Australasian/Oceanian Diptera Catalog — Web Version

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22. Family CULICIDAE

[Originally] By Neal L. Evenhuis & Samuel M. Gon III

Culicids, or mosquitoes, are represented in the Australasian/ Oceanian Regions by the 3 known subfamilies (Anophelinae, Toxorhynchitinae, and Culicinae) comprising over 600 valid species and subspecies in almost 70 genera and subgenera.

Adult females of most species are obligate blood feeders and play an important medico-veterinary role as transmitters of various diseases, while others are important biting pests. In the Australasian/Oceanian Regions certain species are known to transmit malaria, filariasis, and viral diseases such as dengue and encephalitis. In contrast, adults of Toxorhynchites are non­biting. Their larvae are predaceous on other aquatic invertebrates (Steffan & Evenhuis 1981) and some species have been introduced onto Pacific islands such as Hawaii and Tahiti for bio­logical control of culicine mosquitoes.

Larvae and pupae of culicids are aquatic, occupying standing or slow-moving waters under conditions ranging from temporary accumulations of water held by plants, tires, cans, and other containers, to vast estuarine wetlands. Because of the medical importance of the fam­ily, a large body of literature exists regarding the natural history, epidemiological role, and control of many of the species.
Refs.: Russel et al. (1943, keys, lists, world Anopheles), Bohart (1957, Micronesian spp.), Steffan (1966a, checklist Papuan Subregion spp.), Knight & Stone (1977, cat. world spp., bibliogr.), Knight (1978, cat. world spp.), Lee et al. (1980, cat., bibliogr. Australasian spp.; 1982, 1984, 1987a,b, rev. Australian spp.), Steffan et al. (1980, bibliogr. Toxorhynchites), Manning et al. (1982, bibliogr. Toxorhynchites), Ward (1984, cat. world spp.).

Genus ANOPHELES Meigen

ANOPHELES Meigen, 1818: 10. Type species: Anopheles maculipennis Meigen, 1818, des. I.C.Z.N., 1959a: 155.

Subgenus ANOPHELES Meigen

ANOPHELES Meigen, 1818: 10 (as gen.). Type species: Anopheles maculipennis Meigen, 1818, des. I.C.Z.N., 1959a: 155.
MYZORHYNCHUS Blanchard, 1902: 795 (as gen.). Type species: Anopheles sinensis Wiedemann, 1828, orig. des.
NEOSTETHOPHELES James, 1910: 98 (as gen.). Type species: Anopheles aitkenii James, 1903, orig. des.
aitkenii James—not Australasian/Oceanian.
albotaeniatus Theobald, 1903b: 88 (Myzorhynchus). Malaysia (Pen); ?Indonesia (Irian Jaya, Maluku); Indonesia (Java, Kalimantan, Sulawesi, Sumatra), Malaysia (Sabah, Sarawak).
argyropus Swellengrebel, 1914: 334 (Myzorhynchus). Indonesia (Sumatra); ?Indonesia (Maluku); Oriental Reg.
atratipes Skuse, 1889c: 1755. Australia (NSW) [original type locality]; Australia (Qld) [neotype locality; see Mackerras, 1927a: 38]; Australia (SA, Vic, WA).
baezai Gater, 1933: 189 [1934: 162]. Malaysia (Pen); Guam, ?Indonesia (Maluku); Cambodia, Indonesia, Philippines, Thailand, Vietnam.
bancroftii Giles, 1902: 511. Australia (Qld); Australia (NT, WA), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG).
     bancrofti, error for bancroftii.
barbirostris Wulp, 1884b: 248. Indonesia (Java); Guam [immigrant], Indonesia (Maluku); widesp. Oriental Reg.
barbumbrosus Strickland & Chowdhury— not Australasian/Oceanian.
bengalensis Puri—not Australasian/Oceanian.
colledgei Marks, 1956: 41. Australia (Qld).
corethroides Theobald, 1907: 35. Australia (Qld).
hyrcanus Pallas—not Australasian/Oceanian.
insulaeflorum Swellengrebel & Swellengrebel-de Graaf, 1919: 2 [1920: 34] (Stethomyia aitkenii var.). Indonesia (Java); Indonesia (Irian Jaya, Maluku); Indonesia (Sumatra), widesp. Oriental Reg.
lesteri Baisas & Hu—not Australasian/Oceanian.
maculipennis Meigen—not Australasian/Oceanian.
nigerrimus Giles, 1900: 162. India; Indonesia (Maluku); Indonesia (Java, Nusa Tenggara, Sulawesi, Sumatra), widesp. Oriental Reg.
     venhuisi Bonne-Wepster, 1951: 284. Indonesia (Java).
palmatus Rodenwaldt, 1926: 794 [1927: 518] (Stetomyia aitkeni var.). Indonesia (Java); Indonesia (Irian Jaya, Maluku); Indonesia (Kalimantan, Sumatra), Malaysia (Pen, Sabah, Sarawak), Thailand.
papuensis Dobrotworsky, 1957: 185. PNG (PNG); ?Indonesia (Irian Jaya).
     stigmaticus, authors, not Skuse, 1889, misid.
peditaeniatus Leicester, 1908: 31 (Myzorhynchus). Malaysia (Pen); ?Indonesia (Maluku); Oriental Reg.
pilinotum Harrison & Scanlon, 1974: 33. Philippines; Indonesia (?Irian Jaya, Maluku).
powelli Lee, 1944a: 21. Australia (NT); Australia (Qld).
pseudobarbirostris Ludlow, 1902: 129. Philippines; Australia (Qld), Indonesia (Irian Jaya, Maluku), PNG (PNG); Indonesia (Sulawesi).
pseudostigmaticus Dobrotworsky, 1957: 184. Australia (Vic); Australia (NSW, Qld).
separatus Leicester, 1908: 36 (Myzorhynchus). Malaysia (Pen); Indonesia (Maluku); Cambodia, Indonesia (Sumatra), Malaysia (Sarawak), Thailand, Vietnam.
sinensis Wiedemann, 1828—not Australasian/Oceanian.
stigmaticus Skuse, 1889c: 1758. Australia (NSW); Australia (Qld, Tas, Vic), Indonesia (Irian Jaya).
tasmaniensis Dobrotworsky, 1966: 130. Australia (Tas); Australia (Vic).
umbrosus Theobald—not Australasian/Oceanian.
vanus Walker, 1859b: 91. Indonesia (Sulawesi); Indonesia (Irian Jaya, Maluku); Malaysia (Sarawak), Philippines.
     barbumbrosus, authors, not Strickland & Chowdhury, 1927, misid.

Subgenus CELLIA Theobald

GRASSIA Theobald, 1902: 181 (as gen.). Type species: Anopheles rossii Giles, 1899 [= Anopheles subpictus Grassi, 1899], orig. des.
CELLIA Theobald, 1902: 181 [1903: 107] (as gen.). Type species: Anopheles pharoensis Theobald, 1901, orig. des.
MYZOMYIA Blanchard, 1902: 795 (as gen.; n. n. for Grassia).
NEOMYZOMYIA Theobald, 1910b: 29 (as gen.). Type species: Myzomyia elegans James, 1903, mon.
PALAEOMYZOMYIA Swellengrebel & Rodenwaldt, 1932: 114 (Anopheles subg.). Type species: Myzomyia parangensis Ludlow, 1914, mon.
aconitus Dönitz, 1903: 70. Indonesia (Sumatra); Indonesia (Maluku); Indonesia (Java, Nusa Tenggara, Sulawesi, Timor), widesp. Oriental Reg.
amictus Edwards, 1921b: 71. Australia (Qld); Australia (NSW, NT, WA).
annulatus Haga, 1930: 368, 373, 380 (keys) (longirostris var.). Indonesia (Irian Jaya).
     annulatus de Rook, 1929: 29, 36, 45 (longirostris var.). Unavailable name; see Ward, 1979: 130.
annulipes Walker, 1856d: 433. Australia (Tas); widesp. Australia, Indonesia (Irian Jaya), Lord Howe I, PNG (PNG).
     musivus Skuse, 1889c: 1754. Australia (NSW).
     mastersi Skuse, 1889c: 1757. Australia (NSW).
     perplexus Taylor, 1943: 153. Australia (NT).
     persimilis Taylor, 1943: 155 (perplexus var.). Australia (Qld).
     derricki Taylor, 1943: 155. Australia (Qld).
     masteri, error for mastersi.
     muscivus, musicus, errors for musivus.
clowi Rozeboom & Knight, 1946: 114. Indonesia (Irian Jaya).
cristatus King & Baisas, 1936: 80. Philippines; ?Indonesia (Irian Jaya).
farauti Laveran, 1902: 908. Vanuatu; Australia (NT, Qld, WA),
Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia (Sulawesi).
     moluccensis Swellengrebel & Swellengrebel-de Graaf, 1919: 1 [1920: 29] (annulipes var.). Indonesia (Maluku).
flavirostris Ludlow, 1914a: 130 (Myzomyia). Philippines; Indonesia (Maluku); widesp. Indonesia.
hilli Woodhill & Lee, 1944: 63. Australia (NT); Australia (Qld, WA), Indonesia (Irian Jaya).
incognitus Brug, 1931a: 136. Indonesia (Irian Jaya).
indefinitus Ludlow, 1904b: 299 (Myzomyia rossi var.). Philippines; Guam [immigrant], Northern Marianas; Oriental Reg.
karwari James in Theobald, 1903b: 102 (Nyssorhynchus). India; Indonesia (Irian Jaya), PNG (PNG); widesp. Oriental Reg.
kochi Dönitz, 1901: 36. Indonesia (Sumatra); Indonesia (Maluku); widesp. SE Oriental Reg.
koliensis Owen, 1945: 236. Solomon Is; Indonesia (Irian Jaya), PNG (Bougainville I, PNG).
litoralis King, 1932: 311. Philippines; Guam [immigrant]; Malaysia (Sabah).
longirostris Brug, 1928a: 278 [1928c: 424; 1928d: 297]. Indonesia (Irian Jaya); Indonesia (Maluku), PNG (Bismarck Arch, PNG).
ludlowae Theobald, 1903b: 42 (Myzomyia; as ludlowii). Philippines; Indonesia (Maluku); China (Hainan I), Indonesia (Sulawesi), Malaysia (Pen, Sarawak), Taiwan.
lungae Belkin & Schlosser, 1944: 269. Solomon Is; PNG (Bougainville I).
maculatus Theobald, 1901b: 99, 171. Hong Kong; ?Indonesia (Maluku); widesp. Oriental Reg.
meraukensis Venhuis, 1932: 1040. Indonesia (Irian Jaya); Australia (NT, Qld, WA), PNG (PNG).
minimus Theobald—not Australasian/Oceanian.
nataliae Belkin, 1945: 315. Solomon Is; PNG (Bougainville I).
novaguinensis Venhuis, 1933: 203 (punctulatus var.). Indonesia (Irian Jaya); Australia (NT, Qld, WA).
     breinli Taylor, 1943: 156. Australia (Qld).
     novaginensis, error for novaguiensis.
paragensis Ludlow, 1914a: 129 (Myzomyia). Philippines; Indonesia (Maluku); Indonesia (Kalimantan, Sulawesi, Sumatra), Malaysia (Pen, Sarawak).
punctipennis Say, 1823: 9 (Culex). e. USA; Hawaiian Is [introduced; not established]; Nearctic Reg.
punctulatus Dönitz, 1901: 36. PNG (Bismarck Arch & PNG); ?Australia (NT), Indonesia (Irian Jaya, Maluku); PNG (Bougainville I), Solomon Is.
rennellensis Taylor & Maffi, 1999: 195. Solomon Is.
solomonis Belkin, Knight & Rozeboom, 1945: 247. Solomon Is; PNG (Bougainville I).
subpictus Grassi in Grassi et al., 1899: 101. India; Guam [immigrant], Indonesia (Irian Jaya, Maluku), Northern Marianas [immigrant], PNG (Bismarck Arch, PNG); widesp. SE Oriental Reg.
sundaicus Rodenwaldt, 1925: 185 (Myzomyia ludlowi var.). "Indonesia"; Indonesia (Maluku); widesp. SE Oriental Reg.
tessellatus Theobald, 1901b: 175. Malaysia (Pen); Indonesia (?Irian Jaya, Maluku); widesp. Oriental Reg.
     tesselatus, error for tessellatus.
     ssp. orientalis Swellengrebel & Swellengrebel-de Graaf, 1919: 3 (Neomyzomyia punctulata var.). Indonesia (Java & Maluku & Sulawesi).
     ssp. tessellatus s. str. Malaysia (Pen); Indonesia (Irian Jaya, Maluku); widesp. Oriental Reg.
vagus Dönitz, 1903: 80. Indonesia (Java & Sulawesi & Sumatra); Guam [immigrant], Indonesia (Maluku), Northern Marianas, ?PNG (PNG); Oriental Reg.

Nomen nudum in Cellia Theobald

melanotarsis Woodhill & Lee in Director General of Medical Services, 1943: 8.

Nomen nudum in Anopheles Meigen

solomonensis Cumpston, 1924: 1402.

Genus BIRONELLA Theobald

BIRONELLA Theobald, 1905c: 69. Type species: Bironella gracilis Theobald, 1905, mon.

Subgenus BIRONELLA Theobald

BIRONELLA Theobald, 1905c: 69 (as gen.). Type species: Bironella gracilis Theobald, 1905, mon.
gracilis Theobald, 1905c: 69. PNG (PNG); "Australia"; Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch). [Conserved by I.C.Z.N., 1965: 164.]
     bironelli Christophers, 1924: 16 (Anopheles; n. n. for gracilis Theobald). [Suppressed by I.C.Z.N., 1965: 124.]
     walchi Söesilo, 1932: 1035 (Anopheles). PNG (PNG).
     occulta Bonne-Wepster, 1951: 71. Indonesia (Irian Jaya).
simmondsi Tenorio, 1977: 331. Australia (Qld).

