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94. Family HELEOMYZIDAE

[Originally] by Brian R. Pitkin

There are over 500 described species of Heleomyzidae in about 65 genera and 22 tribes distributed throughout the world; the greatest number occur in the Holarctic Region (McAlpine 1985b). All 76 species in 22 genera and 10 tribes listed here are from Australia, New Zealand, and Papua New Guinea.

Little is known of the biology of the Australasian species. Adults of Diplogeomyza and related genera (McAlpine 1967) are frequently encountered in wet forest country, some being most prevalent in mountainous areas. Adults of Borboroides and Heleomicra are attracted to carcasses and feces. Austroleria extensa has been reared from the fungus Boletus granulatus (Polyporaceae). The larvae of the Holarctic Suillinae occur principally in fungi, as do those of Tapeigaster, whereas larvae of Cairnsimyia live in borer tunnels in trees.

The Borboropsidae, Chiropteromyzidae, Cnemospathidae, Heteromyzidae, Notomyzidae, Rhinotoridae, and Trixoscelididae have been recognized by some recent authors as distinct from the Heleomyzidae. McAlpine (1985b), however, included all of them as tribes within the Heleomyzidae [Heteromyzidae (= Heleomyzini)].
Refs.: McAlpine (1967, keys Diplogeomyza, Austroleria spp.; 1968, key Cairnsimyia spp., larval descr.; 1984, key Pseudoleria spp.; 1985b, key Australasian gen.), McAlpine & Kent (1982, key Tapeigaster spp., larval descr.).

Tribe ALLOPHYLOPSINI

Genus ALLOPHYLINA Tonnoir & Malloch

ALLOPHYLINA Tonnoir & Malloch, 1927: 87. Type species: Allophylina albitarsis Tonnoir & Malloch, 1927, orig. des.
albitarsis Tonnoir & Malloch, 1927: 88. NZ (NZ).

Genus ALLOPHYLOPSIS Lamb

ALLOPHYLOPSIS Lamb, 1909: 142. Type species: Allophylopsis punctata Lamb, 1909 [= Heteromyza laquei Hutton, 1902], mon.

Subgenus ALLOPHYLELLA Tonnoir & Malloch

ALLOPHYLELLA Tonnoir & Malloch, 1927: 90. Type species: Allophylopsis philpotti Tonnoir & Malloch, 1927, mon.
philpotti Tonnoir & Malloch, 1927: 91. NZ (NZ).

Subgenus ALLOPHYLLATA Harrison

ALLOPHYLLATA Harrison, 1959: 138. Type species: Allophylopsis inconspicua Tonnoir & Malloch, 1927, orig. des.
inconspicua Tonnoir & Malloch, 1927: 95. NZ (NZ).

Subgenus ALLOPHYLOPSIS Lamb

ALLOPHYLOPSIS Lamb, 1909: 142 (as gen.). Type species: Allophylopsis punctata Lamb, 1909 [= Heteromyza laquei Hutton, 1902], mon.
chathamensis Tonnoir & Malloch, 1927: 94. NZ (Chatham Is).
distincta Tonnoir & Malloch, 1927: 98. NZ (NZ).
fulva Hutton, 1901: 82 (Leria). NZ (NZ).
fuscipennis Tonnoir & Malloch, 1927: 97. NZ (NZ).
hudsoni Hutton, 1901: 8 (Helomyza). NZ (NZ).
      proxima Tonnoir & Malloch, 1927: 96. NZ (NZ).
laquei Hutton, 1902a: 173 (Heteromyza). NZ (NZ).
      punctata Lamb, 1909: 143. NZ (NZ).
minuta Tonnoir & Malloch, 1927: 95. NZ (NZ).
obscura Tonnoir & Malloch, 1927: 99. NZ (NZ).

Subgenus HUTTONOMYIA Malloch

HUTTONOMYIA Malloch, 1922c: 227 (as gen.). Type species: Helomyza scutellata Hutton, 1901, orig. des.
PARAPHYLOPSIS Tonnoir & Malloch, 1927: 93. Type species: Helomyza scutellata Hutton, 1901, des. Harrison, 1959: 132.
bivittata Harrison, 1959: 135. NZ (NZ).
scutellata Hutton, 1901: 81 (Helomyza). NZ (NZ).
subscutellata Tonnoir & Malloch, 1927: 93. NZ (NZ).

