Australasian/Oceanian Diptera Catalog -- Web Version

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Among the world mycetophiloids, the family Mycetophilidae is the most diversified, with about 3,000 species in 150 genera and at least twice this number of species awaiting description. The family is cosmopolitan, being well represented in forested areas worldwide. A number of the extant genera have been preserved in Baltic amber (late Eocene to lower Oligocene), and the available paleontological and biogeographical information implies that the family was already well diversified by at least the Middle Cretaceous.

Like the Keroplatidae, the mycetophilid fauna of these regions contains an old Transantarctic element plus pantropical and Indomalayan elements. As far as can be ascertained without phylogenetic and biogeographic studies on Oriental and Australasian mycetophilids (yet to be undertaken), the greatest part of the Australasian fauna seems predominantly of Oriental origin. Morganiella, Austrosciophila, Paramorganiella, Tasmanina, Taxicnemis, and Trizygia (Sciophilinae); Anomalomyia, Ateleia, Caethronia, Cycloneura, Paracycloneura, Paradoxa, Sigmoleia, and Trichoterga (Leiinae); and Pseudalyiisina and Zygophronia (Mycetophilinae) are endemic Australasian genera, many with close relationships to Neotropical genera with a Chilean center of endemism.

Most mycetophilids inhabit wet forests but are quite common in swamps and are present, although in limited numbers of species, in the moister parts of heath and open grassland. Larvae of mycetophilids are mainly bound to the sporophores of fleshy Basidiomycetes (esp. Agaricaeceae, Amanitaceae, Boletaceae, and Russulaceae) inside which they feed on spores and hyphae (or perhaps on spores only). Some species can be bred from Polyporaceae (bracket fungi) and rarely from Ascomycetes and Myxomycetes. A few species are monophagous or polyphagous, most of the other species are restricted to particular genera or families of fungi.

Other larvae (esp. Mycomyinae, Sciophilinae, and Leiinae) spin glutinous webs under sporophores or under bark of dead trunks and branches. Some of these larvae are predaceous and others mycophagous. A few Afrotropical and Palearctic species are reported to live on mosses and liverworts.

Nothing seems to be known of the larval biology of the Australasian mycetophilids. Regarding genera known from these regions and bred elsewhere, larvae of Mycomya spin webs under bark, in cavities of rotten wood, and more rarely under fungi. There is definite evidence of their mycophagy, but some species may be predaceous (Väisänen 1984). It has been demonstrated that the larvae of a Palearctic species of Neoempheria are predaceous. Phthinia and Tetragoneura have been reported from rotting wood associated with fungi, and Manota and Platurocypta from Myxomycetes. All the Australasian genera of Exechiini and Mycetophila have been bred elsewhere on fleshy Basidiomycetes. Other members of the Mycetophilini are surface feeders under encrusting fungi (Trichonta, Phronia) or mycelium on wood (Epicypta) (Stubbs & Chandler 1978, Hackman & Meinander 1979).

Imagos of mycetophilids are found mostly in damp and shady places in forests or along stream banks in open country. They also occur in nests of small mammals and in caves, sometimes in great numbers (Antriadophila petulans). They can be collected by sweeping, occasionally at light traps or windows, and very good results are obtained with yellow-tray and Malaise traps. The orange and black Pseudalysiina mimicans from Tasmania mimics a braconid wasp of the tribe Alysiini (Tonnoir 1929b).

Some of the present families of Sciaroidea have long been treated as mere subfamilies of Mycetophilidae, and are still considered as such by many taxonomists. The main papers on the systematics of Australasian mycetophilids are mostly the same as for keroplatids: Skuse (1888c, 1890d) and Tonnoir (1929b) for Australia, Tonnoir & Edwards (1927) for New Zealand, and Colless (1966) for Micronesia.

The phylogeny of the subfamilies of Mycetophilidae is far from being clarified, and several subfamilies are probably polyphyletic. Therefore, the classification adopted here is rather conventional, following Edwards (1925b) for the main divisions of the family but raising most of his tribes to subfamily rank as proposed by Väisänen (1984).

MYCOMYA Rondani, 1856: 194. Type species: Sciophila marginata Meigen, 1818, orig. des.
MYCOMYIA, error for Mycomya.

CYMOMYA Väisänen, 1984: 271. Type species: Sciophila circumdata Staeger, 1840, orig. des.
rufonigra Matile, 1991: 236. New Caledonia (New Caledonia).

MYCOMYA Rondani, 1856: 194 (as gen.). Type species: Sciophila marginata Meigen, 1818, orig. des.
MYCOMYIA, error for Mycomya.
flavilatera Tonnoir in Tonnoir & Edwards, 1927: 802. NZ (NZ).
furcata Edwards in Tonnoir & Edwards, 1927: 802. NZ (NZ).
humeralis Skuse, 1890d: 609 (Sciophila). Australia (NSW).
maccoyi Skuse, 1890d: 606 (Sciophila). Australia (Vic).
par Walker, 1856d: 417 (Sciophila). Australia (Tas).
pictithorax Skuse, 1890d: 607 (Sciophila). Australia (NSW).
plagiata Tonnoir in Tonnoir & Edwards, 1927: 802. NZ (NZ).
richmondensis Skuse, 1890d: 608 (Sciophila). Australia (NSW).
sylvicola Skuse, 1890d: 610 (Sciophila). Australia (NSW).

MYCOMYOPSIS Väisänen, 1984: 295. Type species: Sciophila trilineata Zetterstedt, 18??, orig. des.
minuscula Matile, 1991: 238. New Caledonia (New Caledonia).
perparva Matile, 1991: 240. New Caledonia (New Caledonia).
yoshimotoi Matile, 1991: 241. New Caledonia (New Caledonia).

EMPHERIA Winnertz, 1863: 738. Type species: Sciophila striata Meigen, 1818, des. Coquillett, 1910: 537. [Preocc. Hagen, 1854.]
NEOEMPHERIA Osten Sacken, 1878: 9 (n. n. for Empheria).
signifera Skuse, 1890d: 611. Australia (NSW).
vicina Colless, 1966: 652. Bonin Is.