Subgenus BRUGELLA Edwards

BRUGELLA Edwards, 1930c: 288. Type species: Anopheles travestitus Brug, 1928, orig. des.
hollandi Taylor, 1934: 229. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (Bougainville I, PNG), Solomon Is.
obscura Tenorio, 1975: 734. PNG (PNG); Indonesia (Irian Jaya).
travestita Brug, 1928b: 921 (Anopheles). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).

Subgenus NEOBIRONELLA Tenorio

NEOBIRONELLA Tenorio, 1977: 345. Type species: Bironella confusa Bonne-Wepster, 1951, orig. des.
confusa Bonne-Wepster, 1951: 67. Indonesia (Irian Jaya); PNG (Bismarck Arch, PNG).
derooki Söesilo & Slooten, 1931a: 1256 [1931b: 126]. Indonesia (Irian Jaya); PNG (PNG).
     soesiloi Strickland & Chowdhury, 1931: 12 (Anopheles). Indonesia (Irian Jaya). N. syn.
papuae Swellengrebel & Swellengrebel-de Graaf, 1919: 3 (Stethomyia aitkenii var.) [1920: 36 (Stethomyia culiciformis var.)]. Indonesia (Irian Jaya); Australia (Qld), Indonesia (Maluku), PNG (Bismarck Arch, PNG)
     brugi Söesilo & Slooten, 1931a: 1256 [1931b: 126]. Indonesia (Irian Jaya).
     pupaae, error for papuae.

Genus AEDEOMYIA Theobald

AEDEOMYIA Theobald, 1901a: 235. Type species: Aedes squammipennis Lynch Arribálzaga, 1878, des. Neveu-Lemaire, 1902a: 223 and Sharp, 1902: 258.
AEDOMYIA, error for Aedeomyia.
catasticta Knab, 1909: 387. Philippines; Australia (NT, Qld, WA), Fiji, Guam, Indonesia (Irian Jaya, Maluku), Kiribati (Line Is), Northern Marianas, PNG (Bougainville I, PNG), Solomon Is; India, Indonesia (Java, Kalimantan, Nusa Tenggara, Sulawesi, Sumatra), SE Asia, Sri Lanka.
venustipes Skuse, 1889c: 1761 (Aedes). Australia (NSW); Australia (Qld, Vic).

Genus AEDES Meigen

AEDES Meigen, 1818: 13. Type species: Aedes cinereus Meigen, 1818, mon.

Subgenus AEDIMORPHUS Theobald

AEDIMORPHUS Theobald, 1903b: 290 (as gen.). Type species: Uranotaenia domestica Theobald, 1901, mon.
REEDOMYIA Ludlow, 1905: 94 (as gen.). Type species: Reedomyia pampangensis Ludlow, 1905, orig. des.
LEPTIDOTOMYIA Theobald, 1905c: 80 (as gen.). Type species: Lepidotomyia alboscutellata Theobald, 1905, mon.
GEITONOMYIA Leicester, 1908: 134 (as gen.). Type species: Culex caecus Theobald, 1901, mon.
alboscutellatus Theobald, 1905c: 80 (Lepidotomyia). PNG (PNG); Australia (NT, Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I), Solomon Is; Japan, Korea, widesp. Oriental Reg.
     pampangensis, authors, not Ludlow, 1905, misid.
     albiscutellatus, alboscutellaris, alboscutellatis, errors for alboscutellatus.
caecus Theobald, 1901b: 413 (Culex). Malaysia (Pen); Indonesia (Irian Jaya, Maluku), Northern Marianas, PNG (PNG); widesp. Oriental Reg.
     imprimens, Senior-White, 1923, not Walker, 1860, misid.
     coecus, error for caecus.
lowisii Theobald, 1910b: 257 (Reedomyia). India (Andaman Is); Indonesia (Maluku), PNG (PNG); Indonesia (Sulawesi), Philippines.
nocturnus Theobald, 1903b: 159 (Culex). Fiji; American Samoa, Australia (WA), Belau, ?Cook Is, Hawaiian Is, Indonesia (Irian Jaya, Maluku), Kiribati (Gilbert Is), Marshall Is, Micronesia, New Caledonia (Loyalty Is, New Caledonia), NZ (NZ), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is, Tonga, Vanuatu, Western Samoa.
     niger Theobald, 1913: 164 (Culex nocturnus var.). New Caledonia (New Caledonia).
     vexans, authors, not Meigen, 1830, misid.
oakleyi Stone, 1939: 163. Guam; Northern Marianas.
senyavinensis Knight & Hurlbut, 1949: 27. Micronesia.
trukensis Bohart, 1957: 63. Micronesia.
vexans Meigen—prob. not Australasian/Oceanian.


CHAETOCRUIOMYIA Theobald, 1910b: 195 (as gen.). Type species: Chaetocruiomyia sylvestris Theobald, 1910 [preocc., = Aedes spinosipes Edwards, 1922], mon.
calabyi Marks, 1963a: 204. Australia (WA); Australia (SA).
elchoensis Taylor, 1929: 276. Australia (NT); Australia (Qld).
humeralis Edwards, 1922b: 93. Australia (Qld).
macmillani Marks, 1964a: 138. Australia (NSW); Australia (Tas, Vic).
moloiensis Taylor, 1929: 276. Australia (Qld).
spinosipes Edwards, 1922b: 92 (n. n. for sylvestris Theobald). Australia (Qld).
     sylvestris Theobald, 1910b: 196 (Chaetocruiomyia). Australia (Qld). [Preocc. Theobald, 1901.]
tulliae Taylor, 1929: 275. Australia (Qld).
wattensis Taylor, 1929: 275. Australia (Qld); Australia (NSW, NT, SA, Vic).


CHRISTOPHERSIOMYIA Barraud, 1923a: 786 (as gen.). Type species: Stegomyia thomsoni Theobald, 1905, orig. des.
chionodes Belkin, 1962: 434. PNG (Bougainville I); Indonesia (Maluku), PNG (PNG).

Subgenus EDWARDSAEDES Belkin

EDWARDSAEDES Belkin, 1962: 408. Type species: Culex imprimens Walker 1860, orig. des.
imprimens Walker, 1860b: 144 (Culex). Indonesia (Maluku); Australia (Qld), Indonesia (Irian Jaya), PNG (PNG); Indonesia (Java), Korea, Malaysia (Pen), Philippines, Thailand.
     brugi Edwards, 1924d: 371. Indonesia (Irian Jaya).

Subgenus FINLAYA Theobald

FINLAYA Theobald, 1903b: 281 (as gen.). Type species: Finlaya poicilius Theobald, 1903, des. Sharp, 1904: 308.
DANIELSIA Theobald, 1904b: 78 (as gen.). Type species: Danielsia albotaeniata Leicester, 1904, mon.
HULECOETEOMYIA Theobald, 1904c: 163 (as gen.). Type species: Hulecoeteomyia trilineata Leicester, 1904 [preocc., = Howardina chrysolineata Theobald, 1907], mon.
MOLPEMYIA Theobald, 1910b: 479 (as gen.). Type species: Molpemyia purpurea Theobald, 1910, mon.
CALOMYIA Taylor, 1914b: 684 (as gen.). Type species: Calomyia priestleyi Taylor, 1914 [= Molpemyia purpurea Theobald, 1910], mon. [Preocc. Rossi, 1848.]
aculeatus Theobald, 1903b: 233 (Gilesia). Australia (Qld); Australia (NSW).
albilabris Edwards, 1925a: 258. Solomon Is; PNG (Bougainville I, PNG).
alboannulatus Macquart, 1850: 314(10) (Culex). Australia, "côte orientale"; Australia (Qld, SA, Tas, Vic, WA).
alocasicola Marks, 1947a: 35. Australia (Qld); Australia (NSW).
alticola Bonne-Wepster, 1948: 313. Indonesia (Irian Jaya).
anggiensis Bonne-Wepster, 1937: 97. Indonesia (Irian Jaya); PNG (PNG).
antipodeus Edwards, 1920: 132 (Ochlerotatus). NZ (NZ); Norfolk Is.
argenteitarsis Brug, 1932: 76. Indonesia (Irian Jaya).
argyronotum Belkin, 1962: 354. Solomon Is.
     papuensis, Belkin, 1962, not Taylor, 1914, misid.
aureostriatus Doleschall, 1857: 385 (Culex). Indonesia (Maluku); Indonesia (Irian Jaya); China, India, Philippines, Ryukyu Is, Sri Lanka, Taiwan.
auridorsum Edwards, 1922b: 93. Australia (Qld).
australiensis Theobald, 1910b: 313 (Leucomyia). Australia (Qld).
avistylus Brug, 1939: 107 (flavipennis var.). Indonesia (Sulawesi); Indonesia (Maluku); Malaysia (Sabah, Sarawak).
biocellatus Taylor, 1914d: 463 (Culex). Australia (NSW); Australia (Qld).
bougainvillensis Marks, 1947a: 19. PNG (Bougainville I); Solomon Is.
britteni Marks & Hodgkin, 1958: 33. Australia (WA); Australia (NT).
burnetti Belkin, 1962: 363. Fiji.
buxtoni Belkin, 1962: 356. PNG (Bougainville I).
candidoscutellum Marks, 1947b: 1. Australia (Qld); Australia (ACT, NSW), Indonesia (Irian Jaya), PNG (PNG).
clintoni Taylor, 1946: 211. PNG (PNG).
derooki Brug, 1932: 75. Indonesia (Maluku).
dobodurus King & Hoogstraal, 1946a: 37. PNG (PNG); Indonesia (Irian Jaya).
dobrotworskyi Marks, 1958b: 65. Australia (Vic).
dorseyi Knight, 1946: 277. Belau.
edgari Stone & Rosen, 1952: 425. French Polynesia (Society Is).
fijiensis Marks, 1947a: 26. Fiji.
flavipennis Giles, 1904a: 366 (Finlaya). Philippines; Indonesia (Irian Jaya); Indonesia (Java).
franclemonti Belkin, 1962: 367. Solomon Is.
freycinetiae Laird, 1957: 343. Fiji.
fuscipalpis Belkin, 1962: 370. Solomon Is.
fuscitarsus Belkin, 1962: 371. Solomon Is.
gahnicola Marks, 1947a: 43. Australia (Qld); Australia (NSW).
gani Bonne-Wepster, 1940: 158. Indonesia (Irian Jaya).
gracilelineatus Bonne-Wepster, 1937: 93. Indonesia (Irian Jaya); PNG (PNG).
gressitti Bohart, 1957: 23. Belau.
hollandius King & Hoogstraal, 1946a: 38. Indonesia (Irian Jaya); PNG (PNG).
hollingsheadi Belkin, 1962: 372. Solomon Is; PNG (Bougainville I).
horotoi Taylor, 1972: 317. Solomon Is.
hui Bohart, 1957: 25. Belau.
inexpectatus Bonne-Wepster, 1948: 318 (Aedimorphus). Indonesia (Irian Jaya); PNG (PNG).
     inexpectabilis, error for inexpectatus.
iwi Marks, 1955: 11. Australia (Qld).
josephinae Marks, 1958b: 58. Australia (Qld); Indonesia (Irian Jaya), ?PNG (PNG).
keefei King & Hoogstraal, 1946f: 311. PNG (PNG); Australia (Qld).
knighti Stone & Bohart, 1944: 210. Solomon Is.
kochi Dönitz, 1901: 38 (Culex). PNG (PNG); Australia (NT, Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch); ?Indonesia (Timor).
lauriei Carter, 1921a: 623 (Ochlerotatus; as laurei). Lord Howe I.
lewelleni Starkey & Webb, 1946: 179. Belau.
mackerrasi Taylor, 1927: 68. Australia (Qld).
maffii Taylor & Tenorio, 1974: 577. Solomon Is.
mallochi Taylor, 1944: 121. Australia (Qld); Australia (NSW, NT, SA, Vic).
mcdonaldi Belkin, 1962: 396. Solomon Is; PNG (Bougainville I, PNG).
milsoni Taylor, 1915a: 179 (Culicada). Australia (NSW); Australia (Qld, Vic).
     milsoni Taylor, 1916b: 566 (Hulecoeteomyia). Australia (NSW). [Preocc. Taylor, 1915.]
monocellatus Marks, 1948: 14. Australia (Qld); Australia (NSW).
neogeorgianus Belkin, 1962: 375. Solomon Is.
notoscriptus Skuse, 1889c: 1738 (Culex). Australia (NSW); widesp. Australia, Indonesia (Irian Jaya, Maluku), New Caledonia (Loyalty Is, New Caledonia), NZ (NZ), PNG (Bismarck Arch, PNG); Oriental Reg.
novalbitarsis King & Hoogstraal, 1946c: 146 (n. n. for albitarsis Taylor). PNG (PNG); Indonesia (Irian Jaya).
     albitarsis Taylor, 1914c: 194 (Leucomyia). PNG (PNG). [Preocc. Ludlow, 1905.]
occidentalis Skuse, 1889c: 1729 (Culex). Australia (WA); ?Australia (NT).
oceanicus Belkin, 1962: 377. American Samoa; Tonga, Western Samoa.
palmarum Edwards, 1924d: 382. Australia (Qld).
     novalbitarsis, authors, not King & Hoogstraal, 1946, misid.
papuensis Taylor, 1914c: 193 (Leucomyia australiensis var.). PNG (PNG); Indonesia (Irian Jaya); PNG (Bismarck Arch).
pecuniosus Edwards, 1922b: 94 (Aedes). Australia (NT); Australia (WA).
plagosus Marks, 1959b: 21. Australia (NSW).
plumiferus King & Hoogstraal 1946f: 306. Indonesia (Irian Jaya); PNG (PNG).
poicilius Theobald 1903b: 283 (Finlaya). Malaysia (Pen); ?Indonesia (Irian Jaya), ?PNG (PNG); SE Oriental Reg.
purpuraceus Brug, 1932: 77. Indonesia (Irian Jaya).
purpureus Theobald, 1910b: 479 (Molpemyia). Australia (Qld).
     priestleyi Taylor, 1914d: 684 (Calomyia). Australia (Qld).
quasirubithorax Theobald, 1910b: 327 (Culex). Australia (Qld); Australia (NSW, NT), Indonesia (Irian Jaya), PNG (PNG).
     quasirubrithorax, error for quasirubithorax.
quinquelineatus Edwards, 1922b: 93. Australia (Qld); Australia (NSW).
roai Belkin, 1962: 353. Solomon Is.
rubiginosus Belkin, 1962: 346. ?Solomon Is; PNG (Bismarck Arch).
rubrithorax Macquart, 1850: 313(9) (Culex). Australia (Tas); widesp. Australia (excl. NT).
     similis Strickland, 1911: 132 (Culicelsa). Australia (SA). [Preocc. Theobald, 1910.]
     queenslandis Strickland, 1911: 179 (Culicelsa). Australia (Qld).
     demansis Strickland, 1911: 202 (Culicada). Australia (Tas).
     cumpstoni Taylor, 1914b: 692 (Culicada). Australia (Vic).
     hybrida Taylor, 1916b: 568 (Culicada). Australia (NSW).
     rubithorax, error for rubrithorax.
rupestris Dobrotworsky, 1959: 136. Australia (Vic); Australia (ACT, NSW, Qld, Tas).
samoanus Grünberg, 1913: 130 (Finlaya). Western Samoa; American Samoa.
savoryi Bohart, 1957: 31. Bonin Is.
schlosseri Belkin, 1962: 382. Solomon Is.
shehzadae Qutubuddin, 1972: 119. PNG (PNG).
solomonis Stone & Bohart, 1944: 208. Solomon Is; PNG (Bougainville I).
stanleyi Peters, 1963a: 1. PNG (PNG).
subalbitarsis King & Hoogstraal, 1946c: 144. Indonesia (Irian Jaya); PNG (PNG).
subauridorsum Marks, 1948: 28. Australia (Qld); Australia (NSW, Vic).
subbasalis Dobrotworsky, 1963: 298. Australia (ACT); Australia (NSW, Qld, Vic).
togoi Theobald, 1907: 379 (Culicelsa). Japan; Bonin Is, Marcus I, ?Northern Marianas, Volcano Is; Canada, USA, E Asia from Siberia to Malaysia (Pen).
toxopeusi Bonne-Wepster, 1948: 315. Indonesia (Irian Jaya).
tsiliensis King & Hoogstraal, 1946f: 305. PNG (PNG).
tubbutiensis Dobrotworsky, 1959: 139. Australia (Vic).
tutuilae Ramalingam & Belkin, 1965: 3. American Samoa; Western Samoa.
wallacei Edwards, 1926e: 105. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG).
wasselli Marks, 1947b: 9. Australia (Qld).

