Sciaridae are common faunal elements in nearly all regions of the world
but are poorly known because their small size and superficial homogeneity
have not attracted collectors or taxonomists. Immature stages can be found
in rotten wood, under bark of fallen trees, in other kinds of decaying plant
material, feeding in fungi or animal excrement or, in Austrosciara termitophila,
as inquilines in termite nests. One genus (Vulagisciara) and new species has been found as a guest in rubiaceous ant-plants in Fiji (Evenhuis et al., 2007.). Some species feed on living plant tissue
and can cause economically important damage, especially in greenhouses and
mushroom houses.
Recent studies indicate that the Australasian/Oceanian fauna is rich in taxa. The regional literature, with the exception of
the faunal studies of Hawaii (Hardy 1960) and Micronesia (Steffan 1969),
consists primarily of descriptions of new species. Skuse (1888b, 1890d)
described 61 Sciaridae from Australia, but most of the collections were
from New South Wales only. The Papua New Guinean fauna has been studied by Mohrig (1999, 2004). The New Zealand fauna was treated by Tonnoir & Edwards
(1927) and most recently revised by Mohrig & Jaschhof (1999), and thirteen new species of Bradysia were described by Vilkamaa et al. (2012) from New Caledonia.
This catalog is essentially a list of published names, although some necessarily
obvious taxonomic changes have been made. Many older generic placements undoubtedly
are incorrect, and types need to be examined before correct generic assignments
can be made. Most of the species listed under Sciara should be transferred
to other genera after examination of the types.
Refs.: Hardy (1960, Hawaiian spp.), Freeman (1983), Frey (1942, 1948),
Steffan (1966b, rev. Nearctic gen.; 1969, Micronesian spp.), Tuomikoski
(1960), Evenhuis (2007 et al., Fiji sciarids), Vilkamaa et al. (2012, New Caledonian Bradysia).
Genus AERUMNOSA Mohrig
AERUMNOSA Mohrig, 1999: 158. Type species: Aerumnosa noctinigra Mohrig, 1999, orig. des.
furcillata Mohrig, 1999: 163. PNG (PNG).
noctinigra Mohrig, 1999: 159. PNG (PNG).
nocturna Mohrig, 1999: 161. PNG (PNG).
perfida Mohrig, 1999: 160. PNG (PNG).
rhinocerata Mohrig, 1999: 165. PNG (PNG).
trichovenosa Mohrig, 1999: 164. PNG (PNG).
AUSTROSCIARA Schmitz & Mjöberg, 1924: 1. Type species:
Austrosciara termitophila Schmitz & Mjöberg, 1924, mon.
CTENOSCIARA Tuomikoski, 1960: 110. Type species: Sciara hyalipennis Meigen, 1804, orig. des.
abdita Mohrig, 2014: 134 (Ctenosciara). PNG (PNG).
certa Mohrig, 2014: 124 (Ctenosciara). PNG (PNG).
constrictans Edwards in Tonnoir & Edwards, 1927: 792 (Sciara).
NZ (NZ).
nubeculosa Edwards in Tonnoir & Edwards, 1927: 793 (Sciara). NZ
(NZ).
cracensVilkamaa et al., 2012: 38 (Ctenosciara). New Caledonia (New Caledonia).
crinita Vilkamaa et al., 2012: 41 (Ctenosciara). New Caledonia (New Caledonia).
depilis Vilkamaa et al., 2012: 41 (Ctenosciara). New Caledonia (New Caledonia).
depressa Vilkamaa et al., 2012: 44 (Ctenosciara). New Caledonia (New Caledonia).
etorutao Köhler in Köhler & Mohrig, 2016: 9 (Ctenosciara). NZ (NZ).
exilis Vilkamaa et al., 2012: 45 (Ctenosciara). New Caledonia (New Caledonia).
griseinervis Edwards in Tonnoir & Edwards, 1927: 793 (Sciara).