ALLOCOTOCERA Mik, 1886a: 102 (n. n. for Eurycera).
EURYCERA Dziedzicki, 1885: 166. Type species: Eurycera flava Dziedzicki, 1885 [= Leia pulchella Curtis, 1837], mon. [Preocc. de Laporte, 1833.]
EURYCERAS Marshall, 1896b: 291. Type species: Euryceras anaclinoides Marshall, 1896, mon.
anaclinoides Marshall, 1896b: 292 (Euryceras). NZ (NZ).
cephasi Edwards in Tonnoir & Edwards, 1927: 803. NZ (NZ).
crassipalpis Tonnoir in Tonnoir & Edwards, 1927: 804. NZ (NZ).
dilatata Tonnoir in Tonnoir & Edwards, 1927: 804. NZ (NZ).

ANEURA Marshall, 1896b: 287. Type species: Aneura boletinoides, Marshall, 1896, mon.
appendiculata Tonnoir in Tonnoir & Edwards, 1927: 808. NZ (NZ).
bispinosa Edwards in Tonnoir & Edwards, 1927: 808. NZ (NZ).
boletinoides Marshall, 1896b: 288. NZ (NZ); NZ (Auckland Is).
defecta Edwards in Tonnoir & Edwards, 1927: 809. NZ (NZ).
fagi Marshall, 1896b: 282 (Sciophila). NZ (NZ).
filiformis Tonnoir in Tonnoir & Edwards, 1927: 807. NZ (NZ).
fusca Tonnoir in Tonnoir & Edwards, 1927: 807. NZ (NZ).
longicauda Tonnoir in Tonnoir & Edwards, 1927: 809. NZ (NZ).
longipalpis Tonnoir in Tonnoir & Edwards, 1927: 809. NZ (NZ).
nitida Tonnoir in Tonnoir & Edwards, 1927: 807. NZ (NZ).
pallida Edwards in Tonnoir & Edwards, 1927: 810. NZ (NZ).

EUDICRANA Loew, 1870: 142 [1872b: 180]. Type species: Eudicrana obumbrata Loew, 1870, mon.
PAREUDICRANA Tonnoir, 1929b: 600. Type species: Pareudicrana monticola Tonnoir, 1929, orig. des.
araucariae Matile, 1991: 242. New Caledonia (New Caledonia).
monticola Tonnoir, 1929b: 600 (Pareudicrana). Australia (Tas).
nicholsoni Tonnoir, 1929b: 600 (Pareudicrana). Australia (NSW).

AUSTROSCIOPHILA Tonnoir, 1929b: 604 (Sciophila subg.). Type species: Sciophila solitaria Tonnoir, 1929, orig. des.
solitaria Tonnoir, 1929b: 604 (Sciophila). Australia (Tas).

MORGANIELLA Tonnoir & Edwards, 1927: 817. Type species: Morganiella fusca Tonnoir in Tonnoir & Edwards, 1927, orig. des.
fusca Tonnoir in Tonnoir & Edwards, 1927: 817. NZ (NZ).

NEOALLOCOTOCERA Tonnoir, 1929b: 601. Type species: Neoallocotocera fusca Tonnoir, 1929, orig. des.
fusca Tonnoir, 1929b: 601. Australia (Tas).

APHELOMERA Skuse, 1888c: 1206. Type species: Aphelomera sydneyensis Skuse, 1888, mon. [Preocc. Stephens, 1833.]
NEOAPHELOMERA Miller, 1945: 72 (n. n. for Aphelomera).
elongata Tonnoir in Tonnoir & Edwards, 1927: 815 (Aphelomera). NZ (NZ).
forcipata Edwards in Tonnoir & Edwards, 1927: 815 (Aphelomera). NZ (NZ).
longicauda Edwards in Tonnoir & Edwards, 1927: 815 (Aphelomera). NZ (NZ).
majuscula Edwards in Tonnoir & Edwards, 1927: 814 (Aphelomera). NZ (NZ).
marshalli Edwards in Tonnoir & Edwards, 1927: 816 (Aphelomera). NZ (NZ).
opaca Tonnoir in Tonnoir & Edwards, 1927: 814 (Aphelomera). NZ (NZ).
seclusa Matile, 1991: 244. New Caledonia (New Caledonia).
skusei Marshall, 1896b: 296 (Aphelomera). NZ (NZ); NZ (Auckland Is).
sydneyensis Skuse, 1888c: 1207 (Aphelomera). Australia (NSW).

NEOTRIZYGIA Tonnoir & Edwards, 1927: 816. Type species: Neotrizygia obscura Tonnoir in Tonnoir & Edwards, 1927, orig. des.
obscura Tonnoir in Tonnoir & Edwards, 1927: 816. NZ (NZ).

PARAMORGANIELLA Tonnoir, 1929b: 606. Type species: Paramorganiella adventurosa Tonnoir, 1929, orig. des.
adventurosa Tonnoir, 1929b: 606. Australia (Tas).

PARATRIZYGIA Tonnoir, 1929b: 605. Type species: Paratrizygia conformis Tonnoir, 1929, orig. des.
conformis Tonnoir, 1929b: 605. Australia (Tas).

PARVICELLULA Marshall, 1896b: 284. Type species: Parvicellula triangula Marshall, 1896, mon.
apicalis Tonnoir in Tonnoir & Edwards, 1927: 811. NZ (NZ).
fascipennis Edwards in Tonnoir & Edwards, 1927: 811. NZ (NZ).
gracilis Tonnoir in Tonnoir & Edwards, 1927: 811. NZ (NZ).
hamata Edwards in Tonnoir & Edwards, 1927: 812. NZ (NZ).
nigricoxa Tonnoir in Tonnoir & Edwards, 1927: 812. NZ (NZ).
obscura Tonnoir in Tonnoir & Edwards, 1927: 811. NZ (NZ).
ruficoxa Tonnoir in Tonnoir & Edwards, 1927: 812. NZ (NZ).
subhamata Tonnoir in Tonnoir & Edwards, 1927: 812. NZ (NZ).
triangula Marshall, 1896b: 284. NZ (NZ).