Subgenus GEOSKUSEA Edwards

GEOSKUSEA Edwards, 1929g: 342. Type species: Aedes fimbripes Edwards, 1924, orig. des.
becki Belkin, 1962: 334. Solomon Is.
daggyi Stone & Bohart, 1944: 215. Vanuatu.
fimbripes Edwards, 1924d: 390. PNG (Bismarck Arch); PNG (Bougainville I, PNG).
longiforceps Edwards, 1929f: 315. Solomon Is; PNG (Bougainville I).
lunulatus King & Hoogstraal, 1946b: 95 (Pseudoskusea). Indonesia (Irian Jaya); PNG (PNG).
perryi Belkin, 1962: 336. Solomon Is.
tonsus Edwards, 1924d: 390. Indonesia (Maluku).

Subgenus HALAEDES Belkin

CAENOCEPHALUS Taylor, 1914b: 700 (as gen.). Type species: Caenocephalus concolor Taylor, 1914 [= Culex australis Erichson, 1842], mon. [Preocc. Wulp, 1898.]
HALAEDES Belkin, 1962: 328. Type species: Culex australis Erichson, 1842, orig. des.
ashworthi Edwards, 1921b: 75. Australia (WA).
australis Erichson, 1842: 270 (Culex). Australia (Tas); Australia (ACT, NSW, SA, Vic), Lord Howe I, NZ (NZ), Norfolk I.
     crucians Walker, 1856d: 432 (Culex). Australia (Tas).
     tasmaniensis Strickland, 1911: 181 (Culicada). Australia (Tas).
     concolor Taylor, 1914b: 700 (Caenocephalus). Australia (NSW).

Subgenus HUAEDES Huang

HUAEDES Huang, 1968a: 169. Type species: Aedes wauensis Huang, 1968, orig. des.
medialis Brug, 1932: 79. Indonesia (Irian Jaya).
variepictus King & Hoogstraal, 1946b: 98. Indonesia (Irian Jaya).
wauensis Huang, 1968a: 170. PNG (PNG).


LEPTOSOMATOMYIA Theobald, 1905c: 110 (as gen.). Type species: Leptosomatomyia lateralis Theobald, 1905 [preocc., = Aedes aurimargo Edwards, 1922], mon.
aurimargo Edwards, 1922b: 94. Australia (Qld); Indonesia (Irian Jaya), PNG (PNG).
     lateralis Theobald, 1905c: 110 (Leptosomatomyia). PNG (PNG). [Preocc. Meigen, 1818.]
     litoralis, error for lateralis.

Subgenus LEVUA Stone & Bohart

LEVUA Stone & Bohart, 1944: 214. Type species: Aedes suvae Stone & Bohart, 1944 [= Aedes geoskusea Amos, 1944], orig. des.
geoskusea Amos, 1944: 32. Fiji.
     suvae Stone & Bohart, 1944: 214. Fiji.

Subgenus LORRAINEA Belkin

LORRAINEA Belkin, 1962: 430. Type species: Aedes dasyorrhus King & Hoogstraal, 1946, orig. des.
amesii Ludlow—not Australasian/Oceanian.
dasyorrhus King & Hoogstraal, 1946b: 100. Indonesia (Irian Jaya); Australia (?NT, Qld), PNG (PNG), ?Solomon Is, Vanuatu.
fumidus Edwards, 1928a: 274. Singapore; ?Indonesia (Maluku); Indonesia (Sulawesi, Sumatra), Malaysia (Pen), Philippines.
lamelliferus Bohart & Ingram, 1946: 29. Micronesia; ?Belau.

Subgenus MACLEAYA Theobald

MACLEAYA Theobald, 1903a: 154 (as gen.). Type species: Macleaya tremula Theobald, 1903, mon.
littlechildi Taylor, 1933: 370. PNG (PNG); Australia (Qld).
stoneorum Marks, 1977: 33. Australia (Qld); Australia (NT, WA).
tremulus Theobald, 1903a: 155 (Macleaya). Australia (Qld); Australia (NT, SA, Vic, WA), Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).
     minuta Taylor, 1912b: 30 (Danielsia). Australia (NT). [Preocc. Theobald, 1910.]
     alboannulata Taylor, 1912b: 31 (Danielsia). Australia (NT). [Preocc. Macquart, 1850.]
     australis Taylor, 1914d: 457 (Aedimorphus). Australia (Qld). [Preocc. Erichson, 1842.]
     darwini Taylor, 1914d: 458 (Aedimorphus australis var.). Australia (NT).
     doddi Taylor, 1919a: 831 (Mimetomyia). PNG (PNG).

Subgenus MUCIDUS Theobald

MUCIDUS Theobald, 1901a: 235 (as gen.). Type species: Mucidus africanus Theobald, 1901 [preocc., = Mucidus grahamii Theobald, 1909], subs. mon. Theobald, 1901d: ii.
PARDOMYIA Theobald, 1907: 280 (as gen.). Type species: Pardomyia aurantia Theobald, 1907, mon.
alternans Westwood, 1836: 681 (Culex). "Nov Hollandi "; Australia (NSW, NT, Qld, Vic, WA), Indonesia (Irian Jaya), New Caledonia (Loyalty Is, New Caledonia), PNG (PNG), Vanuatu; Indonesia (Timor).
     commovens Walker, 1856d: 432 (Culex). "New Holland."
     hispidosus Skuse, 1889c: 1726 (Culex). Australia (NSW).
     kermorganti Laveran, 1901: 568 (Culex). New Caledonia (New Caledonia).
aurantius Theobald, 1907: 280 (Pardomyia). Malaysia (Sarawak); Australia (Qld), Indonesia (Irian Jaya), PNG (PNG); Indonesia (Kalimantan, Sumatra), Malaysia (Pen), SE Asia.
     ssp. aurantius s. str. Malaysia (Sarawak); Indonesia (Irian Jaya); Indonesia (Kalimantan, Sumatra), SE Asia.
     auranticus, error for aurantius.
     ssp. chrysogaster Taylor, 1927: 67 (Mucidus; as sp.). Australia (Qld); Indonesia (Irian Jaya), PNG (PNG).
painei Knight, 1948: 68 (aurantius ssp.; n. n. for nigrescens Edwards). Solomon Is; Indonesia (Irian Jaya), PNG (Bismarck Arch, Bougainville I).
     nigrescens Edwards, 1929f: 314 (Pardomyia aurantia var.). Solomon Is. [Preocc. Theobald, 1907.]


NEOMELANICONION Newstead in Newstead, Dutton & Todd, 1907: 31 (as Neomelaniconion palpaleNewstead, 1907, mon.
lineatopennis Ludlow, 1905: 133 (Taeniorhynchus). Philippines; Australia (NT, Qld, WA), Indonesia (Maluku); Indonesia (Java, Sulawesi, Sumatra, Timor), widesp. Afrotrop., Oriental Regs.
     linealis Taylor, 1913: 57 (Pseudohowardina). Australia (Qld).

Subgenus NOTHOSKUSEA Dumbleton

NOTHOSKUSEA Dumbleton, 1962: 20. Type species: Aedes chathamicus Dumbleton, 1962, orig. des.
chathamicus Dumbleton, 1962: 20. NZ (Chatham Is).

Subgenus OCHLEROTATUS Lynch Arribálzaga

OCHLEROTATUS Lynch Arribálzaga, 1891a: 353, 367 [1891b: 143] (as gen.). Type species: Ochlerotatus confirmatus Lynch Arribálzaga, 1891 [= Aedes scapularis Rondani, 1848], des. Coquillett, 1910: 577.
OCHLEROTHATUS. Incorr. orig. spell. of Ochlerotatus (Lynch Arribálzaga, 1891a: 374).
TAENIORHYNCHUS Lynch Arribálzaga, 1891a: 374 [1891b: 147] (as gen.). Suppressed by I.C.Z.N., 1959c: 187.
GILESIA Theobald, 1903b: 233 (as gen.). Type species: Gilesia aculeata Theobald, 1903, mon.
CULICELSA Felt, 1904: 391b (as gen.). Type species: Culex taeniorhynchus Wiedemann, 1821, orig. des.
CULICADA Felt, 1904: 391b (as gen.). Type species: Culex canadensis Theobald, 1901, orig. des.
ANDERSONIA Strickland, 1911: 250 (as gen.). Type species: Andersonia tasmaniensis Strickland, 1911 [preocc., = Aedes andersoni Edwards, 1926], mon. [Preocc. Boulenger, 1900.]
andersoni Edwards, 1926e: 112 (n. n. for tasmaniensis Strickland). Australia (Tas); Australia (NSW, Vic).
     tasmaniensis Strickland, 1911: 250 (Andersonia). Australia (Tas). [Preocc. Strickland, 1911.]
     tasmaniensis Taylor, 1914d: 466 (Menolepis). Australia (Tas). [Preocc. Strickland, 1911.]
burpengaryensis Theobald, 1905d: 26 (Culicada; as bupengaryensis). Australia (Qld); Australia (NSW).
cacozelus Marks, 1963b: 45. Australia (WA).
calcariae Marks, 1957b: 74. Australia (SA); Australia (Tas, Vic).
camptorhynchus Thomson, 1869: 443 (Culex). Australia (NSW); Australia (SA, Tas, Vic, WA).
     labeculosus Coquillett, 1905a: 116 (Culex). Australia (Vic).
     westralis Strickland, 1911: 130 (Culicelsa). Australia (WA).
     inornata Strickland, 1911: 201 (Culicada). Australia (Tas).
     nigra Taylor, 1914b: 688 (Culicada). Australia (Tas). [Preocc. Giles, 1904.]
     annulipes Taylor, 1914b: 693 (Culicada). Australia (Tas). [Preocc. Meigen, 1830.]
     victoriensis Taylor, 1914d: 460 (Culicada). Australia (Vic).
clelandi Taylor, 1914b: 690. Australia (Tas); Australia (SA, Vic, WA).
continentalis Dobrotworsky, 1960a: 71. Australia (Vic); Australia (SA, Tas).
cunabulanus Edwards, 1924d: 378. Australia (Tas).
eidsvoldensis Mackerras, 1927b: 295 (theobaldi f.). Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
     eidsvolaensis, error for eidsvoldensis.
explorator Marks, 1964b: 60. Australia (NT).
flavifrons Skuse, 1889c: 1735 (Culex). Australia (NSW & Qld); Australia (SA, Tas, Vic).
     vandema Strickland, 1911: 202 (Culicada). Australia (Tas).
     variegatans Strickland, 1911: 204 (Culicada vandema var.). Australia (Tas).
hesperonotius Marks, 1959a: 131. Australia (WA).
hodgkini Marks, 1959a: 112. Australia (WA).
imperfectus Dobrotworsky, 1963: 296. Australia (Vic); Australia (Qld, NSW).
linesi Marks, 1964b: 63. Australia (SA).
luteifemur Edwards, 1926e: 112. Australia (Tas); Australia (SA, Vic).
macintoshi Marks, 1959a: 117. Australia (WA).
nigrithorax Macquart, 1847a: 9 [1847b: 25] (Culex). Australia (Tas); Australia (NSW, SA, Vic).
     macleayanus Mackerras, 1927b: 291. Australia (Tas).
nivalis Edwards, 1926e: 112. Australia (Vic); Australia (NSW, Qld, Tas).
normanensis Taylor, 1915a: 182 (Culex). Australia (Qld); Australia (NT, WA).
perkinsi Marks, 1949: 32. Australia (Qld); Australia (Vic).
phaecasiatus Marks, 1964b: 64. Australia (NT); Australia (Qld).
procax Skuse, 1889c: 1742 (Culex). Australia (NSW); Australia (Qld, Vic).
pseudonormanensis Marks, 1949: 26. Australia (Qld); Australia (NSW, NT, WA).
purpureifemur Marks, 1959a: 132. Australia (WA).
purpuriventris Edwards, 1926e: 113. Australia (Tas); Australia (SA, Vic).
ratcliffei Marks, 1959a: 123. Australia (WA).
sagax Skuse, 1889c: 1744 (Culex). Australia (NSW); Australia (Qld, SA, Vic, WA).
     wilsoni Taylor, 1919a: 833 (Culicada). Australia (Vic).
sapiens Marks, 1964b: 66. Australia (NSW); Australia (Qld, WA).
silvestris Dobrotworsky, 1961a: 248 (n. n. for waterhousei Dobrotworsky). Australia (Vic); Australia (ACT, NSW, SA, Tas).
     waterhousei Dobrotworsky, 1960a: 57. Australia (Vic). [Preocc. Theobald, 1905.]
spilotus Marks, 1963b: 31. Australia (Vic); Australia (SA).
stricklandi Edwards, 1912b: 523. Australia (WA); Australia (SA, Tas), NZ (NZ).
     australis Strickland, 1911: 133 (Grabhamia). Australia (WA). [Preocc. Erichson, 1842.]
     flindersi Taylor, 1914b: 686 (Grabhamia). Australia (Tas).
subalbirostris Klein & Marks, 1960: 115. NZ (NZ).
theobaldi Taylor, 1914a: 751 (Grabhamia). Australia (Qld); Australia (NSW, ?NT, SA, Vic).
     flavifrons, Theobald, 1907, not Skuse, 1889, misid.
turneri Marks, 1963b: 42. Australia (WA).
vigilax Skuse, 1889c: 1731 (Culex). Australia (NSW & Qld); Australia (NT, SA, Vic, WA), Fiji, Indonesia (Irian Jaya), New Caledonia (Loyalty Is, New Caledonia), PNG (PNG), Solomon Is, Vanuatu; Ryukyu Is, Seychelles, widesp. Oriental Reg. [Conserved by I.C.Z.N., 1972: 11.]
     albirostris Macquart, 1850: 314(10) (Culex). [Suppressed by I.C.Z.N., 1972: 11.]
     marinus Theobald, 1901b: 396 (Culex). Australia (Qld).
     pseudovigilax Theobald, 1907: 382 (Culicelsa). Australia (Qld).
     uniformis Strickland, 1911: 131 (Culicelsa). Australia (WA). [Preocc. Theobald, 1910.]
vittiger Skuse, 1889c: 1728 (Culex). Australia (NSW); Australia (NT, Qld, SA, Vic).