NZ (NZ).
hawaiiensis Hardy, 1956c: 78 (Sciara). Hawaiian Is.
hyalipennis Meigen, 1804: 99 (Sciara). Not given [Central
Europe; see Gerbachevskaya-Pavluchenko, 1986: 43]; NZ (Campbell I, NZ) [immigrant];
Taiwan, widesp. Palearctic Reg.
annulata Meigen, 1818: 284 (Sciara). Not given [Germany].
inferma Mohrig, 2014: 130 (Ctenosciara). PNG (PNG).
inflata Vilkamaa et al., 2012: 45 (Ctenosciara). New Caledonia (New Caledonia).
infrequens Skuse, 1888b: 719 (Sciara). Australia (NSW).
lobigera Vilkamaa et al., 2012: 47 (Ctenosciara). New Caledonia (New Caledonia).
lubrica Mohrig, 2014: 133 (Ctenosciara). PNG (PNG).
montivaga Skuse, 1888b: 710 (Sciara). Australia (NSW).
multiformis Mohrig, 2014: 129 (Ctenosciara). PNG (PNG).
multispinosa Steffan, 1969: 712 (Ctenosciara). Micronesia.
nigrostyla Mohrig in Mohrig & Jaschhof, 1999: 1 (Ctenosciara)7. NZ (NZ).
nudopteryx Mohrig in Mohrig & Jaschhof, 1999: 19 (Ctenosciara). NZ (NZ).
obesa Vilkamaa et al., 2012: 49. New Caledonia (New Caledonia).
ovalis Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ (NZ).
pedibusa Mohrig, Kauscke & Keller, 2017: 362. Australia (Vic).
pullata Mohrig, 2014: 127 (Ctenosciara). PNG (PNG).
semipilosa Mohrig, 2014: 131 (Ctenosciara). PNG (PNG).
spectabilis Skuse, 1888b: 716 (Sciara). Australia (NSW).
subabdita Mohrig, 2014: 136 (Ctenosciara). PNG (PNG).
subpullata Mohrig, 2014: 128 (Ctenosciara). PNG (PNG).
termitophila Schmitz & Mjöberg, 1924: 2. Australia (NSW).
uniocolorata Mohrig, 2014: 126 (Ctenosciara). PNG (PNG).
winnertzi Skuse, 1888b: 709 (Sciara). Australia (NSW); NZ(NZ); China.
rufulenta Edwards in Tonnoir & Edwards, 1927: 794 (Sciara). NZ
(NZ).
vicarians Edwards in Tonnoir & Edwards, 1927: 793 (Sciara). NZ
(NZ).
xanthonota Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ
(NZ).
BRADYSIA Winnertz, 1867: 180. Type species: Bradysia angustipennis
Winnertz, 1867, des. Enderlein, 1911c: 127.
aequispina Vilkamaa et al., 2012: 26. New Caledonia (New Caledonia).
amabilis Skuse, 1888b: 712 (Sciara). Australia (NSW).
angustopalpa Mohrig, 2016: 29. PNG (PNG).
aspera Mohrig, 2016: 6. PNG (PNG).
aspercerca Mohrig, 2016: 5. PNG (PNG).
beata Mohrig, 2016: 25. PNG (PNG).
bishopi Steffan, 1973: 353. Hawaiian Is; American Samoa, Fiji, French Polynesia, New Caledonia (New Caledonia); Puerto Rico, Seychelles, Thailand.
boninensis Steffan, 1969: 717. Bonin Is.
bullata Mohrig, 2016: 21. PNG (PNG).
campbellensis Steffan, 1964: 296. NZ (Campbell I).
centidens Vilkamaa et al., 2012: 28. New Caledonia (New Caledonia).
circumfilata Mohrig, 2016: 26. PNG (PNG).
clavistylis Vilkamaa et al., 2012: 28. New Caledonia (New Caledonia).
conjuncta Skuse, 1890c: 403 (Sciara). Australia (NSW).
serenipennis Skuse, 1890c: 402 (Sciara). Australia (NSW).
crassicornis Skuse, 1890c: 407 (Sciara). Australia (NSW).
credula Mohrig, 2016: 29. PNG (PNG).
cristata Mohrig, 2016: 17. PNG (PNG).
exsequialis Skuse, 1890c: 409 (Sciara). Australia (NSW).
fantula Mohrig, 2016: 11. PNG (PNG).
filigera Vilkamaa et al., 2012: 30. New Caledonia (New Caledonia).
flaveola Mohrig, 2016: 14. PNG (PNG).
flavocristata Mohrig, 2016: 15. PNG (PNG).
frequens Skuse, 1888b: 682 (Sciara). Australia (NSW).
