g: 31]. Type species: Chironomus virescens Meigen, 1818, des. Harnisch, 1923: 304.
curtivalva Kieffer, 1917a: 213 (Chironomus). Australia (NSW); Australia (ACT, NT, SA, Vic, WA), NZ (NZ).
     cylindricus, Forsyth, 1971, not Freeman, 1959, misid.
     pulcher, Skuse, 1889, not Wiedemann, 1830, misid.

Genus CONOCHIRONOMUS Freeman

CONOCHIRONOMUS Freeman, 1961: 701. Type species: Endochironomus acutistilus Freeman, 1955 (as "acutistillus"), orig. des.
acutistilus Freeman, 1955b: 288 (Endochironomus). Zaire; Australia (Qld); Afrotrop. Reg.
australiensis Cranston & Hare, 1995: 260. Australia (NT); Australia (Qld, WA).
cervus Cranston & Hare, 1995: 262. Australia (NT); Australia (WA).
cygnus Cranston & Hare, 1995: 262. Australia (Qld); Australia (NT, WA).
kakadu Cranston & Hare, 1995: 261. Australia (NT); Australia (Qld, WA).

Genus CRYPTOCHIRONOMUS Kieffer

CRYPTOCHIRONOMUS Kieffer, 1918a: 46 (Chironomus subg.). Type species: Chironomus chlorolobus Kieffer, 1918, orig. des.
griseidorsum Kieffer, 1917a: 218 (Chironomus). Australia (NSW); widesp. Australia (excl. ACT), PNG (PNG).
javae Kieffer, 1924a: 264. Indonesia (Java); Belau.
ponapensis Tokunaga, 1964a: 576 (Chironomus). Micronesia.

Genus DICROTENDIPES Kieffer

DICROTENDIPES Kieffer, 1913e: 23. Type species: Dicrotendipes pictipennis Kieffer, 1913 [= Chironomus quatuordecimpunctatus Goetghebuer, 1936], mon.
balciunasi Epler, 1988: 113. Australia (NT); Australia (WA).
bilobatus Kieffer, 1917a: 222. Australia (NSW); Australia (ACT).
candidibasis Edwards, 1924f: 573 (Chironomus). Fiji; Western Samoa.
     melanocnemis Edwards, 1928c: 65 (Chironomus). Western Samoa.
conjunctus Walker, 1856d: 425 (Chironomus). Australia (Tas); Australia (ACT, NSW, Qld, Tas, WA).
cumberlandensis Epler, 1988: 125. Australia (Vic); Australia (Qld).
flexus Johannsen, 1932c: 530 (Chironomus). Indonesia (Java); Australia (NT).
jobetus Epler, 1988: 130. Australia (NT); Australia (WA).
jonmartini Epler, 1988: 131. Australia (SA); Australia (WA).
leei Freeman, 1961: 691 (Chironomus). Australia (NSW).
lindae Epler, 1988: 133. Australia (Qld); Australia (NT).
nervosus Staeger, 1839: 567 (Chironomus). Denmark; Bonin Is; Europe, Japan, Nearctic Reg..
pelochloris Kieffer, 1912d: 39 (Tendipes). Taiwan; Australia (NSW, NT, Qld); Oriental, Palearctic Regs.
     wirthi Freeman, 1961: 692 (Chironomus). Australia (NSW).
pseudoconjunctus Epler, 1988: 139. Australia (Tas); Australia (SA, Vic, WA).
sarinae Epler, 1988: 142. Australia (Qld); Australia (WA).
septemmaculatus Becker, 1908: 77 (Chironomus). Canary Is: Australia (NSW, NT, Qld); Afrotrop., Oriental, Palearctic Regs.
     pilosimanus Kieffer, 1914: 262. South Africa.
taylori Freeman, 1961: 692 (Chironomus). Australia (Qld); PNG (PNG).
tenuiforceps Kieffer, 1913d: 136 (Tendipes). India; Australia (Qld).
     innisfailensis Freeman, 1961: 694 (Chironomus). Australia (Qld).

Genus EINFELDIA Kieffer

EINFELDIA Kieffer, 1924c: 393. Type species: Einfeldia pectoralis Kieffer, 1924, mon.
vitellina Freeman, 1961: 686 (Chironomus). Australia (NT); Australia (NSW, Qld), Fiji, PNG (PNG).