PHTHINIA Winnertz, 1863: 779. Type species: Phthinia humilis Winnertz, 1863, des. Johannsen, 1909: 83.
longiventris Tonnoir in Tonnoir & Edwards, 1927: 813. NZ (NZ).

Genus SCIOPHILA Meigen

SCIOPHILA Meigen, 1818: 245. Type species: Sciophila hirta Meigen, 1818, des. Curtis, 1837: 641.
ocreata Philippi, 1865: 625. Chile; NZ (NZ) [introduced].
parviareolata Santos Abreu, 1920: 28. Canary Is; NZ (NZ) [introduced].

HOMASPIS Skuse, 1888c: 1191. Type species: Homaspis meridiana Skuse, 1888, mon. [Preocc. Förster, 1868.]
STENOPHRAGMA Skuse, 1890d: 612 (n. n. for Homaspis).
bickeli Oliveira & Amorim, 2012: 150. Australia (WA).
collessi Oliveira & Amorim, 2012: 154. Australia (WA).
hirtipenne Skuse, 1890d: 614. Australia (NSW).
meridianum Skuse, 1888c: 1192 (Homapsis). Australia (SA).
paponorum Matile, 1991: 245. New Caledonia (New Caledonia).
picticorne Skuse, 1890d: 613. Australia (SA).

TASMANINA Tonnoir, 1929b: 602. Type species: Tasmanina gracilis Tonnoir, 1929, orig. des.
gracilis Tonnoir, 1929b: 603. Australia (Tas).

TAXICNEMIS Tonnoir & Edwards, 1927: 805. Type species: Sciophila hirta Marshall, 1896 [preocc., = Taxicnemis marshalli Matile,1989], orig. des.
flava Edwards in Tonnoir & Edwards, 1927: 805. NZ (NZ).
marshalli Matile, 1989 (n. n. for hirta Marshall). NZ (NZ).
     hirta Marshall, 1896b: 283 (Sciophila). NZ (NZ). [Preocc. Meigen, 1818.]
     ssp. bivittata Edwards in Tonnoir & Edwards, 1927: 805 (hirta var.). NZ (NZ).
     ssp. marshalli s.str. NZ (NZ).

TRIZYGIA Skuse, 1888c: 1204. Type species: Trizygia flavipes Skuse, 1888, mon.
flavipes Skuse, 1888c: 1205. Australia (NSW).

Subfamily METANEPSINAE

Genus CHALSTONEPSIA Søli

CHALSTONEPSIA Søli, 1996: 79. Type species: Chalastonepsia orientalis Søli, 1996, orig. des.
hokkaidensis Kallweit, 1998: 341. Japan; PNG (PNG); Brunei, Indonesia, Malaysia, Taiwan, Thailand.
orientalis Søli, 1996: 81. Malaysia; PNG (PNG); Brunei, Indonesia, Thailand.

AUSTROSYNAPHA Tonnoir, 1929b: 607. Type species: Austrosynapha hirta Tonnoir, 1929, orig. des.
SYNAPHA, Tonnoir & Edwards, 1927, not Meigen, 1818, misid.
apicalis Tonnoir in Tonnoir & Edwards, 1927: 818 (Synapha). NZ (NZ).
cawthroni Tonnoir in Tonnoir & Edwards, 1927: 819 (Synapha). NZ (NZ).
claripennis Tonnoir in Tonnoir & Edwards, 1927: 818 (Synapha). NZ (NZ).
gracilis Tonnoir in Tonnoir & Edwards, 1927: 818 (Synapha). NZ (NZ).
hirta Tonnoir, 1929b: 607. Australia (Tas).
parva Tonnoir in Tonnoir & Edwards, 1927: 819 (Synapha). NZ (NZ).
pulchella Tonnoir in Tonnoir & Edwards, 1927: 819 (Synapha). NZ (NZ).
similis Tonnoir in Tonnoir & Edwards, 1927: 818 (Synapha). NZ (NZ).

SYNAPHA Meigen, 1818: 227. Type species: Synapha fasciata Meigen, 1818, mon.
consueta Matile, 1991: 234. New Caledonia (New Caledonia).

ACRODICRANIA Skuse, 1888c: 1194. Type species: Acrodicrania atricauda Skuse, 1888, des. Johannsen, 1909: 82.
angustifurca Skuse, 1890d: 616. Australia (NSW).
atricauda Skuse, 1888c: 1195. Australia (NSW); Australia (Qld).
fasciata Skuse, 1888c: 1198. Australia (NSW); Australia (Qld).
setosicauda Skuse, 1888c: 1196. Australia (NSW).

ALLACTONEURA de Maijere, 1907: 201. Type species: Allactoneura cincta de Meijere, 1907, mon.
neocaledonica Matile, 1993: 205. New Caledonia (New Caledonia).
obscurata Walker, 1865a: 130 (Mycetophila). Indonesia (Irian Jaya); "Borneo," Indonesia (Sula Arch, Sulawesi).
papuensis Bechev, 1995: 81. Indonesia (Irian Jaya).

ANOMALA Marshall, 1896b: 293. Type species: Mycetophila guttata Hutton, 1881, mon. [Preocc. Block, 1799.]
ANOMALOMYIA Hutton, 1904a: 134 (n. n. for Anomala).
affinis Tonnoir in Tonnoir & Edwards, 1927: 822. NZ (NZ).
basalis Tonnoir in Tonnoir & Edwards, 1927: 821. NZ (NZ).
flavicauda Edwards in Tonnoir & Edwards, 1927: 822. NZ (NZ).
guttata Hutton, 1881: 11 (Mycetophila). NZ (Auckland Is).
immaculata Edwards in Tonnoir & Edwards, 1927: 820. NZ (NZ).
intermedia Matile, 1993: 170. New Caledonia (New Caledonia).
minor Marshall, 1896b: 295 (Anomala). NZ (NZ).
nasuta Matile, 1993: 169. New Caledonia (New Caledonia).
obscura Tonnoir in Tonnoir & Edwards, 1927: 821. NZ (NZ).
picta Matile, 1993: 171. New Caledonia (New Caledonia).
subobscura Tonnoir in Tonnoir & Edwards, 1927: 821. NZ (NZ).
thompsoni Tonnoir in Tonnoir & Edwards, 1927: 822. NZ (NZ).
viatoris Edwards in Tonnoir & Edwards, 1927: 822. NZ (NZ).