froggatti Skuse, 1888b: 681 (Sciara). Australia (NSW).
gemellata Vilkamaa et al., 2012: 32. New Caledonia (New Caledonia).
gibbosa Vilkamaa et al., 2012: 33. New Caledonia (New Caledonia); PNG (PNG).
impatiens Johannsen, 1912: 136 (Sciara). USA; Austrralia (ACT, NSW, Qld, SA, Tas, Vic, WA); Hawaiian I, Norfolk I; Brazil, China, Japan.
hardyi Shaw, 1952: 493 (Sciara). Hawaiian Is.
inaequispina Mohrig, 2016: 10. PNG (PNG).
incohata Mohrig, 2016: 6. PNG (PNG).
irwini Köhler & Menzel, 2013: 66. New Caledonia (New Caledonia).
kraussi Steffan, 1969: 718. Guam.
luctifica Skuse, 1888b: 680 (Sciara). Australia (SA); New Caledonia (New Caledonia), PNG (PNG)..
planistylata Vilkamaa et al., 2012: 38. New Caledonia (New Caledonia).
maesta Skuse, 1888b: 691 (Sciara). Australia (NSW).
mastersi Skuse, 1888b: 723 (Trichosia). Australia (NSW).
megahypopygialis Mohrig, 2016: 23. PNG (PNG).
melina Vilkamaa et al., 2012: 34. New Caledonia (New Caledonia).
microdentata Mohrig, 2016: 24. PNG (PNG).
molokaiensis Grimshaw, 1901: 2 (Sciara). Hawaiian Is.
stonei Shaw, 1952: 495 (Sciara). Hawaiian Is.
mutuata Mohrig, 2016: 12. PNG (PNG).
novaeseelandiae Köhler & Mohrig, 2016: 2. NZ (NZ).
nudilobata Vilkamaa et al., 2012: 36. New Caledonia (New Caledonia).
obnoxia Mohrig, 2016: 11. PNG (PNG).
ocellaris Comstock, 1882: 202 (Sciara). USA; Australia (ACT, NSW, Qld, SA, Tas, Vic, WA), Hawaiian Is, Micronesia; cosmopol.
tritici Coquillett, 1895a: 408 (Sciara). Not given [USA.
johannseni Shaw, 1952: 493 (Sciara). Hawaiian Is. [Preocc.
Enderlein, 1912.]
garreti Shaw, 1952: 494 (Sciara). Hawaiian Is.
laffooni Shaw, 1952: 494 (Sciara). Hawaiian Is.
ornatula Skuse, 1888b: 711 (Sciara). Australia (NSW).
pallipes Fabricius, 1787: 326 (Tipula). Germany; NZ (NZ) [immigrant]; Europe, Algeria.
paudicens Vilkamaa et al., 2012: 37. New Caledonia (New Caledonia).
pellistyla Mohrig, 2016: 4. PNG (PNG).
perfrigida Mohrig, 2016: 9. PNG (PNG).
pernitida Skuse, 1888b: 685 (Sciara). Australia (NSW).
pictipes Skuse, 1888b: 721 (Sciara). Australia (NSW).
notata Skuse, 1888b: 720 (Sciara). Australia (NSW).
seticornis Vilkamaa et al., 2012: 39. New Caledonia (New Caledonia).
praebullata Mohrig, 2016: 19. PNG (PNG).
pseudocampestris Mohrig--not Australasian/Oceanian.
quinquespina Mohrig, 2016: 7. PNG (PNG).
radicum Brunetti, 1912b: 139 (Sciara). India; American Samoa,
Fiji, French Polynesia (Marquesas, ?Society Is), Hawaiian Is, Western Samoa;
England.
robusta Mohrig, 2016: 27. PNG (PNG).
rubra Harrison, 1955: 213 (Sciara). NZ (Campbell I).
saetibullata Mohrig, 2016: 21. PNG (PNG).
semicristata Mohrig, 2016: 18. PNG (PNG).
similigibbosa Köhler & Menzel, 2013: 64. New Caledonia (New Caledonia).
snyderi Steffan, 1969: 723. Bonin Is.
spatitergum Hardy, 1956c: 85 (Sciara). Hawaiian Is; PNG (PNG); Brazil,
India, Madagascar, Panama, Zimbabwe.