Genus ENDOCHIRONOMUS Kieffer

ENDOCHIRONOMUS Kieffer, 1918b: 69. Type species: Chironomus alismatus Kieffer, 1915 [= Tipula tendens Fabricius, 1775], orig. des.
palauensis Tokunaga, 1964a: 571 (Chironomus). Belau.
tenuicaudus Tokunaga, 1964a: 572 (Chironomus). Micronesia.

Genus FISSIMENTUM Cranston & Nolte

FISSIMENTUM Cranston & Nolte, 1996: 2. Type species: Fissimentum desiccatum Cranston & Nolte, 1996, orig. des.
Undescribed sp.--Australia (Qld, Vic.) (Cranston & Nolte, 1996: 12).

Genus GILLOTIA Kieffer

GILLOTIA Kieffer, 1921c: 272 [1921g: 31]. Type species: Gillotia fuscipes Kieffer, 1922 [preocc., = Chironomus trifidus Freeman, 1957], subs. mon. Kieffer, 1922c: 62.
Undescribed sp.--Australia (NSW).

Genus GOELDICHIRONOMUS Fittkau

GOELDICHIRONOMUS Fittkau, 1965: 210. Type species: Chironomus holoprasinus Goeldi, 1905, orig. des.
holoprasinus Goeldi, 1905: 135 (Chironomus). Brazil; Hawaiian Is; New World.

Genus HARNISCHIA Kieffer

HARNISCHIA Kieffer, 1921c: 273 [1921e: 69]. Type species: Harnischia fuscimana Kieffer, 1921, subs. mon. Kieffer, 1921e: 69.
curtilamellata Malloch, 1915a: 474 (Chironomus). USA; Australia (ACT); Afrotrop., Holarctic, Oriental Regs.
     ?dycei Freeman, 1961: 701 (Chironomus). Australia (ACT).

Genus HARRISIUS Freeman

HARRISIUS Freeman, 1959: 426. Type species: Harrisius pallidus Freeman, 1959, orig. des.
montanus Skuse, 1889e: 268 (Tanytarsus). Australia (NSW); Australia (Vic).
     cereolus Skuse, 1889e: 270 (Tanytarsus). Australia (NSW).
pallidus Freeman, 1959: 426. NZ (NZ).

Genus IMPARIPECTEN Freeman

IMPARIPECTEN Freeman, 1961: 703. Type species: Imparipecten pictipes Freeman, 1961, orig. des.
pictipes Freeman, 1961: 703. Australia (Tas); Australia (ACT, NSW).

Genus KIEFFERULUS Goetghebuer

CARTERIA Kieffer, 1921d: 590. Type species: Tendipes longilobus Kieffer, 1916, orig. des. [Preocc. Diesing, 1866.]
KIEFFERULUS Goetghebuer, 1922: 40. Type species: Tanytarsus tendipediformis Goetghebuer, 1921, orig. des.
CARTERONICA Strand, 1928: 48 (n. n. for Carteria).
NILODORUM , authors, not Kieffer, 1921, misid.
intertinctus Skuse, 1889e: 234 (Chironomus). Australia (NSW); widesp. Australia (excl. NT).
     hexhamensis Skuse, 1889e: 237 (Chironomus). Australia (NSW).
     lativalva Kieffer, 1917a: 211 (Chironomus). Australia (NSW).
longilobus Kieffer, 1916: 107 (Tendipes). Taiwan; Australia (NSW, Qld, WA), Belau, Guam, Micronesia, Northern Marianas; Japan.
martini Freeman, 1961: 690 (Chironomus). Australia (Vic); Australia (NSW, NT, Qld, Tas).
opalensis Forsyth, 1975: 215. NZ (NZ).
paratinctus Martin, 1964: 285 (Chironomus). Australia (Vic); Australia (NSW, Qld).
tainanus Kieffer, 1912d: 36 (Tendipes). Taiwan; Australia (NT, Qld); Japan, Oriental Reg.
     biroi Kieffer, 1918b: 108 (Chironomus). Sri Lanka.

Genus MICROCHIRONOMUS Kieffer

MICROCHIRONOMUS Kieffer, 1918b: 113 (Chironomus subg.). Type species: Chironomus lendli Kieffer, 1918, orig. des.
tener Kieffer, 1918a: 48 (Chironomus). Poland; Australia (Qld); Australia (ACT, NT, Vic); Afrotrop., Oriental, Palearctic Regs.
     forcipatus Freeman, 1954a: 20 (Chironomus). South Africa.