ATELEIA Skuse, 1888c: 1201. Type species: Ateleia spadicithorax Skuse, 1888, mon.
spadicithorax Skuse, 1888c: 1202. Australia (NSW).

CALEDONILEIA Matile, 1993: 179. Type species: Caledonileia pusilla Matile, 1993, orig. des.
pusilla Matile, 1993: 181. New Caledonia (New Caledonia).

CAWTHRONIA Tonnoir & Edwards, 1927: 825. Type species: Cawthronia nigra Tonnoir in Tonnoir & Edwards, 1927, orig. des.
nigra Tonnoir in Tonnoir & Edwards, 1927: 826. NZ (NZ).

CLASTOBASIS Skuse, 1890d: 617. Type species: Clastobasis tryoni Skuse, 1890, mon.
RHYMOLEIA Edwards, 1924f: 570 (Leiomyia subg.). Type species: Leiomyia fijiana Edwards, 1924, orig. des.
fijiana Edwards, 1924f: 570 (Leiomyia). Fiji.
     fijiensis, error for fijiana.
ridens Colless, 1966: 654 (Leia). Bonin I.
stylata Matile, 1993: 176. New Caledonia (New Caledonia).
tryoni Skuse, 1890d: 619. Australia (Qld).
vicina Matile, 1993: 178. New Caledonia (New Caledonia).
yoshimotoi Matile, 1993: 172. New Caledonia (New Caledonia).

CYCLONEURA Marshall, 1896b: 289. Type species: Cycloneura flava Marshall, 1896, mon.
flava Marshall, 1896b: 289. NZ (NZ).
peterjohnsi Jaschhof & Kallweit, 2009: 17. NZ (NZ).
separata Jaschhof & Kallweit, 2009: 14. NZ (NZ).
similis Jaschhof & Kallweit, 2009: 12. NZ (NZ).
triangulata Tonnoir in Tonnoir & Edwards, 1927: 825. NZ (NZ).

GRACILILEIA Matile, 1993: 181. Type species: Gracilileia redunda Matile, 1993, orig. des.
aurata Matile, 1993: 186. New Caledonia (New Caledonia).
lydiae Matile, 1993: 169. New Caledonia (New Caledonia).
montana Matile, 1993: 184. New Caledonia (New Caledonia).
redunda Matile, 1993: 183. New Caledonia (New Caledonia).
tillierorum Matile, 1993: 186. New Caledonia (New Caledonia).

INDOLEIA Edwards, 1928b: 6 (Leia subg.). Type species: Leia bisetosa Edwards, 1928, orig. des.
demeijerei Matile, 1989: 140 (n. n. for nigra de Meijere). Indonesia (Maluku).
     nigra de Meijere, 1913a: 46 (Leia). Indonesia (Maluku). [Preocc. Zetterstedt, 1838.]

LEIA Meigen, 1818: 253. Type species: Leia fascipennis Meigen, 1818, des. Curtis, 1837a: 645.
LEJOMYA Rondani, 1856: 195 (unjust. n.n. for Leia Meigen).
GLAPHYROPTERA Winnertz, 1863: 771. Type species: Leia fascipennis Meigen, 1818, des. Coquillett, 1910: 547. [Preocc. Herr, 1852.]
NEOGLAPHYROPTERA Osten Sacken, 1878: 10, 216 (n.n. for Glaphyroptera Winnertz).
LEJA Scudder, 1891: 630 (unjust. n.n. for Leia Meigen).
arsona Hutson, 1978: 123. Soth Africa; NZ (NZ) [introduced]; Europe, Kenya, St. Helena.
delobeli Matile, 1993: 175. New Caledonia (New Caledonia).

Genus LEPTOMORPHUS Curtis

LEPTOMORPHUS Curtis, 1831: pl. 365. Type species: Leptomorphus walkeri Curtis, 1831, orig. des.
alienus Papp & Sevcik, 2011: 141.
papua Papp & Sevcik, 2011: 152.

PARACYCLONEURA Tonnoir & Edwards, 1927: 825. Type species: Paracycloneura apicalis Tonnoir in Tonnoir & Edwards, 1927, orig. des.
apicalis Tonnoir in Tonnoir & Edwards, 1927: 825. NZ (NZ).
inopinata Jaschhof & Kallweit, 2009: 26. NZ (NZ).

PARADOXA Marshall, 1896b: 290. Type species: Paradoxa fusca Marshall, 1896, mon.
fusca Marshall, 1896b: 290. NZ (NZ); NZ (Auckland Is).

PARALEIA Tonnoir, 1929b: 611. Type species: Paraleia fulvescens Tonnoir, 1929, orig. des.
fulvescens Tonnoir, 1929b: 611. Australia (Tas); Australia (NSW, Qld).

SIGMOLEIA Tonnoir & Edwards, 1927: 826. Type species: Sigmoleia melanoxantha Edwards in Tonnoir & Edwards, 1927, orig. des.
melanoxantha Edwards in Tonnoir & Edwards, 1927: 826. NZ (NZ).
minuta Matile, 1993: 173. New Caledonia (New Caledonia).
spinosistyla Matile, 1993: 172. New Caledonia (New Caledonia).

STICHOLEIA Söli, 1996: 2. Type species: Sticholeia cheesmanae Söli, 1996, orig. des.
cheesmanae Söli, 1996: 4. Vanuatu.
dolichostyla Söli, 1996: 6. PNG (PNG); Indonesia (Irian Jaya).