radicum, Hardy, 1960, not Brunetti, 1912, misid.
spicotegmata Mohrig, 2016: 28. PNG (PNG).
spissa Vilkamaa et al., 2012: 41. New Caledonia (New Caledonia).
stricta Vilkamaa et al., 2012: 42. New Caledonia (New Caledonia).
subbullata Mohrig, 2016: 20. PNG (PNG).
subcristata Mohrig, 2016: 18. PNG (PNG).
suspecta Mohrig, 2016: 8. PNG (PNG).
tenuitegmata Mohrig, 2016: 15. PNG (PNG).
trispinea Mohrig, 2016: 8. PNG (PNG).
unica Skuse, 1888b: 708 (Sciara). Australia (NSW).
validospina Mohrig, 2016: 13. PNG (PNG).
vevnusta Mohrig, 2016: 22. PNG (PNG).
violenta Mohrig, 2016: 16. PNG (PNG).
watsoni Colless, 1962b: 955. Macquarie I.
Genus CHAETOSCIARA Frey
CHAETOSCIARA Frey, 1942:33 (as Neosciara Pettey subg.). Type species: Sciara fenestralis Zetterstedt, 1838 [misidentification, = Sciara estlandica Lengersdorf, 1929], by orig. des.
NEOSCIARA, authors, not Pettey, 1918, misid.
ingrata Mohrig, 1999: 199. PNG (PNG).
recondita Mohrig & Kausckhke, 2017: 453, Australia (Qld).
recta Mohrig, 1999: 198. PNG (PNG).
CHIASMATA Mohrig, 2016: 30. Type species: Chiasmata duubioproterva Mohrig, 2016, orig. des.
dubioproterva Mohrig, 2016: 30. PNG (PNG).
CORYNOPTERA Winnertz, 1867: 177. Type species: Corynoptera
perpusilla Winnertz, 1867, des. Enderlein, 1911c: 127.
alpina Mohrig—not Australasian/Oceanian.
aggregata Köhler & Mohrig, 2016: 4. NZ (NZ).
ancylospina Mohrig in Mohrig & Jaschhof, 1999: 77. NZ (NZ).
basisetosa Mohrig in Mohrig & Jaschhof, 1999: 53. NZ (NZ).
brevipalpis Steffan, 1969: 695. Belau; Hawaiian Is.
catrinjaschhofae Köhler & Mohrig, 2016: 7. NZ (NZ).
coronospina Mohrig in Mohrig & Jaschhof, 1999: 84. NZ (NZ).
cowanorum Mohrig in Mohrig & Jaschhof, 1999: 81. NZ (NZ).
denospica Mohrig in Mohrig & Jaschhof, 1999: 74. NZ (NZ).
densisetosa Mohrig in Mohrig & Jaschhof, 1999: 54. NZ (NZ).
didymistyla Mohrig in Mohrig & Jaschhof, 1999: 67. NZ (NZ).
diversicalcaria Mohrig, 2004: 163. PNG (PNG).
dividospica Mohrig in Mohrig & Jaschhof, 1999: 70. NZ (NZ).
expressospina Mohrig in Mohrig & Jaschhof, 1999: 79. NZ (NZ).
facticia Mohrig in Mohrig & Jaschhof, 1999: 49. NZ (NZ).
fatigans Johannsen, 1912: 132 (Sciara). USA; New Caledonia (New Caledonia), NZ (NZ) [immigrant]; Holarctic.
filisetosa Mohrig in Mohrig & Jaschhof, 1999: 63. NZ (NZ).
filispica Mohrig in Mohrig & Jaschhof, 1999: 75. NZ (NZ).
fuscispica Mohrig in Mohrig & Jaschhof, 1999: 71. NZ (NZ).
gladiota Mohrig, 2004: 162. PNG (PNG).
harrisi Edwards in Tonnoir & Edwards, 1927: 797 (Sciara). NZ (NZ).
hemisetosa Mohrig in Mohrig & Jaschhof, 1999: 64. NZ (NZ).
heterochela Steffan, 1969: 697. Micronesia; Belau.
latistylata Hardy, 1956c: 82 (Sciara). Hawaiian Is; Marshall Is,
Northern Marianas.
microsetosa Mohrig in Mohrig & Jaschhof, 1999: 62. NZ (NZ).