Subgenus PARAEDES Edwards

PARAEDES Edwards in Barraud, 1934: 446 (as gen.). Type species: Paraedes barraudi Edwards, 1934, orig. des.
ostentatio Leicester, 1908: 193 (Aioretomyia). Malaysia (Pen); Indonesia (Maluku); Indonesia (Java, Kalimantan, Sulawesi, Sumatra), widesp. Oriental Reg.

Subgenus PSEUDOSKUSEA Theobald

PSEUDOSKUSEA Theobald, 1907: 192 (as gen.). Type species: Skusea multiplex Theobald, 1903, mon.
bancroftianus Edwards,1921c: 74. Australia (Qld); Australia (NSW, NT, SA, Vic, WA).
culiciformis Theobald, 1905c: 77 (Skusea). PNG (PNG); Australia (NT, Qld), PNG (Bismarck Arch).
multiplex Theobald, 1903b: 293 (Skusea). Australia (Qld); Australia (NSW, Vic).
postspiraculosis Dobrotworsky, 1961b: 261. Australia (Vic); Australia (NSW, SA, Tas).
     postspiraculosus, error for postspiraculosis.

Subgenus RHINOSKUSEA Edwards

RHINOSKUSEA Edwards, 1929g: 342. Type species: Ficalbia longirostris Leicester, 1908, orig. des.
longirostris Leicester, 1908: 228 (Ficalbia). Malaysia (Pen); Australia (NT, Qld), PNG (PNG); Indonesia (Java, Sulawesi, Sumatra), widesp. Oriental Reg.
     hilli Taylor, 1919a: 841 (Uranotaenia). Australia (NT).

Subgenus SCUTOMYIA Theobald

SCUTOMYIA Theobald, 1904b: 77 (as gen.). Type species: Scutomyia albolineatus Theobald, 1904, mon.
albolineatus Theobald, 1904b: 77 (Scutomyia). Malaysia (Pen); Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG), Solomon Is; SE Oriental Reg.

Subgenus STEGOMYIA Theobald

STEGOMYIA Theobald in Howard, 1901: 235 [Theobald, 1901d: iii] (as gen.). Type species: Culex fasciatus Fabricius, 1805 [= Culex aegypti Linnaeus, 1762], des. Neveu-Lemaire, 1902a: 212.
PSEUDOSTEGOMYIA Theobald, 1910b: 10 (as gen.). Type species: Stegomyia gardinerii Ludlow, 1905, mon.
aegypti Linnaeus in Hasselquist, 1762: 470 (Culex). Egypt [original type locality], Malaysia (Pen) [neotype locality; see I.C.Z.N., 1964: 246]; American Samoa, Australia (NSW, NT, Qld, WA), Bonin Is, Cook Is, Fiji, widesp. French Polynesia, Guam, Hawaiian Is, Indonesia (Irian Jaya, Maluku), Kiribati, Marcus I, Marshall Is, Micronesia, Nauru, New Caledonia (New Caledonia), Niue, Northern Marianas, PNG (Bismarck Arch, Bougainville I, PNG), Pitcairn Is, Solomon Is, Tokelau, Tonga, Tuvalu, Vanuatu, Volcano Is, Wake I, Wallis & Futuna, Western Samoa; cosmopol. between 10 °C winter isotherms. [Conserved by I.C.Z.N., 1964: 246.]
     fasciatus Fabricius, 1805: 36 (Culex). "Americae insulis" [= West Indies].
     bancrofti Skuse, 1889c: 1740 (Culex). Australia (Qld).
     queenslandensis Theobald, 1901b: 297 (Stegomyia fasciata var.). Australia (Qld).
     pulcherrima Taylor 1919a: 830 (Mimetomyia). Australia (Qld).
agrihanensis Bohart, 1957: 39. Northern Marianas.
albopictus Skuse, 1894: 20 (Culex). India; Australia (NT, Qld), French Polynesia (Tuamotu Arch) [introduced; not established], Guam, Hawaiian Is, Indonesia (Irian Jaya, Maluku), Marshall I, Micronesia, Northern Marianas, PNG (PNG), Solomon Is; Brazil [immigrant], Korea, Ryukyu Is, USA [immigrant], widesp. Afrotrop., Oriental Regs.
alorensis Bonne-Wepster & Brug, 1932: 92 (variegatus var.). Indonesia (Maluku).
annandalei Theobald, 1910a: 10 (Stegomyia). India; Indonesia (Irian Jaya, Maluku), ?PNG (PNG); China, Taiwan, SE Oriental Reg.
aobae Belkin, 1962: 452. Vanuatu.
cooki Belkin, 1962: 454. Niue; Tonga.
dybasi Bohart, 1957: 53. Belau; Guam.
futunae Belkin, 1962: 455. Wallis & Futuna.
gardinerii Ludlow, 1905a: 99 (Stegomyia). Philippines; Indonesia (Maluku); Indonesia (Nusa Tenggara, Sulawesi), Malaysia (Sabah).
guamensis Farner & Bohart, 1944: 117. Guam; Northern Marianas.
gurneyi Stone & Bohart, 1944: 217. PNG (Bougainville I); Solomon Is.
hakanssoni Knight & Hurlburt, 1949: 23. Micronesia.
hebrideus Edwards, 1926e: 102. Vanuatu; PNG (PNG), Solomon Is.
hensilli Farner, 1945: 59. Micronesia; Belau.
     scutellaris, authors, not Walker, 1851, ?misid.
hoguei Belkin, 1962: 460. Solomon Is.
horrescens Edwards, 1935a: 128. Fiji.
katherinensis Woodhill, 1949: 141 (scutellaris ssp.). Australia (NT); Australia (Qld, WA).
kesseli Huang & Hitchcock, 1980: 23. Tonga.
maehleri Bohart, 1957: 40. Micronesia.
marshallensis Stone & Bohart, 1944: 218. Marshall Is; ?Guam, Kiribati (Gilbert Is), Micronesia.
neopandani Bohart, 1957: 36. Northern Marianas; Guam.
palauensis Bohart, 1957: 51. Belau.
pandani Stone, 1939: 162. Guam.
paullusi Stone & Farner, 1945: 155. Philippines; Indonesia (Irian Jaya, Maluku); Malaysia (Pen, Sabah).
pernotatus Farner & Bohart, 1944: 118. Vanuatu.
polynesiensis Marks, 1951: 138. Fiji; American Samoa, Cook Is, Easter I, French Polynesia (Austral Is, Marquesas, Society Is, Tuamotu Arch), Pitcairn Is, Tokelau, Tuvalu, Wallis & Futuna, Western Samoa.
pseudalbopictus Borel—not Australasian/Oceanian.
pseudoscutellaris Theobald, 1910c: 156 (Stegomyia). Fiji.
quasiscutellaris Farner & Bohart, 1944: 120. Solomon Is; PNG (Bougainville I).
robinsoni Belkin, 1962: 447. Solomon Is; Vanuatu.
rotanus Bohart & Ingram, 1946: 39. Northern Marianas; Guam.
rotumae Belkin, 1962: 472. Fiji.
saipanensis Stone, 1945: 66. Northern Marianas; Guam.
scutellaris Walker, 1858b: 77 (Culex). Indonesia (Maluku); Australia (Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG); Indonesia (Timor).
     variegatus Doleschall, 1858: 77 (Culex). Indonesia (Maluku). [Preocc. Schrank, 1781.]
     zonatipes Walker, 1861b: 229 (Culex). Indonesia (Irian Jaya).
scutoscriptus Bohart & Ingram, 1946: 27. Micronesia.
tabu Ramalingam & Belkin, 1965: 1. Tonga.
tongae Edwards, 1926e: 103. Tonga.
tulagiensis Edwards, 1926e: 109. Solomon Is; Vanuatu.
upolensis Marks, 1957a: 51. Western Samoa; American Samoa.
varuae Belkin, 1962: 478. Solomon Is; Vanuatu.
     tongae, authors, not Edwards, 1926, misid.
wadai Tanaka, Mizusawa & Saugstad, 1979: 400. Bonin Is.

Unplaced species of AEDES Meigen

burnsi Basio & Reisen, 1971: 57. Guam. Nomen dubium.
butleri Theobald, 1901c: 230. Malaysia (Pen); Indonesia (Maluku); Indonesia (Java, Kalimantan, Sulawesi, Sumatra), Oriental Reg.
cunninghami Taylor, 1944: 123 (n. n. for bancrofti Taylor). Australia (Qld).
     bancrofti Taylor, 1914d: 465 (Skusea). Australia (Qld). [Preocc. Skuse, 1889.]
daliensis Taylor, 1916b: 565 (Stegomyia). Australia (NT).

Genus ARMIGERES Theobald

ARMIGERES Theobald, 1901a: 235. Type species: Culex obturbans Walker, 1859, des. Brunetti, 1914: 55.

Subgenus ARMIGERES Theobald

ARMIGERES Theobald, 1901a: 235 (as gen.). Type species: Culex obturbans Walker, 1859, des. Sharp, 1902: 259.
DESVOIDYA Blanchard, 1901: 1046 (as gen.; unnec. n. n. for Armigeres). [Preocc. Meade, 1892.]
BLANCHARDIOMYIA Brunetti, 1912a: 440 (as gen.; n. n. for Desvoidya).
NEOSQUAMOMYIA Taylor, 1914c: 186 (as gen.). Type species: Neosquamomyia breinli Taylor, 1914, mon.
breinli Taylor, 1914c: 186 (Neosquamomyia). PNG (PNG); Indonesia (Irian Jaya), PNG (Bismarck Arch, Bougainville I), Solomon Is, Vanuatu.
denbesteni Brug, 1925: 345. Indonesia (Maluku); Indonesia (Irian Jaya).
     spathulatus Brug, 1939: 101. Indonesia (Maluku).
lacuum Edwards, 1922b: 97. PNG (Bismarck Arch); ?Indonesia (Maluku).
malayi Theobald, 1901b: 258 (Uranotaenia). Malaysia (Pen); Indonesia (Irian Jaya, Maluku), ?PNG (PNG); Oriental Reg.
milnensis Lee, 1944b: 215. PNG (PNG); Indonesia (Irian Jaya).
papuensis Peters, 1963a: 4. PNG (PNG); Indonesia (Irian Jaya).
subalbatus Coquillett, 1898: 302 (Culex). Japan; Guam; E Oriental, E Palearctic Regs.

Subgenus LEICESTERIA Theobald

LEICESTERIA Theobald, 1904d: 211 (as gen.). Type species: Leicesteria longipalpis Leicester, 1904, mon.
BREVIRHYNCHUS Theobald, 1908: 293 (as gen.). Type species: Brevirhynchus magnus Theobald, 1908, mon.
annulipalpis Theobald, 1910b: 6 (Brevirhynchus). India; Indonesia (Maluku); widesp. Oriental Reg.