TETRAGONEURA Winnertz, 1846: 18. Type species: Tetragoneura distincta Winnertz, 1846 [= Sciophila sylvatica Curtis, 1837], des. Johannsen, 1909: 34.
annulicornis Matile, 1993: 189. New Caledonia (New Caledonia).
auriculata Matile, 1993: 191. New Caledonia (New Caledonia).
baylaci Matile, 1993: 193. New Caledonia (New Caledonia).
boucheti Matile, 1993: 193. New Caledonia (New Caledonia).
bourgoini Matile, 1993: 195. New Caledonia (New Caledonia).
chazeaui Matile, 1993: 196. New Caledonia (New Caledonia).
exigua Matile, 1989: 140 (n. n. for minuta Tonnoir). NZ (NZ).
     minuta Tonnoir in Tonnoir & Edwards, 1927: 832. NZ (NZ). [Preocc. Meunier, 1904.]
falliciosa Matile, 1993: 198. New Caledonia (New Caledonia).
flexa Edwards in Tonnoir & Edwards, 1927: 833. NZ (NZ).
fusca Tonnoir in Tonnoir & Edwards, 1927: 829. NZ (NZ).
hirsuticauda Matile, 1993: 199. New Caledonia (New Caledonia).
minima Tonnoir in Tonnoir & Edwards, 1927: 830. NZ (NZ); NZ (Auckland Is, Campbell I).
nigra Marshall, 1896b: 286. NZ (NZ).
obliqua Edwards in Tonnoir & Edwards, 1927: 831. NZ (NZ).
obscura Tonnoir in Tonnoir & Edwards, 1927: 832. NZ (NZ).
occulta Matile, 1993: 203. New Caledonia (New Caledonia).
opaca Tonnoir in Tonnoir & Edwards, 1927: 829. NZ (NZ).
pervigila Matile, 1993: 200. New Caledonia (New Caledonia).
proxima Tonnoir in Tonnoir & Edwards, 1927: 831. NZ (NZ).
reducta Matile, 1993: 204. New Caledonia (New Caledonia).
rufipes Tonnoir in Tonnoir & Edwards, 1927: 833. NZ (NZ); NZ (Auckland Is, Campbell I).
spinipes Edwards in Tonnoir & Edwards, 1927: 830. NZ (NZ).
tonnoiri Matile, 1989: 141 (n. n. for distincta Tonnoir). NZ (NZ).
     distincta Tonnoir in Tonnoir & Edwards, 1927: 832. NZ (NZ). [Preocc. Winnertz, 1846.]
ultima Tonnoir in Tonnoir & Edwards, 1927: 833. NZ (NZ)
usitata Matile, 1993: 201. New Caledonia (New Caledonia).
venusta Tonnoir in Tonnoir & Edwards, 1927: 831. NZ (NZ)

TRICHOTERGA Tonnoir & Edwards, 1927: 827. Type species: Trichoterga monticola Tonnoir in Tonnoir & Edwards, 1927, orig. des.
monticola Tonnoir in Tonnoir & Edwards, 1927: 827. NZ (NZ).
     ssp. incisurata Edwards in Tonnoir & Edwards, 1927: 828 (monticola var.). NZ (NZ).
     ssp. monticola s.str. NZ (NZ).

Genus TONNWARDSIA Jaschhof & Kallweit

TONNWARDSIA Jaschhof & Kallweit, 2009: 27. Type species: Cycloneura aberrans Tonnoir, 1927, orig. des.
aberrans Tonnoir in Tonnoir & Edwards, 1927: 824 (Cycloneura). NZ (NZ).

Genus WAIPAPOMYIA Jaschhof & Kallweit

WAIPAPOMYIA Jaschhof & Kallweit, 2009: 31. Type species: Waipapomyia dentata Jaschhof & Kallweit, 2009, orig. des.
dentata Jaschhof & Kallweit, 2009: 33. NZ (NZ).
elongata Jaschhof & Kallweit, 2009: 33. NZ (NZ).
truncata Jaschhof & Kallweit, 2009: 34. NZ (NZ).

fulva Walker, 1856d: 416 (Leia). Australia (Tas).

Subfamily MANOTINAE
Genus EUMANOTA Edwards

EUMANOTA Edwards, 1933: 231. Type species: Eumanota leucrura Edwards, 1933, orig. des.
jani Papp, 2004: 228. PNG (PNG).
malukuensis Søli, 2002: 50. Indonesia (Maluku); Indonesia (Nusa Tenggara).
vilkamaai Hippa & Sevcik, 2010: 19. PNG (PNG).

Genus MANOTA Williston

MANOTA Williston, 1896b: 260. Type species: Manota defecta Williston, 1896, mon.
acris Kurina & Hippa, 2015: 262. PNG (PNG).
alulata Kurina & Hippa, 2015: 260. PNG (PNG).
apentachaeta Kurina & Hippa, 2015: 264. PNG (PNG).
bicuspis Hippa, 2007: 32. PNG (PNG); Solomon Is.
biunculata Hippa, 2007: 25. PNG (PNG).
clivicola Kurina & Hippa, 2015: 267. PNG (PNG).
cordata Kurina & Hippa, 2015: 267. PNG (PNG).
ctenophora Matile, 1993: 208. New Caledonia (New Caledonia).
evexa Hippa, 2007: 36. PNG (PNG).
explicans Hippa, 2007: 17. PNG (PNG).
feminea Kurina & Hippa, 2015: 269. PNG (PNG).
gemella Hippa, 2007: 28. Indonesia (Maluku).
hamulata Colless, 1966: 656. Belau; PNG (PNG).
hirsuta Hippa, 2007: 23. PNG (PNG).
kaindienis Kurina & Hippa, 2015: 271. PNG (PNG).
lunata Kurina & Hippa, 2015: 272. PNG (PNG).
maorica Edwards in Tonnoir & Edwards, 1927: 801. NZ (NZ).
nimia Kurina & Hippa, 2015: 275. Solomn Is.
orthacantha Hippa, 2007: 38. PNG (PNG).
pacifica Edwards, 1928c: 30. Samoa.
parilis Hippa, 2007: 34. PNG (PNG); Vanuatu.
pentacantha Hippa, 2007: 42. Solomon Is.
pentachaeta Kurina & Hippa, 2015: 275. PNG (PNG).
perissochaeta Hippa, 2007: 14. Solomon Is; PNG (PNG).
serawei Hippa, 2007: 8. PNG (PNG).
siciliculata Kurina & Hippa, 2015: 278. PNG (PNG).
sicula Hippa, 2007: 40. PNG (PNG).
sigma Kurina & Hippa, 2015: 280. PNG (PNG).
spathula Hippa, 2007: 19. PNG (PNG).
subspathula Hippa, 2007: 21. PNG (PNG); Australia (Qld).
taedia Matile, 1993: 206. New Caledonia (New Caledonia).
tricuspis Hippa, 2007: 17. Fiji.
unisetata Kurina & Hippa, 2015: 282. Solomon Is.