Genus MICROTENDIPES Kieffer

MICROTENDIPES Kieffer, 1915: 70. Type species: Tendipes abbreviatus Kieffer, 1913 [= Chironomus chloris Meigen, 1818], orig. des.
umbrosus Freeman, 1955a: 32. Kenya; Australia (Qld), PNG (PNG); Afrotrop. Reg.


Genus NANDEVA Wiedenbrug, Reiss, and Fittkau

NANDEVA Wiedenbrug, Reiss, & Fittkau, 1998: 60. Type species: Nandeva gaucha Wiedenbrug, Reiss & Fittkau, 1998, orig. des.
fittkaui Cranston, 1999: 296. Australia (Qld).

Genus NILOTHAUMA Kieffer

NILOTHAUMA Kieffer, 1921c: 270. Type species: Nilothauma pictipenne Kieffer, 1921, subs. mon. Kieffer, 1921g: 37.
Unidentified sp.--Australia (WA).

Genus OPHRYOPHORUS Freeman

OPHRYOPHORUS Freeman, 1959: 427. Type species: Ophryophorus ramiferus Freeman, 1959, orig. des.
ramiferus Freeman, 1959: 428. NZ (NZ).

Genus PARABORNIELLA Freeman

PARABORNIELLA Freeman, 1961: 717. Type species: Paraborniella tonnoiri Freeman, 1961, orig. des.
tonnoiri Freeman, 1961: 718. Australia (ACT); Australia (NSW, Vic, WA).

Genus PARACHIRONOMUS Lenz

PARACHIRONOMUS Lenz, 1921: 160. Type species: Chironomus cryptomus Kieffer, 1915, des. Townes, 1945: 152.
cylindricus Freeman, 1959: 425 (Chironomus). NZ (NZ).
delinificus Skuse, 1889e: 239 (Chironomus). Australia (NSW); Australia (Tas, Vic).

Genus PARACLADOPELMA Harnisch

PARACLADOPELMA Harnisch, 1923: 304. Type species: Tendipes camptolabis Kieffer, 1913c, orig. des.
angustum Freeman, 1961: 699 (Chironomus). Australia (NSW).

Genus PARALAUTERBORNIELLA Lenz

PARALAUTERBORNIELLA Lenz, 1941: 48. Type species: Chironomus brachylabis Edwards, 1929 [= Chironomus nigrohalteralis Malloch, 1915], mon.
Unidentified sp.--Australia (Qld).

Genus PARATENDIPES Kieffer

PARATENDIPES Kieffer, 1911e: 41. Type species: Chironomus albimanus Meigen, 1818, orig. des.
Unidentified sp.--Australia (Qld).

Genus PARVITERGUM Freeman

PARVITERGUM Freeman, 1961: 716. Type species: Parvitergum fuscipenne Freeman, 1961, orig. des.
fuscipenne Freeman, 1961: 717. Australia (Vic).

Genus PAUCISPINIGERA Freeman

PAUCISPINIGERA Freeman, 1959: 428. Type species: Paucispinigera approximata Freeman, 1959, orig. des.
approximata Freeman, 1959: 429. NZ (NZ).

Genus PHAENOPSECTRA Kieffer

PHAENOPSECTRA Kieffer, 1921c: 276. Type species: Chironomus leucolabis Kieffer, 1915 [= Chironomus flavipes Meigen, 1818], des. Goetghebuer, 1938: 80.
gressitti Tokunaga, 1964a: 579. Micronesia; Guam, Marshall Is.

Genus POLYPEDILUM Kieffer

POLYPEDILUM Kieffer, 1912d: 41 [1913a: 15]. Type species: Polypedilum pelostolum Kieffer, 1912 [= Chironomus nubifer Skuse, 1889], des. Ashe, 1981: 58.

Subgenus PENTAPEDILUM Kieffer

PENTAPEDILUM Kieffer, 1913b: 25 (as gen.). Type species: Pentapedilum stratiotale Kieffer, 1913 [= Chironomus tritus Walker, 1856], des. Edwards, 1929i: 376.
convexum Johannsen, 1932c: 540 (Pentapedilum). Indonesia (Sumatra); Belau, Micronesia; Japan.
elongatum Tokunaga, 1964a: 600. Micronesia.
esakii Tokunaga, 1964a: 600 (Pentapedilum). Micronesia; Guam.
leei Freeman, 1961: 713. Australia (NSW); Australia (NT, Qld, SA, WA).
nodosum Johannsen, 1932c: 541 (Pentapedilum). Indonesia (Java); Belau, Micronesia, Northern Marianas, PNG (PNG); Indonesia (Sumatra).
palauense Tokunaga, 1964a: 601. Belau.
trukense Tokunaga, 1940a: 223 (Pentapedilum). Micronesia; Belau.