Unplaced species of ARMIGERES Theobald

ventralis Walker, 1860b: 144 (Culex). Indonesia (Maluku).


HEIZMANNIA Ludlow, 1905a: 130. Type species: Heizmannia scintillans Ludlow, 1905, mon.
HEINZMANNIA Brunetti, 1914: 61 (unjust. emend. of Heizmannia).
aurea Brug, 1932: 75. Indonesia (Maluku).

Genus OPIFEX Hutton

OPIFEX Hutton, 1902b: 188. Type species: Opifex fuscus Hutton, 1902, mon.
fuscus Hutton, 1902b: 188. NZ (NZ); NZ (Kermadec Is).

Genus VERRALLINA Theobald

VERRALLINA Theobald, 1903b: 295. Type species: Aedes butleri Theobald, 1901, des. Blanchard, 1905: 417.

Subgenus NEOMACLEAYA Theobald

NEOMACLEAYA Theobald, 1907: 238 (as gen.). Type species: Neomacleaya indica Theobald, 1907, mon.
Laffoon, 1946: 240 (Aedes). Philippines; Indonesia (Irian Jaya).
neomacrodixoa King & Hoogstraal, 1947c: 124 (Aedes). Indonesia (Irian Jaya); Indonesia (Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia (Sulawesi).
panayensis Ludlow, 1914c: 159 (Aedes). Philippines; Indonesia (Irian Jaya, Maluku); Indonesia (Kalimantan, Nusa Tenggara, Sulawesi), Malaysia (Sabah).

Subgenus VERRALLINA Theobald

VERRALLINA Theobald, 1903b: 295 (as gen.). Type species: Aedes butleri Theobald, 1901, des. Blanchard, 1905: 417.
AIORETOMYIA Leicester, 1908: 185 (as gen.). Type species: Aioretomyia varietas Leicester, 1908, des. Brunetti, 1914: 55
azureosquamata Bonne-Wepster, 1948: 320 (Aedes). Indonesia (Irian Jaya).
bifoliata King & Hoogstraal, 1947c: 120 (Aedes). Indonesia (Irian Jaya); PNG (PNG).
carmenti Edwards, 1924d: 388 (Aedes). Solomon Is; Australia (NT, Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is.
     ceramensis Brug, 1934: 512 (Aedes). Indonesia (Maluku).
cuccioi Belkin, 1962: 414 (Aedes). Solomon Is; PNG (Bougainville I, PNG).
embiensis Huang, 1968b: 24 (Aedes). PNG (PNG).
foliformis King & Hoogstraal, 1947c: 121 (Aedes). PNG (PNG); Indonesia (Irian Jaya).
funerea Theobald, 1903b: 292 (Skusea). Australia (Qld); Australia (NSW, NT), Indonesia (Irian Jaya), PNG (PNG).
     basalis Taylor, 1912a: 27 (Pseudoskusea). Australia (Qld).
killertonis Huang, 1968b: 31. PNG (PNG).
leilae King & Hoogstraal, 1947c: 121 (Aedes). Indonesia (Irian Jaya); Australia (Qld).
lineata Taylor, 1914c: 191 (Lepidotomyia). PNG (PNG); Australia (Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I), Solomon Is, Vanuatu; Indonesia (Nusa Tenggara, Timor).
     ornata Theobald, 1905c: 79 (Skusea funerea var.). PNG (PNG). [Preocc. Meigen, 1818.]
mccormicki Belkin, 1962: 420 (Aedes). Solomon Is; PNG (Bismarck Arch, Bougainville I).
milnensis King & Hoogstraal, 1947c: 122 (Aedes). PNG (PNG).
multifolium King & Hoogstraal, 1947c: 122 (Aedes). Indonesia (Irian Jaya) Australia (Qld), PNG (PNG).
obsoleta Huang, 1968b: 45 (Aedes). PNG (PNG).
parasimilis King & Hoogstraal, 1947c: 125 (Aedes). Indonesia (Irian Jaya); Indonesia (Maluku), PNG (PNG).
pipkini Bohart, 1957: 65 (Aedes). Micronesia; Belau.
quadrifolium Brug, 1934: 512 (Aedes). Indonesia (Irian Jaya); ?Australia (Qld), PNG (PNG).
quadrispinata King & Hoogstraal, 1947c: 126 (Aedes). Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).
reesi King & Hoogstraal, 1947c: 127 (Aedes). PNG (PNG); Australia (NT, Qld, WA), Indonesia (Irian Jaya).
sentania King & Hoogstraal, 1947c: 128 (Aedes). Indonesia (Irian Jaya); PNG (PNG).
similis Theobald, 1910b: 189 (Pseudoskusea). Australia (Qld); Indonesia (Irian Jaya, Maluku); PNG (PNG).
simplus King & Hoogstraal, 1947c: 130 (Aedes). Indonesia (Irian Jaya).
trispinata King & Hoogstraal, 1947c: 130 (Aedes). PNG (PNG); Indonesia (Irian Jaya).
vanapa Huang, 1968b: 63 (Aedes). PNG (PNG).
variabilis Huang, 1968b: 65 (Aedes). PNG (PNG).

Genus CULEX Linnaeus

CULEX Linnaeus, 1758: 602. Type species: Culex pipiens Linnaeus, 1758, des. Latreille, 1810: 442.

Subgenus ACALLYNTRUM Stone & Penn

ACALLYNTRUM Stone & Penn, 1948: 109. Type species: Culex perkinsi Stone & Penn, 1948, orig. des.
axillicola Steffan, 1979: 133. PNG (PNG).
belkini Stone & Penn, 1948: 114. Solomon Is.
bicki Stone & Penn, 1947: 89. Indonesia (Irian Jaya).
binigrolineatus Knight & Rozeboom, 1945: 289. Indonesia (Irian Jaya).
bougainvillensis Steffan, 1979: 138. PNG (Bougainville I).
pallidiceps Theobald, 1905c: 101 (Melanoconion). PNG (PNG).
perkinsi Stone & Penn, 1948: 110. Solomon Is, PNG (Bougainville I).

Subgenus CULEX Linnaeus

CULEX Linnaeus, 1758: 602 (as gen.). Type species: Culex pipiens Linnaeus, 1758, des. Latreille, 1810: 442.
LEUCOMYIA Theobald, 1907: 372 (as gen.). Type species: Culex gelidus Theobald, 1901, orig. des. [Preocc. Brauer & Bergenstamm, 1891.]
THEOBALDIOMYIA Brunetti, 1912a: 462 (as gen.). Type species: Culex gelidus Theobald, 1901, orig. des.
albinervis Edwards, 1929g: 338. Fiji.
alis Theobald, 1903b: 167. Christmas I (Indian Ocean); Indonesia (Irian Jaya, Maluku); Indonesia (Kalimantan), Malaysia (Pen, Sabah), Philippines, Taiwan, Thailand, Vietnam.
annulirostris Skuse, 1889c: 1737. Australia (NSW); American Samoa,
Australia (NT, Qld), Belau, Cook Is, Fiji, French Polynesia (Austral Is, Society Is, Tuamotu Arch), Guam, Indonesia (Irian Jaya, Maluku), Kiribati (Gilbert Is), Marshall Is, Micronesia, ?Nauru, New Caledonia (Loyalty Is, New Caledonia), Northern Marianas, PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is, Tonga, Tuvalu, Vanuatu, Western Samoa; Indonesia (Nusa Tenggara, Sulawesi), Philippines.
     ssp. annulirostris s. str. Australia (NSW); American Samoa, Australia (NT, Qld), Belau, Cook Is, Fiji, French Polynesia (Austral Is, Society Is, Tuamotu Arch), Indonesia (Irian Jaya, Maluku), Kiribati (Gilbert Is), Micronesia, ?Nauru, New Caledonia (Loyalty Is, New Caledonia), PNG (Bougainville I, PNG), Solomon Is, Tonga, Tuvalu, Vanuatu, Western Samoa.
     bancroftii Theobald, 1901b: 367 (annulirostris ssp.). Australia (Qld).
     somerseti Taylor, 1912a: 28. Australia (Qld).
     palpalis Taylor, 1912b: 29. Australia (NT).
     jepsoni Bahr, 1912: 18. Fiji.
     consimilis Taylor, 1913: 55 (Culicelsa). Australia (Qld). [Preocc. Newstead, 1907.]
     simplex Taylor, 1914b: 698 (Culicelsa). Australia (Qld). [Preocc. Theobald, 1903.]
     ssp. marianae Bohart & Ingram, 1946: 42. Northern Marianas; Guam.
asteliae Belkin, 1968a: 66. NZ (NZ).
atriceps Edwards, 1926e: 105. French Polynesia (Society Is).
australicus Dobrotworsky & Drummond, 1953: 143 (pipiens ssp.). Australia (Vic); Australia (NSW), New Caledonia (New Caledonia), Vanuatu.
banksensis Maffi & Tenorio, 1977: 503. Vanuatu.
barraudi Edwards—not Australasian/Oceanian.
crinicaudus Edwards, 1921b: 77. Australia (NT).
     parvus Taylor, 1912b: 27. Australia (NT).
edwardsi Barraud, 1923b: 507. India; Australia (Qld), PNG (PNG); Philippines, Sri Lanka.
fuscocephalus Theobald, 1907: 420. Sri Lanka; Guam [immigrant]; widesp. Oriental Reg.
gelidus Theobald, 1901b: 20. Malaysia (Pen); Australia (Qld), Indonesia (Irian Jaya); Oriental Reg.
globocoxitus Dobrotworsky, 1953: 357. Australia (Vic).
incognitus Baisas, 1938: 203. Philippines, Indonesia (Maluku); Indonesia (Sulawesi).
iyengari Mattingly & Rageau, 1958: 244. New Caledonia (Loyalty Is); New Caledonia (New Caledonia).
kesseli Belkin, 1962: 200. French Polynesia (Society Is).
litoralis Bohart, 1946b: 43. Northern Marianas; Guam.
marquesensis Stone & Rosen, 1953: 354. French Polynesia (Marquesas).
mimulus Edwards, 1915a: 284. Malaysia (Sarawak); Australia (Qld), Indonesia (Irian Jaya), PNG (PNG); widesp. Oriental Reg.
     mossmani Taylor, 1915a: 181. Australia (Qld).
miraculosus Bonne-Wepster, 1937: 92. Indonesia (Irian Jaya).
molestus Forskål, 1775: 85. Egypt; widesp. in warmer Australasian/Oceanian Regs.; pantrop.
omani Belkin, 1962: 210. Solomon Is; Indonesia (Irian Jaya).
pacificus Edwards, 1916a: 360. Vanuatu.
pervigilans Bergroth, 1889: 295. NZ (NZ); NZ (Auckland Is, Chatham Is, Kermadec Is), ?Norfolk I.
pipiens Linnaeus—not Australasian/Oceanian.
pseudovishnui Colless, 1957a: 88. Singapore; Indonesia (Irian Jaya, Maluku); widesp. Oriental Reg.
quinquefasciatus Say, 1823: 10. USA; widesp. Australasian/Oceanian Regs.; pantrop.
     fatigans Wiedemann, 1828: 10. "Ostindien."
     acer Walker, 1848: 8. NZ (NZ).
     macleayi Skuse, 1889c: 1746. Australia (WA).
     skusii Giles, 1900: 292. "Australia."
     quasilinealis Theobald, 1907: 415. Australia (SA).
     fuscus Taylor, 1914b: 699 (Culicelsa). Australia (Qld). [Preocc. Theobald, 1905.]
     townsvillensis Taylor, 1919a: 836 (n. n. for fuscus Taylor).
     hutchinsoni, Basio, 1972, not Barraud, 1924, ?misid.
roseni Belkin, 1962: 203. French Polynesia (Society Is); ?Tonga.
rotoruae Belkin, 1968a: 64. NZ (NZ).
samoanensis Theobald, 1914: 36 (Pseudotaeniorhynchus). Western Samoa.
sitiens Wiedemann, 1828: 542. Indonesia (Sumatra); American Samoa, Fiji, Guam, Indonesia (Irian Jaya, Maluku), Micronesia, Nauru, New Caledonia (New Caledonia), Niue, PNG (Bougainville I), Solomon Is, Tonga, Tuvalu, Vanuatu, Wallis & Futuna, Western Samoa; widesp. Afrotrop., Oriental Regs.
solitarius Bonne-Wepster, 1938: 394. Indonesia (Irian Jaya); Indonesia (?Java).
squamosus Taylor, 1914b: 691. Australia (Qld); Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is.
     annulata Theobald, 1905c: 98 (Trichopronomyia). PNG (PNG). [Preocc. Schrank, 1776.]
     annulirostris Taylor, 1914b: 696 (Leucomyia). Australia (Qld). [Preocc. Skuse, 1889.]
     taylori Edwards, 1921b: 78 (n. n. for annulirostris Taylor).
starckeae Stone & Knight, 1958: 69 (n. n. for basicinctus Edwards). Australia (Qld); Australia (NSW), New Caledonia (New Caledonia), Vanuatu; Indonesia (Timor).
     basicinctus Edwards, 1922b: 96. Australia (Qld). [Preocc. Edwards, 1921.]
toviiensis Klein, Rivière & Séchan, 1984: 72. French Polynesia (Marquesas).
tritaeniorhynchus Giles, 1901: 606. India; Bonin Is, Guam, Micronesia, Northern Marianas; Afrotrop., S & E Oriental, Palearctic Regs.
     summorosus Dyar, 1920: 180. Philippines.
vicinus Taylor, 1916b: 569. Australia (NT); PNG (PNG).
     annulata Taylor, 1914b: 695 (Leucomyia). Australia (Qld). [Preocc. Schrank, 1776.]
     basicinctus Edwards, 1921b: 78 (n. n. for annulata Taylor).
vishnui Theobald, 1901b: 355. India; Indonesia (Irian Jaya, Maluku); Indonesia (Java, Kalimantan, Sulawesi, Sumatra, Timor), widesp. SE Oriental Reg.
whitmorei Giles, 1904a: 367 (Taeniorhynchus). Philippines; Australia (Qld), PNG (PNG); Oriental, E Palearctic Regs.
whittingtoni Belkin, 1962: 204. Solomon Is.