ALLODIA Winnertz, 1863: 826. Type species: Mycetophila ornaticollis Meigen, 1818 [= Mycetophila lugens Wiedemann, 1817], des. Johannsen, 1909: 104.
sabroskyi Colless, 1966: 659. Belau.

ANATELLA Winnertz, 1863: 854. Type species: Anatella gibba Winnertz, 1863, des. Johannsen, 1909: 101.
Unidentified sp.--New Caledonia (New Caledonia).

BRACHYRADIA Sevcik & Kjaerandsen, 2012: 118. Type species: Brachyradia asiatica Sevcik & Kjaerandsen, 2012, orig. des.
australis Sevick &Kjaerandsen, 2012: 121. PNG (PNG); Indonesia (Sulawesi).

BREVICORNU Marshall, 1896b: 307. Type species: Brevicornu flavum Marshall, 1896, des. Tuomikoski, 1966a: 184.
antennata Harrison, 1964: 304 (Allodia). NZ (Campbell I); NZ (Auckland Is).
brunnea Harrison, 1964: 30 (Allodia). NZ (Campbell I).
flavum Marshall, 1896b: 307. NZ (NZ).
fragile Marshall, 1896b: 308. NZ (NZ).
maculatum Tonnoir in Tonnoir & Edwards, 1927: 834 (Allodia). NZ (NZ).
marshalli Zaitzev, 2002: 12. NZ (NZ).
matilei Zaitzev, 2002: 12. NZ (NZ).
quadriseta Edwards in Tonnoir & Edwards, 1927: 835 (Allodia). NZ (NZ).
rufithorax Tonnoir in Tonnoir & Edwards, 1927: 834 (Allodia). NZ (NZ).
subrufithorax Zaitzev, 2002: 13. NZ (NZ).
tongariro Zaitzev, 2002: 13. NZ (NZ).

Genus CORDYLA Meigen

CORDYLA Meigen, 1803: 263. Type species: Cordyla fusca Meigen, 1804, subs. mon. Meigen, 1804: 93.
australicaKurtina & Oliveria, 2013: 31. Australia (Tas); Australia (NSW, Qld Vic).
jani Kurina, 2005: 403. PNG (PNG).
toraia Kurina, 2005: 405. Indonesia (Sulawesi); PNG (PNG).

EXECHIA Winnertz, 1863: 879. Type species: Tipula fungorum De Geer, 1776 [= Mycetophila fusca Meigen, 1804], des. Johannsen, 1909: 106.
BRACHYDICRANIA Skuse, 1888c: 1215. Type species: Brachydicrania fumosa Skuse, 1888, des. Matile, 1989: 142.
abbreviata Skuse, 1888c: 1219 (Brachydicrania). Australia (NSW).
biseta Edwards in Tonnoir & Edwards, 1927: 837. NZ (NZ).
fascipennis Skuse, 1890d: 628 (Brachydicrania). Australia (NSW).
filata Edwards in Tonnoir & Edwards, 1927: 836. NZ (NZ).
fumosa Skuse, 1888c: 1218 (Brachydicrania). Australia (NSW).
hiemalis Marshall, 1896b: 300 (Brachydicrania). NZ (NZ); NZ (Auckland Is, Campbell I).
howesi Edwards in Tonnoir & Edwards, 1927: 836. NZ (NZ).
lutacea Edwards, 1928c: 31. Western Samoa.
novaezelandiae Tonnoir in Tonnoir & Edwards, 1927: 836. NZ (NZ).
pictiventris Skuse, 1888c: 1217 (Brachydicrania). Australia (NSW).
pullicauda Skuse, 1888c: 1216 (Brachydicrania). Australia (NSW).
snyderi Colless, 1966: 657. Bonin Is.
thomsoni Miller, 1918: 4. NZ (NZ).

SYNPLASTA Skuse, 1890d: 629. Type species: Synplasta annuliventris Skuse, 1890, mon.
ALLODIOPSIS Tuomikoski, 1966a: 171. Type species: Rymosia rustica Edwards, 1941, orig. des.
annuliventris Skuse, 1890d: 630. Australia (NSW).
sydneyensis Skuse, 1890d: 627 (Dynatosoma). Australia (NSW).

ASPIDIONIA Colless, 1966: 664. Type species: Aspidionia palauensis Colless, 1966, orig. des.
palauensis Colless, 1966: 664. Belau.

EPICYPTA Winnertz, 1863: 909. Type species: Mycetophila scatophora Perris, 1849, des. Johannsen, 1909: 110.
DELOPSIS Skuse, 1890d: 623. Type species: Delopsis flavipennis Skuse, 1890, mon.
angusticollaris Edwards 1929h: 74 (Delopsis). Vanuatu; Philippines.
boninensis Colless, 1966: 664. Bonin Is.
buxtoni Edwards, 1928c: 32 (Delopsis). Western Samoa; Indonesia (Sumatra).
flavipennis Skuse, 1890d: 624 (Delopsis). Australia (NSW).
greenwoodi Edwards, 1924f: 570 (Delopsis). Fiji.
palauensis Colless, 1966: 662. Belau.
scutellaris Edwards, 1928c: 33 (Delopsis). Western Samoa.