Subgenus POLYPEDILUM Kieffer

POLYPEDILUM Kieffer, 1912d: 41 [1913a: 15] (as gen.). Type species: Polypedilum pelostolum Kieffer, 1912 [= Chironomus nubifer Skuse, 1889], des. Ashe, 1981: 58.
albicorpus Tokunaga, 1964a: 594. Belau.
alternans Forsyth, 1971: 133. NZ (NZ).
canum Freeman, 1959: 434. NZ (NZ).
     ?ignavus Hutton, 1902b: 183 (Chironomus). NZ (NZ).
concomitatum Johannsen, 1932c: 517 (Chironomus). Indonesia (Java); Belau.
cumberi Freeman, 1959: 434. NZ (NZ).
digitulum Freeman, 1959: 433. NZ (NZ).
dybasi Tokunaga, 1964a: 587. Belau; Micronesia.
griseoguttatum Kieffer, 1921f: 100. Cameroon; ?Australia (NSW, NT, Qld, Vic); India, widesp. Afrotrop. Reg.
harrisi Freeman, 1959: 433. NZ (NZ).
insulanum Tokunaga, 1964a: 586. Micronesia.
longicrus Kieffer, 1921f: 101. Zaire; NZ (NZ); Afrotrop. Reg.
luteum Forsyth, 1971: 133. NZ (NZ).
medivittatum Tokunaga, 1964a: 588. Belau; Guam, Micronesia.
nigribasale Tokunaga, 1964a: 590. Micronesia.
nubiferum Skuse, 1889e: 249 (Chironomus). Australia (NSW); widesp. Australia, Bonin Is, Hawaiian Is; Afrotrop., Oriental, Palearctic Regs.
     albiceps Johannsen, 1932c: 515 (Chironomus). Indonesia (Java).
     ?novemmaculatum Hardy, 1960: 117. Hawaiian Is.
opimum Hutton, 1902b: 182 (Chironomus). NZ (NZ).
oresitrophum Skuse, 1889e: 247 (Chironomus). Australia (NSW); Australia (Tas, Vic).
     brevis Skuse, 1889e: 249 (Chironomus). Australia (NSW).
pavidum Hutton, 1902b: 183 (Chironomus). NZ (NZ).
perturbans Johannsen, 1932c: 518 (Chironomus stictopterus Kieffer, var.). Indonesia (Sumatra); Guam.
ponapense Tokunaga, 1964a: 585. Micronesia.
prasiogaster Freeman, 1961: 713. Australia (NSW); Australia (ACT, Tas, Vic).
seorsum Skuse, 1889e: 241 (Chironomus). Australia (NSW); Australia (Vic).
     orarius Skuse, 1889e: 242 (Chironomus). Australia (NSW).
spadix Tokunaga, 1964a: 593. Belau; Micronesia.
tonnoiri Freeman, 1961: 710. Australia (Tas); Australia (NSW, Vic, WA).
vespertinum Skuse, 1889e: 248 (Chironomus). Australia (NSW).
     venustulus Skuse, 1889e: 257 (Orthocladius). Australia (NSW).
watsoni Freeman, 1961: 710. Australia (WA); Australia (ACT, NSW, NT, Vic).
wirthi Freeman, 1961: 712. Australia (NSW).
yapense Tokunaga, 1964a: 594. Micronesia; Belau.

Unplaced species of Polypedilum Kieffer

lentum Hutton, 1902b: 182 (Chironomus). NZ (NZ). Nomen dubium.

Genus SKUSELLA Freeman

SKUSELLA Freeman, 1961: 718. Type species: Chironomus subvittatus Skuse, 1889, orig. des.
subvittata Skuse, 1889e: 246 (Chironomus). Australia (NSW).

Genus STENOCHIRONOMUS Kieffer

STENOCHIRONOMUS Kieffer in Kieffer & Thienemann, 1919: 44. Type species: Tipula flexilis Linnaeus, 1767 [misid., = Chironomus gibbus Fabricius, 1794], des. Goetghebuer, 1937: 12.