Subgenus CULICIOMYIA Theobald

TRICHORHYNCHUS Theobald, 1905b: 241 (as gen.). Type species: Trichorhynchus fuscus Theobald, 1905 [= Culex fragilis Ludlow 1903], mon. [Preocc. Balbiani, 1887.]
CULICIOMYIA Theobald, 1907: 227 (as gen.). Type species: Culiciomyia inornata Theobald, 1907 [= Culex fragilis Ludlow, 1903], des. Edwards, 1912a: 33.
PECTINOPALPUS Theobald, 1909: 11 (as gen.). Type species: Trichorhynchus fuscus Theobald, 1905 [= Culex fragilis Ludlow, 1903], mon.
TRICHORHYNCHOMYIA Brunetti, 1912a: 477 (as gen.; n. n. for Trichorhynchus).
bailyi Barraud, 1934: 382. India; Indonesia (Irian Jaya); Sri Lanka, Thailand.
ceramensis Sirivanakarn & Kurihara, 1973: 220. Indonesia (Maluku).
fragilis Ludlow, 1903: 142. Philippines; Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; India, Malaysia (Pen, Sabah), Sri Lanka, Thailand.
     inornata Theobald, 1907: 227 (Culiciomyia). Malaysia (Sarawak). [Preocc. Theobald, 1905.]
fuscicinctus King & Hoogstraal, 1946e: 145. Indonesia (Irian Jaya).
maplei Knight & Hurlburt, 1949: 31 (pullus ssp.). Micronesia.
nailoni King & Hoogstraal, 1946e: 143. Indonesia (Irian Jaya).
nigropunctatus Edwards, 1926e: 121 (n. n. for annulata Theobald). Malaysia (Sarawak); Belau, Micronesia; Ryukyu Is, Oriental Reg.
     annulata Theobald, 1907: 230 (Culiciomyia). Malaysia (Sarawak). [Preocc. Schrank, 1776.]
pallidothorax Theobald, 1905d: 32. India; Indonesia (Irian Jaya, Maluku); China, Japan, Ryukyu Is, widesp. Oriental Reg.
papuensis Taylor, 1914c: 201 (Melanoconion). PNG (PNG); PNG (Bismarck Arch), Solomon Is; Malaysia (Pen), Philippines, Thailand.
pullus Theobald, 1905c: 87. PNG (PNG); Australia (Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, Bougainville I), Solomon Is; Indonesia (Timor).
     muticus Edwards, 1923c: 6. PNG (Bismarck Arch).
ruthae Peters, 1958: 91 (as ruthii). PNG (PNG).
spathifurcus Edwards, 1915a: 284 (Culiciomyia). Malaysia (Sarawak); Indonesia (Irian Jaya, Maluku); SE Oriental Reg.

Subgenus EUMELANOMYIA Theobald

EUMELANOMYIA Theobald, 1909: 10 (as gen.). Type species: Culex inconspicuosa Theobald, 1909, mon.
MOCHTHOGENES Edwards, 1930c: 305. Type species: Aedes malayi Leicester, 1908, orig. des.
brevipalpis Giles, 1902: 384 (Stegomyia). India; Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG); widesp. Oriental Reg.
cataractarum Edwards, 1923c: 7. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG), Philippines.
femineus Edwards, 1926e: 107. Vanuatu.
foliatus Brug, 1932: 82. Indonesia (Java); ?Indonesia (Maluku); widesp. Oriental Reg.
malayi Leicester, 1908: 184 (Aedes). Malaysia (Pen); Indonesia (Irian Jaya); widesp. Oriental Reg.
manusensis Sirivanakarn, 1975: 394. PNG (Bismarck Arch).

Subgenus LOPHOCERAOMYIA Theobald

LOPHOCERAOMYIA Theobald, 1905b: 245 (as gen.). Type species: Lophoceraomyia uniformis Theobald, 1905, mon.
LOPHOCERATOMYIA Theobald, 1905c: 93 (as gen.) Type species: Lophoceratomyia fraudatrix Theobald, 1905, des. Brunetti, 1914: 64.
CYATHOMYIA de Meijere, 1910b: 921 (as gen.). Type species: Cyathomyia jenseni de Meijere, 1910, mon.
Colless, 1960: 385. Solomon Is; PNG (Bismarck Arch, Bougainville I).
     franclemonti Belkin, 1962: 256. Solomon Is.
Brug, 1939: 112 (minor var.). Indonesia (Java); Indonesia (Maluku).
becki Belkin, 1962: 265. Solomon Is.
bergi Belkin, 1962: 258. Solomon Is.
bolii Sirivanakarn, 1968: 100. PNG (PNG).
buxtoni Edwards, 1926e: 106 (hilli var.). Vanuatu; Indonesia (Maluku).
carolinensis Bohart & Ingram, 1946: 30. Micronesia.
castaneus Sirivanakarn, 1973: 212. PNG (PNG).
christiani Colless, 1960: 389. PNG (PNG); Indonesia (Irian Jaya).
collessi Sirivanakarn, 1968: 170. PNG (Bismarck Arch); PNG (PNG).
cottlei Sirivanakarn, 1968: 180. PNG (Bismarck Arch).
crowei Sirivanakarn, 1968: 115. PNG (PNG).
cubitatus Colless, 1965a: 273. Singapore; Indonesia (Irian Jaya, Maluku), PNG (PNG); Malaysia (Pen, Sabah), Philippines.
cylindricus Theobald, 1903b: 202. Australia (Qld); PNG (Bismarck Arch).
digoelensis Brug, 1932: 81. Indonesia (Irian Jaya); PNG (PNG).
     caeruleus King & Hoogstraal, 1947b: 67. PNG (PNG).
durhami Sirivanakarn, 1968: 177. PNG (PNG).
fraudatrix Theobald, 1905c: 94 (Lophoceratomyia). PNG (PNG); Australia (NT, Qld), Indonesia (Irian Jaya); Indonesia (Sumatra).
     molestus Wiedemann, 1828: 542. Indonesia (Sumatra). [Preocc. Forskål, 1775.]
     annulata Taylor 1916b: 571 (Lophoceratomyia). Australia (NT). [Preocc. Schrank, 1776.]
     cairnsensis Taylor, 1919a: 837 (Lophoceratomyia). Australia (Qld). [Preocc. Taylor, 1919.]
gagnei Evenhuis in Evenhuis & Gon, 1989: 209 (n.n. for ornatus Theobald). PNG (PNG); Indonesia (Irian Jaya), PNG (Bismarck Arch).
     ornatus Theobald, 1905c: 100 (Melanoconion). PNG (PNG). [Preocc. Meigen, 1818.]
gossi Bohart, 1957: 68. Micronesia.
gressitti Sirivanakarn, 1968: 153. PNG (PNG).
hilli Edwards, 1922b: 95. Australia (NT).
     australis Taylor, 1915a: 178 (Neomacleaya). Australia (Qld). [Preocc. Erichson, 1842.]
hurlbuti Belkin, 1962: 266. Solomon Is; PNG (Bismarck Arch, Bougainville I).
insularis Sirivanakarn, 1968: 166. PNG (Bismarck Arch).
kaviengensis Sirivanakarn, 1968: 167. PNG (Bismarck Arch).
kowiroensis Sirivanakarn, 1968: 133. PNG (PNG).
kuhnsi King & Hoogstraal, 1955: 1. Indonesia (Irian Jaya); Indonesia (Maluku), PNG (PNG); Malaysia (Sabah), Philippines.
     mercedesae Baisas, 1974: 121 (uniformis ssp.). Philippines.
kusaiensis Bohart, 1957: 69. Micronesia.
laffooni Belkin, 1962: 261. Solomon Is.
lairdi Belkin, 1962: 257. Solomon Is; PNG (Bougainville I).
lakei Sirivanakarn, 1968: 120. PNG (PNG).
leei King & Hoogstraal, 1955: 4. Indonesia (Irian Jaya); PNG (PNG).
marksae King & Hoogstraal, 1955: 6. Indonesia (Irian Jaya).
minjensis Sirivanakarn, 1968: 151. PNG (PNG).
minor Leicester, 1908: 126 (Lophoceratomyia). Malaysia (Pen); Indonesia (Irian Jaya); Indonesia (Java, Kalimantan), Philippines, SE Oriental Reg.
muruae Sirivanakarn, 1968: 130. PNG (PNG).
orbostiensis Dobrotworsky, 1958a: 317. Australia (Vic).
oweni Belkin, 1962: 259. Solomon Is.
perryi Belkin, 1962: 268. Solomon Is.
petersi Colless, 1960: 388. PNG (PNG).
pseudornatus Colless, 1960: 386. PNG (PNG).
pseudorubithoracis Sirivanakarn, 1968: 178. PNG (PNG).
rajaneeae Sirivanakarn, 1968: 174. PNG (Bismarck Arch).
schilfgaardei Sirivanakarn, 1968: 163. PNG (PNG).
sedlacekae Sirivanakarn, 1968: 180. PNG (PNG).
shanahani Sirivanakarn, 1968: 117. PNG (PNG); Indonesia (Irian Jaya).
singuawaensis Sirivanakarn, 1969: 119 (n. n. for confusus Sirivanakarn). PNG (PNG).
     confusus Sirivanakarn, 1968: 105. PNG (PNG). [Preocc. Baisas, 1938.]
solomonis Edwards, 1929f: 316 (fraudatrix var.). Solomon Is; Indonesia (Irian Jaya), PNG (Bougainville I, PNG).
steffani Sirivanakarn, 1968: 112. PNG (PNG).
submarginalis Sirivanakarn, 1973: 212. PNG (PNG).
versabilis Sirivanakarn, 1968: 126. PNG (PNG).
walukasi Belkin, 1962: 264. Solomon Is; PNG (Bougainville I).
wamanguae Sirivanakarn, 1968: 135. PNG (PNG).
winkleri Belkin, 1962: 260. Solomon Is.

Subgenus LUTZIA Theobald

LUTZIA Theobald, 1903b: 155 (as gen.). Type species: Culex bigoti Bellardi, 1862, mon.
JAMESIA Christophers, 1906: 12 (as gen.). Type species: Culex concolor Robineau-Desvoidy, 1827 [= Culex fuscanus Wiedemann, 1820], des. Edwards, 1932d: 190. [Preocc. Jeckel, 1861.]
fuscanus Wiedemann, 1820b: 9 [1821a: 9; 1821b: 9]]. India; ?"Australia," Belau, Guam, Micronesia, Northern Marianas; Indonesia (Java, Sumatra, Timor), widesp. Oriental Reg.
     concolor Robineau-Desvoidy, 1827: 405. Not given [unknown].
     setulosus Doleschall, 1857: 384. Indonesia (Java).
     luridus Doleschall, 1857: 384. Indonesia (Java).
halifaxii Theobald, 1903b: 231. Malaysia (Pen); Australia (NSW, NT, Qld), Bonin Is, Indonesia (Irian Jaya), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is, Volcano Is; Japan, Korea, Ryukyu Is, Oriental Reg.
     vorax Edwards, 1921d: 327 (Lutzia). Japan.
     tigripes, Mochizuki, 1913, not de Grandpré & de Charmoy, 1901, misid.
     halifaxi, error for halifaxii.
shinonagai Tanaka, Misusawa & Saugstad, 1979: 245. Bonin Is.

Subgenus NEOCULEX Dyar

NEOCULEX Dyar, 1905: 48 (as gen.). Type species: Culex territans Walker, 1856, orig. des.
chaetoventralis Theobald, 1910b: 461 (Neomelanoconion). Australia (Qld).
cheesmanae Mattingly & Marks, 1955: 166. New Caledonia (New Caledonia); New Caledonia (Loyalty Is).
crassistylus Brug, 1934: 516. PNG (PNG).
douglasi Dobrotworsky, 1956: 110. Australia (Vic).
dumbletoni Belkin, 1962: 244. New Caledonia (New Caledonia).
fergusoni Taylor, 1914d: 459 (Culicada). Australia (NSW).
gaufini Belkin, 1962: 243. New Caledonia (New Caledonia).
latus Dobrotworsky, 1956: 108. Australia (WA).
leonardi Belkin, 1962: 245. Solomon Is; PNG (Bougainville I).
millironi Belkin, 1962: 242. New Caledonia (New Caledonia).
pedicellus King & Hoogstraal, 1947b: 65. Indonesia (Irian Jaya).
postspiraculosus Lee, 1944b: 221. Australia (NSW).
pseudomelanoconius Theobald, 1907: 416. Australia (Qld).

Subgenus OCULEOMYIA Theobald

OCULEOMYIA Theobald, 1907: 515 (as gen.). Type species: Oculeomyia sarawakae Thoelbald, 1907, orig. des.
bitaeniorhynchus Giles, 1901: 607. India; Australia (NT, Qld), Belau, Indonesia (Irian Jaya), New Caledonia (New Caledonia), PNG (PNG); widesp. Afrotrop., Oriental, S Palearctic Regs.
     abdominalis Taylor, 1913: 53 (Culicelsa). Australia (Qld).


SIRAVANAKARNIUS Tanaka, 2004: 224. Type species: Culex boninensis Bohart, 1957, orig. des.
boninensis Bohart, 1957: 75. Bonin Is.

Subgenerically unplaced species of Culex Linnaeus

cairnsensis Taylor, 1919a: 829. Australia (Qld).