MYCETOPHILA Meigen, 1803: 263. Type species: Tipula agarici Villers, 1789, des. Johannsen, 1909: 110.
aequalis Walker, 1856d: 415. Australia (Tas).
campbellensis Harrison, 1964: 307. NZ (Campbell I); NZ (Auckland Is).
clara Tonnoir in Tonnoir & Edwards, 1927: 855. NZ (NZ).
colorata Tonnoir in Tonnoir & Edwards, 1927: 854. NZ (NZ).
conica Tonnoir in Tonnoir & Edwards, 1927: 845. NZ (NZ).
consobrina Tonnoir in Tonnoir & Edwards, 1927: 845. NZ (NZ).
crassitarsis Edwards in Tonnoir & Edwards, 1927: 853. NZ (NZ).
curtisi Edwards in Tonnoir & Edwards, 1927: 841. NZ (NZ).
diffusa Tonnoir in Tonnoir & Edwards, 1927: 849. NZ (NZ).
dilatata Tonnoir in Tonnoir & Edwards, 1927: 854. NZ (NZ).
elegans Tonnoir in Tonnoir & Edwards, 1927: 842. NZ (NZ).
elongata Tonnoir in Tonnoir & Edwards, 1927: 845. NZ (NZ).
fagi Marshall, 1896b: 303. NZ (NZ).
     variabilis Marshall, 1896b: 304. NZ (NZ).
     robusta Marshall, 1896b: 305. NZ (NZ).
filicornis Tonnoir in Tonnoir & Edwards, 1927: 857. NZ (NZ).
fumosa Tonnoir in Tonnoir & Edwards, 1927: 852. NZ (NZ); NZ (Auckland Is).
furtiva Tonnoir in Tonnoir & Edwards, 1927: 857. NZ (NZ).
grandis Tonnoir in Tonnoir & Edwards, 1927: 851. NZ (NZ).
griseofusca Tonnoir in Tonnoir & Edwards, 1927: 852. NZ (NZ).
     ssp. griseofusca s.str. NZ (NZ).
     ssp. nigriclava Tonnoir in Tonnoir & Edwards, 1927: 852 (griseofusca var.). NZ (NZ).
grisescens Edwards in Tonnoir & Edwards, 1927: 850. NZ (NZ).
harrisi Edwards in Tonnoir & Edwards, 1927: 859. NZ (NZ).
howletti Marshall, 1896: 302. NZ (NZ).
impunctata Edwards in Tonnoir & Edwards, 1927: 856. NZ (NZ).
integra Tonnoir in Tonnoir & Edwards, 1927: 858. NZ (NZ).
intermedia Edwards in Tonnoir & Edwards, 1927: 860. NZ (NZ).
latifascia Edwards in Tonnoir & Edwards, 1927: 843. NZ (NZ).
lomondensis Edwards in Tonnoir & Edwards, 1927: 850. NZ (NZ).
luteolateralis Edwards in Tonnoir & Edwards, 1927: 853. NZ (NZ).
marginepunctata Tonnoir in Tonnoir & Edwards, 1927: 847. NZ (NZ); NZ (Auckland Is).
     ssp. marginepunctata s.str. NZ (NZ); NZ (Auckland Is).
     ssp. rotundipennis Tonnoir in Tonnoir & Edwards, 1927: 847 (marginepunctata var.). NZ (NZ).
     ssp. ruapehensis Tonnoir in Tonnoir & Edwards, 1927: 847 (marginepunctata var.). NZ (NZ).
marshalli Enderlein, 1910a: 174 (n. n. for maculata Marshall). NZ (NZ).
     maculata Marshall, 1896b: 306. NZ (NZ). [Preocc. Macquart, 1834.]
media Tonnoir in Tonnoir & Edwards, 1927: 858. NZ (NZ).
minima Edwards in Tonnoir & Edwards, 1927: 846. NZ (NZ).
nigricans Tonnoir in Tonnoir & Edwards, 1927: 850. NZ (NZ).
nigripalpis Edwards in Tonnoir & Edwards, 1927: 848. NZ (NZ).
nigriventris Skuse, 1890d: 622. Australia (NSW).
nitens Tonnoir in Tonnoir & Edwards, 1927: 847. NZ (NZ).
nitidula Tonnoir in Tonnoir & Edwards, 1927: 848. NZ (NZ).
ornatissima Tonnoir in Tonnoir & Edwards, 1927: 841. NZ (NZ).
phyllura Edwards in Tonnoir & Edwards, 1927: 849. NZ (NZ).
pollicata Edwards in Tonnoir & Edwards, 1927: 853. NZ (NZ).
propria Skuse, 1888c: 1213. Australia (NSW).
pseudomarshalli Tonnoir in Tonnoir & Edwards, 1927: 846. NZ (NZ).
solitaria Tonnoir in Tonnoir & Edwards, 1927: 855. NZ (NZ).
spinigera Tonnoir in Tonnoir & Edwards, 1927: 859. NZ (NZ).
submarshalli Tonnoir in Tonnoir & Edwards, 1927: 846. NZ (NZ).
subnitens Edwards in Tonnoir & Edwards, 1927: 848. NZ (NZ).
subspinigera Tonnoir in Tonnoir & Edwards, 1927: 851. NZ (NZ).
subtenebrosa Tonnoir in Tonnoir & Edwards, 1927: 859. NZ (NZ).
subtillis Tonnoir in Tonnoir & Edwards, 1927: 849. NZ (NZ).
sylvatica Marshall, 1896b: 301. NZ (NZ).
tapleyi Edwards in Tonnoir & Edwards, 1927: 854. NZ (NZ).
tenebrosa Edwards in Tonnoir & Edwards, 1927: 860. NZ (NZ).
tonnoiri Matile, 1989: 144 (n. n. for similis Tonnoir). NZ (NZ).
     similis Tonnoir in Tonnoir & Edwards, 1927: 842. NZ (NZ). [Preocc. Santos Abreu, 1920.]
trispinosa Tonnoir in Tonnoir & Edwards, 1927: 844. NZ (NZ).
unispinosa Tonnoir in Tonnoir & Edwards, 1927: 856. NZ (NZ).
virgata Tonnoir in Tonnoir & Edwards, 1927: 843. NZ (NZ).
viridis Edwards in Tonnoir & Edwards, 1927: 852. NZ (NZ).
vulgaris Tonnoir in Tonnoir & Edwards, 1927: 844. NZ (NZ).