Subgenus PHILPOTTOMYIA Tonnoir & Malloch

PHILPOTTOMYIA Tonnoir & Malloch, 1927: 91. Type species: Allophylopsis lineata Tonnoir & Malloch, 1927, orig. des.
lineata Tonnoir & Malloch, 1927: 92. NZ (NZ).
rufithorax Tonnoir & Malloch, 1927: 92. NZ (NZ).

Genus AUSTROLERIA McAlpine

AUSTROLERIA McAlpine, 1967: 80. Type species: Austroleria extensa McAlpine, 1967, orig. des.
extensa McAlpine, 1967: 80. Australia (NSW); Australia (ACT, SA, Tas, Vic).
truncata McAlpine, 1967: 81. Australia (NSW); Australia (Tas, Vic).

Genus DIPLOGEOMYZA Hendel

DIPLOGEOMYZA Hendel, 1917: 38. Type species: Diplogeomyza diaphora Hendel, 1917, orig. des.
annularis McAlpine, 1967: 96. Australia (NSW); Australia (Vic).
conformis McAlpine, 1967: 88. Australia (ACT); Australia (NSW, Tas, Vic).
diaphora Hendel, 1917: 38. Australia (Vic); Australia (NSW, Qld, SA, Tas).
flavipalpis McAlpine, 1967: 93. Australia (NSW).
hardyi McAlpine, 1967: 90. Australia (NSW); Australia (ACT, SA, Vic).
      ssp. hardyi s. str. Australia (NSW); Australia (ACT, SA, Vic).
      ssp. tasmanica McAlpine, 1967: 92. Australia (Tas).
immaculata McAlpine, 1967: 89. Australia (Qld).
incisa McAlpine, 1967: 85. Australia (Vic).
maculipennis Malloch, 1926d: 552 (Huttonomyia). Australia (NSW); widesp. Australia (excl. NT).
media McAlpine, 1967: 100. Australia (NSW); Australia (Qld, Vic).
pectinervis McAlpine, 1967: 104. Australia (NSW); Australia (Qld).
signata McAlpine, 1967: 99. Australia (NSW); Australia (Qld).
spinosa McAlpine, 1967: 98. Australia (NSW); Australia (Qld, Vic).
tridens McAlpine, 1967: 94. Australia (Qld); Australia (NSW).
victoriae McAlpine, 1967: 96. Australia (Vic).
wirthi McAlpine, 1967: 86. Australia (NSW); Australia (Tas, Vic).

Genus LERIOPSIS McAlpine

LERIOPSIS McAlpine, 1967: 76. Type species: Leriopsis montana McAlpine, 1967, orig. des.
montana McAlpine, 1967: 76. Australia (Tas); Australia (ACT, NSW, Vic).

Genus TRIXOLERIA McAlpine

TRIXOLERIA McAlpine, 1967: 77. Type species: Trixoleria maculata McAlpine, 1967, orig. des.
maculata McAlpine, 1967: 78. Australia (NSW); Australia (ACT, Tas, Vic, WA).

Tribe BORBOROIDINI

Genus BORBOROIDES Malloch

BORBOROIDES Malloch, 1925f: 85. Type species: Borboroides atra Malloch, 1925, orig. des.
atra Malloch, 1925f: 85. Australia (NSW); Australia (Qld, Tas, Vic,
WA).

Genus HELEOMICRA McAlpine

HELEOMICRA McAlpine, 1985b: 210. Type species: Heleomicra collessi McAlpine, 1985, orig. des.
collessi McAlpine, 1985b: 210. Australia (NSW).

Tribe CNEMOSPATHIDINI

Genus PROSOPANTRUM Enderlein

PROSOPANTRUM Enderlein, 1912e: 135. Type species: Prosopantrum austrinum Enderlein, 1912 [= Agromyza diadematum Bigot, 1888], orig. des.
ACROSTICHALIA Tonnoir & Malloch, 1927: 86. Type species: Acrostichalia flavifrons Tonnoir & Malloch, 1927, orig. des.
CNEMOSPATHIS Enderlein, 1938b: 655. Type species: Cnemospathis baeckstroemi Enderlein, 1938, orig. des.
flavifrons Tonnoir & Malloch, 1927: 87 (Acrostichalia). NZ (NZ); Australia (Tas); S. Am, S Africa [introduced].
      baeckstroemi Enderlein, 1938b: 656 (Cnemospathis). Chile.
      schoenemanni Enderlein, 1938b: 657 (Cnemospathis). Chile.