Subgenus PETALOPHOLEUS Borkent

PETALOPHOLEUS Borkent, 1984: 92. Type species: Stenochironomus cinctus Townes, 1945, orig. des.
Undescribed spp.--Australia (Qld), Fiji.

Subgenus STENOCHIRONOMUS Kieffer

STENOCHIRONOMUS Kieffer in Kieffer & Thienemann, 1919: 44 (as gen.). Type species: Tipula flexilis Linnaeus, 1767 [misid., = Chironomus gibbus Fabricius, 1794], des. Goetghebuer, 1937: 12.
watsoni Freeman, 1961: 705. Australia (WA); Australia (Qld, Vic).

Unplaced species of Stenochironomus Kieffer

longilobatus Tokunaga, 1964a: 592 (Polypedilum). Micronesia.
sabroskyi Tokunaga, 1964a: 577. Belau.

Genus STICTOCHIRONOMUS Kieffer

STICTOCHIRONOMUS Kieffer in Kieffer & Thienemann, 1919: 44. Type species: Chironomus pictulus Meigen, 1830, des. Townes, 1945: 77.
fluviaticus Skuse, 1889e: 245 (Chironomus). Australia (NSW); Australia (Qld).
illawarra Freeman, 1961: 716. Australia (NSW); Australia (Vic).
townesi Tokunaga, 1964a: 578. Micronesia.

Genus XENOCHIRONOMUS Kieffer

XENOCHIRONOMUS Kieffer, 1921c: 273. Type species: Chironomus xenolabis Kieffer, 1916, des. Goetghebuer, 1937: 32.
canterburyensis Freeman, 1959: 425 (Chironomus). NZ (NZ); Australia (SA, Vic).


Genus XYLOCHIRONOMUS Cranston

XYLOCHIRONOMUS Cranston, 2006: 228. Type specis: Xylochironomus kakadu Cranston, 20906, orig. des.
kakadu Cranston, 2006: 231. Australia (NT).

Genus YAMA Sublette & Martin

YAMA Sublette & Martin, 1980: 222. Type species: Yama tahitiensis Sublette & Martin, 1980, orig. des.
tahitiensis Sublette & Martin, 1980: 226. French Polynesia (Society Is).

Genus ZAVRELIELLA Kieffer

ZAVRELIELLA Kieffer, 1920: 334. Type species: Chironomus clavaticrus Kieffer, 1913 [= Chironomus marmoratus Wulp, 1885], orig. des.
fuscoguttata Kieffer, 1922c: 37 (Polypedilum). Sudan; Australia (NSW, NT); Afrotrop. Reg.

Unplaced species of Chironomini

anomalus Freeman, 1961: 706 (Stenochironomus). Australia (Tas); Australia (ACT, SA, Vic)
australiensis Freeman, 1961: 696 (Chironomus). Australia (NSW); Australia (ACT, SA, Tas).

Nomina dubia in Chironominae

apicale Kieffer, 1917a: 221 (Polypedilum). PNG (PNG).
blandellus Kieffer, 1906: 16 (Chironomus; n. n. for blandus Skuse). Australia (NSW); Australia (Vic).
     blandus Skuse, 1889e: 238 (Chironomus). Australia (NSW). [Preocc. Wulp, 1858.]
dissimilis Kieffer, 1917a: 210 (Chironomus). PNG (PNG).
dolichopelma Kieffer, 1917a: 215 (Chironomus). PNG (PNG).
erimanus Kieffer, 1917a: 211 (Chironomus). PNG (PNG).
humilis Kieffer, 1917a: 220 (Chironomus). Australia (NSW); PNG (PNG).
huonensis Kieffer, 1917a: 215 (Chironomus). PNG (PNG).
instabilis Walker, 1865a: 103 (Chironomus). "New Guinea" [= Indonesia (Irian Jaya)].
microtomus Kieffer, 1917a: 218 (Chironomus). Australia (NSW).
neglectus Kieffer, 1917a: 220 (Chironomus). PNG (PNG).
pallidiceps Kieffer, 1917a: 215 (Chironomus). PNG (PNG).
pallidiforceps Kieffer, 1917a: 215 (Chironomus). PNG (PNG).
quadrifasciatus Kieffer, 1917a: 216 (Chironomus). Australia (Qld).
sydneyensis Kieffer, 1917a: 213 (Chironomus). Australia (NSW).



This page last revised 17 April 2017