PHRONIA Winnertz, 1863: 857. Type species: Phronia rustica Winnertz, 1863 [= Mycetophila exigua Zetterstedt, 1852], des. Johannsen, 1909: 96.
boninensis Colless, 1966: 660. Bonin Is.

PLATUROCYPTA Enderlein, 1910b: 76. Type species: Platurocypta limbatifemur Enderlein, 1910, orig. des.
dilatata Tonnoir in Tonnoir & Edwards, 1927: 861 (Epicypta). NZ (NZ).
immaculata Tonnoir in Tonnoir & Edwards, 1927: 860 (Epicypta). NZ (NZ).

PSEUDALYSIINA Tonnoir, 1929b: 608. Type species: Pseudalysiina mimicans Tonnoir, 1929, orig. des.
mimicans Tonnoir, 1929b: 609. Australia (Tas).

SCEPTONIA Winnterz, 1863: 907. Type species: Mycetophila nigra Meigen, 1804, des. Johannsen, 1909: 113.
ornatithorax Skuse, 1890d: 621. Australia (NSW).

TRICHONTA Winnertz, 1863: 847. Type species: Mycetophila melanura Staeger, 1840, des. Johannsen, 1909: 94.
illaetabilis Skuse, 1888c: 1210. Australia (NSW).
vegeta Skuse, 1888c: 1209. Australia (NSW).

ZYGOMYIA Winnterz, 1863: 901. Type species: Mycetophila vara Staeger, 1840, des. Johannsen, 1909: 112.
acuta Tonnoir in Tonnoir & Edwards, 1927: 870. NZ (NZ).
albinotata Tonnoir in Tonnoir & Edwards, 1927: 870. NZ (NZ).
apicalis Tonnoir in Tonnoir & Edwards, 1927: 866. NZ (NZ).
bifasciola Matile, 1989: 145 (n.n. for bifasciata Tonnoir). NZ (NZ).
     bifasciata Tonnoir in Tonnoir & Edwards, 1927: 863. NZ (NZ). [Preocc. Garrett, 1925.]
bivittata Tonnoir in Tonnoir & Edwards, 1927: 864. NZ (NZ).
brunnea Tonnoir in Tonnoir & Edwards, 1927: 866. NZ (NZ).
costata Tonnoir in Tonnoir & Edwards, 1927: 865. NZ (NZ).
crassicauda Tonnoir in Tonnoir & Edwards, 1927: 867. NZ (NZ).
crassipyga Tonnoir in Tonnoir & Edwards, 1927: 868. NZ (NZ).
diffusa Tonnoir in Tonnoir & Edwards, 1927: 878, pl. 80, fig. 299. NZ (NZ).
distincta Tonnoir in Tonnoir & Edwards, 1927: 872. NZ (NZ).
egmontensis Zaitzev, 2002: 112. NZ (NZ).
eluta Edwards in Tonnoir & Edwards, 1927: 871. NZ (NZ).
filigera Edwards in Tonnoir & Edwards, 1927: 872. NZ (NZ).
flavicoxa Marshall, 1896b: 297. NZ (NZ).
fusca Marshall, 1896: 302. NZ (NZ).
grisescens Tonnoir in Tonnoir & Edwards, 1927: 865. NZ (NZ).
guttata Tonnoir in Tonnoir & Edwards, 1927: 868. NZ (NZ).
immaculata Tonnoir in Tonnoir & Edwards, 1927: 863. NZ (NZ).
longicauda Tonnoir in Tonnoir & Edwards, 1927: 867. NZ (NZ).
marginata Tonnoir in Tonnoir & Edwards, 1927: 869. NZ (NZ); NZ (Auckland Is).
multiseta Zaitzev, 2002: 114. NZ (NZ).
nigrita Tonnoir in Tonnoir & Edwards, 1927: 871. NZ (NZ).
nigriventris Tonnoir in Tonnoir & Edwards, 1927: 866. NZ (NZ).
nigrohalterata Tonnoir in Tonnoir & Edwards, 1927: 865. NZ (NZ).
obsoleta Tonnoir in Tonnoir & Edwards, 1927: 864. NZ (NZ).
ovata Zaitzev, 2002: 116. NZ (NZ).
penicillata Edwards in Tonnoir & Edwards, 1927: 872. NZ (NZ).
ruficollis Tonnoir in Tonnoir & Edwards, 1927: 866. NZ (NZ).
rufithorax Tonnoir in Tonnoir & Edwards, 1927: 867. NZ (NZ).
similis Tonnoir in Tonnoir & Edwards, 1927: 863. NZ (NZ).
submarginata Zaitzev, 2002: 116. NZ (NZ).
taranakiensis Zaitzev, 2002: 116. NZ (NZ).
tonnoiri Plassmann, 1970: 399 (n.n. for humeralis Tonnoir). NZ (NZ).
     humeralis Tonnoir in Tonnoir & Edwards, 1927: 869. NZ (NZ). [Preocc. Wiedemann, 1817.]
trifasciata Tonnoir in Tonnoir & Edwards, 1927: 871. NZ (NZ).
trispinosaZaitzev, 2002: 119. NZ (NZ).
truncata Tonnoir in Tonnoir & Edwards, 1927: 870. NZ (NZ).
unispinosa Tonnoir in Tonnoir & Edwards, 1927: 870. NZ (NZ).
varipes Edwards in Tonnoir & Edwards, 1927: 868. NZ (NZ).

ZYGOPHRONIA Edwards, 1928c: 31. Type species: Zygophronia pusilla Edwards, 1928, mon.
pusilla edwards, 1928c: 31. Western Samoa.

This page last revised 17 April 2017