Tribe DIACIINI

Genus AMPHIDYSIS McAlpine

AMPHIDYSIS McAlpine, 1985b: 208. Type species: Amphidysis hesperia McAlpine, 1985, orig. des.
hesperia McAlpine, 1985b: 208. Australia (WA).

Tribe FENWICKIINI

Genus ANEURIA Malloch

ANEURIA Malloch, 1930m: 340. Type species: Aneuria bipunctata Malloch, 1930, des. Harrison, 1959: 162. [Aneuria elegans Malloch, 1930, nomen nudum, was originally designated as type species.]
angusta Harrison, 1959: 163. NZ (NZ).
bipunctata Malloch, 1930m: 341. NZ (NZ).
imitatrix Malloch, 1930m: 340. NZ (NZ).
sexpunctata Malloch, 1930m: 342. NZ (NZ).
tripunctata Malloch, 1930m: 342. NZ (NZ).

Nomen nudum in Aneuria Malloch

elegans Malloch, 1930m: 340.

Genus FENWICKIA Malloch

FENWICKIA Malloch, 1930m: 336. Type species: Fenwickia hirsuta Malloch, 1930, orig. des.
affinis Harrison, 1959: 160. NZ (NZ).
caudata Harrison, 1959: 155. NZ (NZ).
claripennis Malloch, 1930m: 337. NZ (NZ).
hirsuta Malloch, 1930m: 338. NZ (NZ).
nuda Malloch, 1930m: 338. NZ (NZ).
similis Malloch, 1930m: 338. NZ (NZ).

Genus XENEURA Malloch

XENEURA Malloch, 1930m: 339. Type species: Milichia picata Hutton, 1902, orig. des.
picata Hutton, 1902b: 195 (Milichia). NZ (NZ).

Tribe HELEOMYZINI

Genus OECOTHEA Haliday

OECOTHEA Haliday in Curtis, 1837: 280. Type species: Heleomyza fenestralis Fallén, 1820, des. Gorodkov, 1984: 18. [Conserved by I.C.Z.N. 1995: 104.]
NEOECOTHEA Peterson & Gill, 1982: 219. Type species: Helomyza fenestralis Fallén, 1820, orig. des.
fenestralis Fallén, 1820a: 5 (Helomyza). Sweden; Australia (NSW, Tas) [introduced], NZ (NZ) [introduced]; Europe, Asia, N. & S. Am.

Genus PSEUDOLERIA Garrett

PSEUDOLERIA Garrett, 1921: 128. Type species: Blepharoptera pectinata Loew, 1872, orig. des.
pectinata Loew, 1872a: 99 [1872b: 275] (Blepharoptera). USA; Australia (ACT, NSW, Vic) [introduced], ?NZ (NZ) [introduced]; ?Chile, South Africa, N. Am.
      pectinerata Garrett, 1921: 128. Canada.
placata Hutton, 1901: 82 (Leria). NZ (NZ); Australia (NSW, Vic) [introduced], NZ (Chatham Is); Canada, USA.
      crassata Garrett, 1925: 3. Canada & USA.


Genus SPILOCHROA Williston

SPILOCHROA Williston, 1907: 2. Type species: Heterochroa ornata Johnson, 1895, mon.
ornata Johnson, 1895: 306 (Heterochroa). USA; Hawaiian Is [immigrant]; s. USA, West Indies.

Genus TEPHROCHLAMYS Loew

TEPHROCHLAMYS Loew, 1862e: 72. Type species: Helomyza rufiventris Meigen, 1830, des. Coquillett, 1910: 613.
rufiventris Meigen, 1830: 58 (Helomyza). Not given [Central Europe]; NZ (NZ) [introduced]; Holarctic Reg.

Tribe PENTACHAETINI

Genus PENTACHAETA McAlpine

PENTACHAETA McAlpine, 1985b: 212. Type species: Pentachaeta physopus McAlpine, 1985, orig. des.
physopus McAlpine, 1985b: 213. Australia (NSW).