Unplaced species of Culicini

novaeguineae Evenhuis in Evenhuis & Gon, 1989: 210 (Culex; n. n. for ventralis Walker). Indonesia (Irian Jaya).
     ventralis Walker, 1865a: 103 (Culex). "New Guinea" [= Indonesia (Irian Jaya)]. [Preocc. Walker, 1860.]


THEOBALDIA Neveu-Lemaire, 1902b: 1331. Type species: Culex annulatus Schrank, 1776, orig. des. [Preocc. Fischer, 1885.]
CULISETA Felt, 1904: 391c. Type species: Culex absobrinus Felt, 1904 [= Culex impatiens Walker, 1848], orig. des.

Subgenus AUSTROTHEOBALDIA Dobrotworsky

AUSTROTHEOBALDIA Dobrotworsky, 1954: 68 (Theobaldia subg.). Type species: Chrysoconops littleri Taylor, 1914, mon.
littleri Taylor, 1914b: 702 (Chrysoconops). Australia (Tas).

Subgenus CLIMACURA Howard, Dyar & Knab

CLIMACURA Howard, Dyar & Knab, 1915: 195, 452 (Culex subg.). Type species: Culex melanurus Coquillett, 1902, orig. des.
antipodea Dobrotworsky, 1963: 291. Australia (Vic).
novaezealandiae Pillai, 1966: 127. NZ (NZ).
tonnoiri Edwards, 1925a: 258 (Theobaldia). NZ (NZ).

Subgenus CULICELLA Felt

CULICELLA Felt, 1904: 391c (as gen.). Type species: Culex dyari Coquillett, 1902, orig. des.
atra Lee, 1944b: 220 (Theobaldia). Australia (WA).
drummondi Dobrotworsky, 1960b: 241 (Theobaldia). Australia (Vic).
inconspicua Lee, 1937: 294 (Theobaldia). Australia (NSW).
otwayensis Dobrotworsky, 1960b: 246 (Theobaldia). Australia (Vic).
sylvanensis Dobrotworsky, 1960b: 245 (Theobaldia). Australia (Vic).
victoriensis Dobrotworsky, 1954: 73 (Theobaldia). Australia (Vic).
weindorferi Edwards, 1926e: 111 (Theobaldia). Australia (Tas).

Subgenus NEOTHEOBALDIA Dobrotworsky

NEOTHEOBALDIA Dobrotworsky, 1954: 68 (Theobaldia subg.). Unavailable name; genus-group name proposed after 1930 without type-species designation.
NEOTHEOBALDIA Dobrotworsky, 1958b: 186 (Theobaldia subg.). Type species: Theobaldia hilli Edwards, 1926, orig. des.
frenchii Theobald, 1901b: 66 (Culex). Australia (Vic).
     ssp. atritarsalis Dobrotworsky, 1954: 73 (Theobaldia). Australia (Vic).
     ssp. frenchii s. str. Australia (Vic).
hilli Edwards, 1926e: 111 (Theobaldia). Australia (Vic).

Unplaced species of Culiseta Felt

arenivaga Marks, 1968: 51. Australia (Qld).

Genus FICALBIA Theobald

FICALBIA Theobald, 1903b: 296. Type species: Uranotaenia minima Theobald, 1901, des. Blanchard, 1905: 418.
minima Theobald, 1901c: 262 (Uranotaenia). India; Indonesia (Irian Jaya), PNG (PNG); China, Indonesia (Java), widesp. Oriental Reg.

Genus MIMOMYIA Theobald

MIMOMYIA Theobald, 1903b: 304. Type species: Mimomyia splendens Theobald, 1903, des. Blanchard, 1905: 419.

Subgenus ETORLEPTIOMYIA Theobald

ETORLEPTIOMYIA Theobald, 1904f: 71 (as gen.). Type species: Etorleptiomyia mediolineata Theobald, 1904, mon.
OREILLIA Ludlow, 1905a: 101 (as gen.; as O’Reillia). Type species: Oreillia luzonensis Ludlow, 1905, mon.
DIXOMYIA Taylor, 1914b: 702 (as gen.). Type species: Dixomyia elegans Taylor, 1914, mon.
ETIORLEPTIOMYIA, error for Etorleptiomyia.
bougainvillensis Belkin, 1962: 294 (Ficalbia). PNG (Bougainville I).
elegans Taylor, 1914b: 703 (Dixomyia). Australia (Qld); Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG), Solomon Is; Indonesia (Java), Philippines, Ryukyu Is, Thailand.
solomonis Belkin, 1962: 292 (Ficalbia). Solomon Is; PNG (Bougainville I).

Subgenus MIMOMYIA Theobald

MIMOMYIA Theobald, 1903b: 304 (as gen.). Type species: Mimomyia splendens Theobald, 1903, des. Blanchard, 1905: 419.
LUDLOWIA Theobald, 1907: 193 (as gen.). Type species: Mimiomyia chamberlaini Ludlow, 1904, des. Sharp, 1909: 360.
CONOPOMYIA Leicester, 1908: 113 (as gen.). Type species: Conopomyia metallica Leicester, 1908, des. Brunetti, 1914: 58.
chamberlaini Ludlow, 1904c: 297. Philippines; Australia (Qld), PNG (PNG); widesp. Oriental Reg.
     ssp. chamberlaini s. str. Philippines; Australia (Qld), PNG (PNG); widesp. Oriental Reg.
     ssp. metallica Leicester, 1908: 113 (Conopomyia; as sp.). Malaysia (Pen); Australia (Qld), Indonesia (Irian Jaya), PNG (PNG); Oriental Reg.
flavens King & Hoogstraal, 1946d: 188 (Ficalbia). Indonesia (Irian Jaya).
gurneyiBelkin, 1962: 296 (Ficalbia). PNG (Bougainville I).
hybrida Leicester, 1908: 115 (Conopomyia). Malaysia (Pen); Indonesia (Irian Jaya); SE Oriental Reg.
modesta King & Hoogstraal, 1946d: 186 (Ficalbia). Indonesia (Irian Jaya).

Genus HODGESIA Theobald

HODGESIA Theobald, 1904a: 17. Type species: Hodgesia sanguinae Theobald, 1904, mon.
cairnsensis Taylor, 1919b: 842. Australia (Qld); PNG (Bismarck Arch, PNG), Solomon Is.
malayi Leicester, 1908: 231. Malaysia (Pen); Indonesia (Maluku); SE Oriental Reg.
quasisanguinae Leicester, 1908: 230. Malaysia (Pen); Australia (Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch); Indonesia (Sulawesi), Philippines.
     triangulatus Taylor, 1914c: 204. PNG (PNG).
solomonis Belkin, 1962: 280. Solomon Is; PNG (Bougainville I).
spoliata Edwards, 1923c: 8. PNG (PNG); Indonesia (Irian Jaya, Maluku); Indonesia (Sulawesi).


COQUILLETTIDIA Dyar, 1905: 47. Type species: Culex perturbans Walker, 1856, orig. des.


AUSTROMANSONIA Belkin, 1968: 76. Type species: Taeniorhynchus tenuipalpis Edwards, 1924, orig. des.
tenuipalpis Edwards, 1924d: 366 (Taeniorhynchus). NZ (NZ).


COQUILLETTIDIA Dyar, 1905: 47 (as gen.). Type species: Culex perturbans Walker, 1856, orig. des.
aurata Dobrotworsky, 1963: 295 (Mansonia). Australia (Vic).
crassipes Wulp, 1881: 9 (Culex). Indonesia (Sumatra); Indonesia (Irian Jaya, Maluku); Ryukyu Is, widesp. Oriental Reg.
     brevicellulus Theobald, 1901b: 212 (Taeniorhynchus). Burma & Malaysia (Pen).
     fuscopteron Theobald, 1911: 239 (Chrysoconops). Indonesia (Irian Jaya).
fijiensis Belkin, 1962: 308 (Mansonia). Fiji; ?Western Samoa.
giblini Taylor, 1914c: 198 (Pseudotaeniorhynchus conopas var.). PNG (PNG); Indonesia (Irian Jaya), PNG (Bismarck Arch).
iracunda Walker, 1848: 6 (Culex). NZ (NZ).
linealis Skuse, 1889c: 1747 (Culex). Australia (NSW).
lutea Belkin, 1962: 309 (Mansonia). Solomon Is; PNG (Bougainville I).
memorans Bonne-Wepster, 1930a: 621 [1930b: 393; 1931: 268] (Taeniorhynchus). Indonesia (Irian Jaya)
nigrochracea Bonne-Wepster, 1930a: 620 [1930b: 392; 1931: 266] (Taeniorhynchus). Indonesia (Irian Jaya); Indonesia (Sumatra).
ochracea Theobald, 1903b: 263 (Taeniorhynchus). Malaysia (Pen); Indonesia (Irian Jaya); Japan, Korea, Ryukyu Is, SE Oriental Reg.
samoaensis Stone, 1966: 331. American Samoa; Western Samoa.
variegata Dobrotworsky, 1963: 293 (Mansonia). Australia (Vic).
xanthogaster Edwards, 1924d: 366 (Taeniorhynchus). Australia (Qld); Australia (NT), New Caledonia (Loyalty Is, New Caledonia), Vanuatu.

Genus MANSONIA Blanchard

PANOPLITES Theobald, 1900: 5. Suppressed by I.C.Z.N., 1959c: 187.
MANSONIA Blanchard, 1901: 1046 (n. n. for Panoplites). Type species: Culex titillans Walker, 1848, des. Neveu-Lemaire, 1902a: 214. [Ratified by I.C.Z.N., 1959c: 187.]
TAENIORHYNCHUS, authors, not Lynch Arribálzaga, 1891, misid.

Subgenus MANSONIOIDES Theobald

MANSONIOIDES Theobald, 1907: 498 (as gen.). Type species: Mansonioides septemguttata Theobald, 1907 [= Panoplites annulifera Theobald, 1901], mon.
MAUSONIOIDES, error for Mansonioides.
annulifera Theobald, 1901b: 183 (Panoplites). India; Indonesia (Irian Jaya); widesp. Oriental Reg.
     septemguttata Theobald, 1907: 499 (Mansonoides). Malaysia (Sarawak).
bonnewepsterae Assem, 1958b: 205. Indonesia (Irian Jaya).
dives Schiner, 1868: 31 (Culex; n. n. for annulipes Walker). Singapore; Australia (Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG); Oriental Reg.
     annulipes Walker, 1856a: 6 (Culex). Singapore. [Preocc. Meigen, 1830.]
indiana Edwards, 1930d: 541. Indonesia (Java); Indonesia (Irian Jaya), PNG (PNG); Oriental Reg.
melanesiensis Belkin, 1962: 314. Solomon Is; PNG (Bougainville I).
papuensis Taylor, 1914c: 200 (Taeniorhynchus). PNG (PNG); Indonesia (Irian Jaya).
septempunctata Theobald, 1905c: 107. PNG (PNG); Australia (Qld), PNG (Bismarck Arch).
uniformis Theobald, 1901b: 180 (Panoplites). India; Australia (NSW, NT, Qld, Vic), Guam, Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG), Solomon Is; Japan, Korea, Ryukyu Is, Afrotrop., Oriental Regs.
     australiensis Giles, 1902: 355 (Panoplites). Australia (Qld).


ORTHOPODOMYIA Theobald, 1904e: 236. Type species: Orthopodomyia albipes Leicester, 1904, mon.
papuensis Zavortink, 1968: 139. PNG (PNG); ?Australia (Qld), Indonesia (Irian Jaya, Maluku).

Genus MALAYA Leicester

MALAYA Leicester, 1908: 258. Type species: Malaya genurostris Leicester, 1908, mon.
HARPAGOMYIA de Meijere, 1909b: 165. Type species: Harpagomyia splendens de Meijere, 1909, mon.
genurostris Leicester, 1908: 258. Malaysia (Pen); Australia (NT, Qld), Indonesia (Irian Jaya); Oriental Reg.
leei Wharton, 1947: 60. PNG (PNG); Indonesia (Irian Jaya); Philippines.
solomonis Wharton, 1947: 63 (Harpagomyia). PNG (Bougainville I).
splendens de Meijere, 1909b: 167 (Harpagomyia). Indonesia (Java); ?Indonesia (Irian Jaya); Indonesia (Kalimantan).


MAORIGOELDIA Edwards, 1930c: 302 (Rachionotomyia subg.). Type species: Culex arygropus Walker, 1848, mon.
argyropa Walker, 1848: 2 (Culex). NZ (NZ).

Genus TOPOMYIA Leicester

TOPOMYIA Leicester, 1908: 238. Type species: Topomyia minor Leicester, 1908, des. Brunetti, 1914: 72.

Subgenus SUAYMYIA Thurman

SUAYMYIA Thurman, 1959: 44 (as gen.). Type species: Suaymyia cristata Thurman, 1959, orig. des.
papuensis Marks, 1960: 91. PNG (PNG).


TRIPTEROIDES Giles, 1904a: 369. Type species: Runchomyia philippinensis Giles, 1904 [= Uranotaenia nitidoventer Giles, 1904], orig. des.