Tribe RHINOTORINI

Genus CAIRNSIMYIA Malloch

CAIRNSIMYIA Malloch, 1931e: 294. Type species: Cairnsimyia cavifrons Malloch, 1931, orig. des.
aroana McAlpine, 1968: 275. PNG (PNG); Australia (Qld).
cavifrons Malloch, 1931e: 294. Australia (Qld); Australia (NSW).
crosskeyi McAlpine, 1968: 274. PNG (PNG).
excavata McAlpine, 1968: 271. Australia (Qld); Australia (NSW).
inscripta Walker, 1865b: 120 (Platystoma). "New Guinea" [= Indonesia (Irian Jaya)]; PNG (PNG).
robusta Walker, 1853: 403 (Heteromyza). Australia (Tas); Australia (NSW, Qld, SA, Vic).
sydneiensis McAlpine, 1968: 270. Australia (NSW).
uniseta McAlpine, 1968: 272. Australia (NSW); Australia (Qld, Vic).
verticalis McAlpine, 1968: 272. Australia (Qld); Australia (NSW).

Genus ZENTULA McAlpine

ZENTULA McAlpine, 1985b: 212. Type species: Zentula vittata McAlpine, 1985, orig. des.
vittata McAlpine, 1985b: 212. Australia (ACT); Australia (NSW).

Genus ZINZA Sinclair & McAlpine

ZINZA Sinclair & McAlpine, 1995: 225. Type species: Zinza grandis Sinclair & McAlpine, 1995, orig. des.
grandis Sinclair & McAlpine, 1995: 227. Australia (Qld).

Tribe TAPEIGASTRINI

Genus OLLIX McAlpine

OLLIX McAlpine, 1985b: 206. Type species: Ollix cogani McAlpine, 1985, orig. des.
cogani McAlpine, 1985b: 207. Australia (WA).

Genus TAPEIGASTER Macquart

TAPEIGASTER Macquart, 1847a: 86 [1847b: 102]. Type species: Tapeigaster annulipes Macquart, 1847, orig. des.
SCIOMYZOPTERA Hendel, 1917: 46. Type species: Sciomyzoptera annulata Hendel, 1917, orig. des.
annulata Hendel, 1917: 47 (Sciomyzoptera). "Australia"; Australia (NSW, Qld).
      fulva Malloch, 1926d: 553, Australia (NSW).
annulipes Macquart, 1847a: 87 [1847b: 103]. "Nouvelle-Hollande"; Australia (NSW, Qld, SA, Tas, Vic).
      cingulipes Walker, 1858a: 220 (Dryomyza). Australia (NSW).
argyrospila Bezzi, 1923a: 77. Australia (NSW); Australia (Qld, SA, Vic).
brunneifrons Malloch, 1927b: 16. Australia (NSW); Australia (ACT, Tas, Vic).
      vernalis Paramonov, 1955f: 457. Australia (NSW).
cinctipes Walker, 1853: 404 (Heteromyza). Australia (Tas); Australia (ACT, NSW, Vic, WA).
      bella Paramonov, 1955f: 454. Australia (ACT & Tas & Vic & WA).
digitata McAlpine & Kent, 1982: 45. Australia (NSW); Australia (ACT, SA, Tas, Vic).
luteipennis Bezzi, 1923a: 76. Australia (NSW); Australia (Qld).
      taylori Malloch, 1935d: 94. Australia (NSW).
nigricornis Macquart, 1851: 277(304) (Sciomyza). Australia (Tas); Australia (NSW, Qld, SA, Vic, WA).
      marginifrons Bezzi, 1923a: 74. Australia (NSW).
paramonovi McAlpine & Kent, 1982: 44. Australia (NSW); Australia (ACT, SA, Tas, Vic).
pulverea McAlpine & Kent, 1982: 50. Australia (NSW); Australia (Qld, Vic).
subglabra McAlpine & Kent, 1982: 52. Australia (NSW); Australia (ACT, Vic).

Tribe WATERHOUSEIINI

Genus WATERHOUSEIA Malloch

WATERHOUSEIA Malloch, 1936b: 260. Type species: Waterhouseia cyclops Malloch, 1936, orig. des.
cyclops Malloch, 1936b: 261. Australia (NSW).


This page last revised 3 June 2007