Subgenus POLYLEPIDOMYIA Theobald

POLYLEPIDOMYIA Theobald, 1905c: 118 (as gen.). Type species: Polylepidomyia argenteiventris Theobald, 1905, mon.
altivallis Bonne-Wepster, 1948: 311. Indonesia (Irian Jaya).
argenteiventris Theobald, 1905c: 118 (Polylepidomyia). PNG (PNG); Indonesia (Irian Jaya), PNG (Bismarck Arch).
     atra Taylor, 1914c: 190 (Stegomyia). PNG (PNG).
     brugi Edwards, 1927d: 356 (Rachionotomyia). Indonesia (Irian Jaya).
caledonicus Edwards, 1922b: 100 (Rachionotomyia). New Caledonia (New Caledonia).
coheni Belkin, 1950: 250. Solomon Is; PNG (Bougainville I).
collessi Lee, 1946: 271. Australia (Qld).
digoelensis Brug, 1934: 506. Indonesia (Irian Jaya).
floridensis Belkin, 1950: 269. Solomon Is.
folicola Belkin, 1955: 230. Vanuatu.
marksae Dobrotworsky, 1965: 58. Australia (Vic).
melanesiensis Belkin, 1955: 233. Vanuatu; New Caledonia (Loyalty Is, New Caledonia), Solomon Is.
microlepis Edwards, 1927d: 353 (Rachionotomyia). Indonesia (Irian Jaya).
perplexus Peters, 1963c: 91. PNG (PNG).
punctolateralis Theobald, 1903a: 156 (Stegomyia). Australia (Qld); Indonesia (Irian Jaya); Indonesia (Timor).
     occidentalis Brug, 1934: 506 (atripes var.). Indonesia (Timor).
rotumanus Edwards, 1929g: 337 (Rachionotomyia). Fiji.
solomonis Edwards, 1924d: 363 (Rachionotomyia). Solomon Is; ?New Caledonia (New Caledonia), PNG (Bougainville I).
standfasti Peters, 1959: 148. PNG (PNG).
tasmaniensis Strickland, 1911: 249 (Stegomyia). Australia (Tas).
     cephasi Edwards, 1923c: 8 (Rachionotomyia). Australia (Tas).

Subgenus RACHISOURA Theobald

RACHISOURA Theobald, 1910b: 207 (as gen.). Type species: Rachisoura sylvestris Theobald, 1910, mon.
adentatus Assem, 1959: 48. PNG (PNG); Indonesia (Irian Jaya).
bisquamatus Lee, 1946: 252. Indonesia (Irian Jaya).
brevirhynchus Brug, 1934: 503. Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).
confusus Lee, 1946: 251. PNG (PNG); Indonesia (Irian Jaya).
cuttsi Assem, 1959: 46. PNG (PNG); Indonesia (Irian Jaya).
exnebulis Bonne-Wepster, 1948: 308. PNG (PNG); Indonesia (Irian Jaya).
felicitatis Bonne-Wepster, 1948: 307. PNG (PNG); Indonesia (Irian Jaya).
filipes Walker, 1861b: 229 (Culex). Indonesia (Irian Jaya); Australia (NT, Qld), Indonesia (Maluku), PNG (Bismarck Arch, PNG).
     hilli Taylor, 1914d: 456 (Stegomyia). Australia (NT).
flabelliger Bonne-Wepster, 1948: 308. PNG (PNG); Indonesia (Irian Jaya).
fuliginosus Lee, 1946: 248. Indonesia (Irian Jaya).
fuscipleura Lee, 1946: 246. Indonesia (Irian Jaya).
kingi Lee, 1946: 256. Indonesia (Irian Jaya).
latisquamus Edwards, 1927d: 354 (Rachionotomyia). Indonesia (Irian Jaya).
leei Peters, 1959: 141. PNG (PNG); Indonesia (Irian Jaya).
longipalpatus Lee, 1946: 249. Indonesia (Irian Jaya).
mathesoni Belkin, 1950: 219. Solomon Is; Indonesia (Irian (Jaya); PNG (Bougainville I).
obscurus Brug, 1934: 504. Indonesia (Irian Jaya).
pallidus Lee, 1946: 255. Indonesia (Irian Jaya).
papua Brug, 1934: 505. Indonesia (Irian Jaya).
pilosus Lee, 1946: 255. Indonesia (Irian Jaya).
plumiger Bonne-Wepster, 1948: 310. Indonesia (Irian Jaya).
simplex Brug, 1934: 507. Indonesia (Irian Jaya).
stonei Belkin, 1950: 288. Solomon Is.
subobscurus Lee, 1946: 273. Australia (Qld).
sylvestris Theobald, 1910b: 208 (Rachisoura). Australia (Qld).
tityae Sloof, 1961: 22. Indonesia (Irian Jaya).
torokinae Belkin, 1950: 236. PNG (Bougainville I).
vanleeuweni Edwards, 1927d: 355 (Rachionotomyia). Indonesia (Irian Jaya).


TRIPTEROIDES Giles, 1904a: 369 (as gen.). Type species: Runchomyia philippinensis Giles, 1904 [= Uranotaenia nitidoventer Giles, 1904], orig. des.
MIMETEOMYIA Theobald, 1910b: 210 (as gen.). Type species: Mimeteomyia apicotriangulata Theobald, 1910 [= Culex atripes Skuse, 1889], mon.
alboscutellatus Lee, 1946: 235. Indonesia (Irian Jaya).
atripes Skuse, 1889c: 1750 (Culex). Australia (NSW); Australia (Qld).
     apicotriangulata Theobald, 1910b: 211 (Mimeteomyia). Australia (Qld).
bimaculipes Theobald, 1905c: 114 (Phoniomyia). PNG (PNG); Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch).
     ornata Taylor, 1914c: 189 (Stegomyia). PNG (PNG).
binotatus Belkin, 1950: 264. PNG (Bougainville I).
bonneti Belkin, 1962: 501. Solomon Is; Vanuatu.
brevipalpis Brug, 1934: 508. Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).
distigma Edwards, 1925a: 257 (Rachionotomyia). Solomon Is.
elegans Brug, 1934: 507. Indonesia (Irian Jaya).
lipovskyi Belkin, 1950: 256. Solomon Is.
littlechildi Edwards, 1930d: 544 (Rachionotomyia). PNG (PNG); Indonesia (Irian Jaya).
lorengaui Peters, 1963b: 67. PNG (Bismarck Arch).
magnesianus Edwards, 1924d: 361 (Rachionotomyia). Australia (Qld).
nissanensis Lee, 1946: 239. PNG (Bismarck Arch).
novohanoverae Peters, 1963b: 63. PNG (Bismarck Arch).
purpuratus Edwards, 1921b: 79 (Rachionotomyia). Fiji; ?American Samoa.
quasiornatus Taylor, 1915a: 177 (Stegomyia). Australia (Qld); ?PNG (Bismarck Arch, PNG).
splendens Lee, 1946: 233. PNG (PNG).

Unplaced species of Tripteroides Giles

concinnus Lee, 1946: 260. Indonesia (Irian Jaya).
subnudipennis Edwards, 1927d: 355 (Rachionotomyia). Indonesia (Irian Jaya).

Genus WYEOMYIA Theobald

WYEOMYIA Theobald, 1901b: 98 [1901c: 267]. Type species: Wyeomyia grayii Theobald, 1901, des. Neveu-Lemaire, 1902a: 223.
mitchellii Theobald, 1905e: 37 (Dendromyia). Jamaica; Hawaiian Is [immigrant]; SE USA, widesp. C. Am., West Indies.
     jamaicensis Theobald, 1905e: 11 (Dendromyia). Jamaica. [Alternate orig. name for mitchellii Theobald.]

Genus URANOTAENIA Lynch Arribálzaga

URANOTAENIA Lynch Arribálzaga, 1891a: 375 [1891b: 163]. Type species: Uranotaenia pulcherrima Lynch Arribálzaga, 1891, des. Neveu-Lemaire, 1902a: 227.

Subgenus PSEUDOFICALBIA Theobald

PSEUDOFICALBIA Theobald, 1911b: 272 [1912: 89] (as gen.). Type species: Ficalbia inornata Theobald, 1908 [= Uranotaenia fusca Theobald, 1907], mon.
atra Theobald, 1905c: 114. PNG (PNG); Australia (Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch).
     nigerrima Taylor, 1914c: 203. PNG (PNG).
     quadrimaculata, authors, not Edwards, 1929, misid.
colocasiae Edwards, 1928a: 281. Fiji.
diagonalis Brug, 1934: 510. Indonesia (Irian Jaya); Australia (Qld), PNG (Bismarck Arch, PNG).
hirsutifemora Peters, 1964: 21. PNG (PNG); Australia (Qld), Indonesia (Irian Jaya); Oriental Reg.
obscura Edwards, 1915a: 285. Malaysia (Sarawak); Indonesia (Irian Jaya, Maluku), PNG (PNG); Oriental Reg.
     papua Brug, 1924: 441. Indonesia (Irian Jaya).
painei Edwards, 1935a: 127. Fiji.
quadrimaculata Edwards, 1929f: 313. Solomon Is; PNG (Bougainville I).

Subgenus URANOTAENIA Lynch Arribálzaga

URANOTAENIA Lynch Arribálzaga, 1891a: 375 [1891b: 163] (as gen.). Type species: Uranotaenia pulcherrima Lynch Arribálzaga, 1891, des. Neveu-Lemaire, 1902a: 227.
ANISOCHELEOMYIA Theobald, 1905a: 52 (as gen.). Type species: Anisocheleomyia nivipes Theobald, 1905, des. Brunetti, 1914: 55.
albescens Taylor, 1914b: 705. Australia (Qld); Australia (NT), Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).
albosternopleura Peters, 1963c: 203. PNG (PNG); Australia (Qld).
amiensis Peters, 1963e: 207. PNG (PNG); Australia (Qld).
antennalis Taylor, 1919a: 840. Australia (Qld); Indonesia (Irian Jaya).
argyrotarsis Leicester, 1908: 214. Malaysia (Pen); Australia (NT, Qld), Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG); Philippines.
     argyritarsis, error for argyrotarsis.
barnesi Belkin, 1953: 332. Solomon Is; PNG (Bougainville I), Vanuatu.
civinskii Belkin, 1953: 341. Solomon Is.
fimbriata King & Hoogstraal, 1947a: 592. Indonesia (Irian Jaya).
gerdae Sloof, 1963: 84. Indonesia (Irian Jaya).
heiseri Baisas, 1935c: 72. Philippines; ?Indonesia (Irian Jaya).
lateralis Ludlow, 1905c: 385 (caeruleocephala var.). Philippines; Australia (NT, Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; widesp. Oriental Reg.
     propria Taylor, 1914b: 704. Australia (Qld).
     cairnsensis Taylor, 1919a: 839. Australia (Qld).
moresbyensis Peters, 1963e: 204. PNG (PNG); Australia (Qld).
King & Hoogstraal, 1947a: 590. Indonesia (Irian Jaya).
nivipes Theobald, 1905a: 53 (Anisocheleomyia). Australia (Qld); Australia (NT).
     albofasciata Taylor, 1920: 164. Australia (NT).
novaguinensis Peters, 1963d: 141. PNG (PNG); Australia (Qld).
     spp. alticola Peters, 1963d: 142. PNG (PNG).
     ssp. novaguinensis s. str. PNG (PNG); Australia (Qld).
Peters, 1964: 19. PNG (PNG); Australia (Qld).
paranovaguinensis Peters, 1963d: 135. PNG (PNG).
pygmaea Theobald, 1901b: 254. Australia (Qld); Australia (NSW).
setosa King & Hoogstraal, 1947a: 585. Indonesia (Irian Jaya).
sexaurei Belkin, 1953: 359. Solomon Is; PNG (Bougainville I).
solomonis Belkin, 1953: 350. Solomon Is; ?Indonesia (Irian (Jaya), PNG (Bougainville I).
subtibioclada King & Hoogstraal, 1947a: 595. Indonesia (Irian Jaya).
tibialis Taylor, 1919a: 839. Australia (Qld); Vanuatu.
tibioclada King & Hoogstraal, 1947a: 593. PNG (PNG).
wysockii Belkin, 1953: 366. Solomon Is; PNG (Bougainville I).


TOXORHYNCHITES Theobald, 1901a: 234 [1901b: 244]. Type species: Toxorhynchites brevipalpis Theobald, 1901, subs. mon. Theobald, 1901b: 245. [Validated by I.C.Z.N., 1959b: 167. Type species conserved by I.C.Z.N., 1982: 122.]

Subgenus LYNCHIELLA Lahille

MEGARHINUS Robineau-Desvoidy, 1827: 412. Type species: Culex haemorrhoidalis Fabricius, 1787, mon. [Preocc. Rafinesque, 1820.]
Culex haemorrhoidalisFabricius, 1787, mon.
MEGARHINA, MEGARRHINA, errors for Megarhinus.
theobaldi Dyar & Knab, 1906: 246. Colombia; Hawaiian Is [laboratory culture; not released]; Neotropical Reg.
     hypoptes Knab, 1907: 50. Nicaragua.

Subgenus TOXORHYNCHITES Theobald

TOXORHYNCHITES Theobald, 1901a: 234 [1901b: 244]. Type species: Toxorhynchites brevipalpis Theobald, 1901, subs. mon. Theobald, 1901b: 245. [Validated by I.C.Z.N., 1959b: 167. Type species conserved by I.C.Z.N., 1982: 122.]
amboinensis Doleschall, 1857: 381 (Culex). Indonesia (Maluku); American Samoa [introduced], Guam [introduced; not established], Hawaiian Is [introduced], Indonesia (Irian Jaya), PNG (PNG), Western Samoa [introduced]; ?Malaysia (Pen), Philippines.
     subulifer Doleschall, 1857: 382 (Culex). Indonesia (Maluku).
     splendens, authors, not Wiedemann, 1819, misid.
brevipalpis Theobald, 1901b: 245. South Africa; American Samoa & Guam [introduced; not established], Hawaiian Is & French Polynesia (Marquesas, Society Is) [introduced]; S Afrotrop. Reg.
inornatus Walker, 1865a: 102 (Megarhina). "New Guinea" [= Indonesia (Irian Jaya)]; Australia (Qld), Fiji [introduced], Hawaiian Is [introduced], ?Indonesia (Maluku), PNG (Bismarck Arch).
nepenthicola Steffan & Evenhuis, 1982: 2. PNG (PNG).
speciosus Skuse, 1889c: 1722 (Megarrhina). Australia (Qld); Australia (NSW, NT), PNG (PNG).
splendens Wiedemann, 1819a: 2 (Culex). Indonesia (Java); Fiji [introduced], Indonesia (Irian Jaya), PNG (PNG); widesp. Oriental Reg.

Last revised 16 February 2011