minutela Skuse, 1888b: 702 (Sciara). Australia (NSW).
nigrospina Mohrig in Mohrig & Jaschhof, 1999: 45. NZ (NZ).
nigrotegminis Mohrig in Mohrig & Jaschhof, 1999: 56. NZ (NZ).
oririclausa Mohrig in Mohrig & Jaschhof, 1999: 65. NZ (NZ).
parasetosa Mohrig in Mohrig & Jaschhof, 1999: 58. NZ (NZ).
pentaspina Mohrig in Mohrig & Jaschhof, 1999: 47. NZ (NZ).
plasiosetosa Mohrig in Mohrig & Jaschhof, 1999: 59. NZ (NZ).
ponapensis Steffan, 1969: 703. Micronesia.
praegladiota Mohrig, 2004: 162. PNG (PNG).
prinospina Mohrig in Mohrig & Jaschhof, 1999: 80. NZ (NZ).
priscospina Mohrig in Mohrig & Jaschhof, 1999: 86. NZ (NZ).
pronospica Mohrig in Mohrig & Jaschhof, 1999: 72. NZ (NZ).
propriospina Mohrig in Mohrig & Jaschhof, 1999: 87. NZ (NZ).
prosopspina Mohrig in Mohrig & Jaschhof, 1999: 82. NZ (NZ).
psilospina Mohrig in Mohrig & Jaschhof, 1999: 83. NZ (NZ).
quasisetosa Mohrig in Mohrig & Jaschhof, 1999: 57. NZ (NZ).
sabroskyi Steffan, 1969: 703. Belau.
semiaggregata Mohrig in Mohrig & Jaschhof, 1999: 61. NZ (NZ).
subantarctica Steffan, 1964: 295. NZ (Campbell I).
sylviae Steffan, 1969: 705. Belau; Micronesia.
tapleyi Edwards in Tonnoir & Edwards, 1927: 797 (Sciara). NZ (NZ).
variospina Mohrig in Mohrig & Jaschhof, 1999: 50. NZ (NZ).
Genus COSMOSCIARA Frey
COSMOSCIARA Frey, 1942: 39. Type species: Plastosciara perniciosa Edwards, 1922 [= Sciara hartii Johannsen, 1912], orig. des.
hartii Johannsen, 1912: 144 (Sciara). USA;
Australia (WA), Fiji, French Polynesia (Marquesas),
Hawaiian Is, Western Samoa; cosmopol..
perniciosa Edwards, 1922c: 160 (Plastosciara). England.
brevicalcarata Hardy, 1956c: 73 (Plastosciara). Hawaiian Is.
Genus CRATYNA Winnerz
CRATYNA Winnertz, 1867: 167. Type species: Cratyna atra Winnertz, 1867, orig. des.
adulterina Mohrig & Kauschke, 2017: 456. Australia ( Qld).
compta Mohrig, 1999: 173. PNG (PNG).
consensa Mohrig, 1999: 172. PNG (PNG).
consentanea Mohrig, 1999: 169. PNG (PNG).
distorta Mohrig, 1999: 168. PNG (PNG).
ficta Mohrig, 1999: 159. PNG (PNG).
flagria Mohrig, 1999: 179. PNG (PNG); Australia (Qld).
flagriola Mohrig & Kauschke, 2017: 457. Australia (Qld).
flagriantennata Mohrig, 1999: 178. PNG (PNG).
flagritissima Mohrig, 1999: 180. PNG (PNG).
flavivaria Mohrig, 1999: 170. PNG (PNG).
flavothoracica Mohrig & Kauschke, 2017: 460. Australia (Qld).
interflagria Mohrig, 1999: 177. PNG (PNG).
livida Mohrig & Kauschke, 2017: 461. Australia (Qld).
longipeda Mohrig & Kauschke, 2017: 462. Australia (Qld).
longitegmentaMohrig, 1999: 167. PNG (PNG).
pilosostyla Mohrig, 1999: 176. PNG (PNG).
praecompta Mohrig, 1999: 171. PNG (PNG).
pullata Mohrig & Kauschke, 2017: 462. Australia (Qld).
subcompta Mohrig, 1999: 174. PNG (PNG).
subvera Mohrig, 2004: 143. PNG (PNG).
variovera Mohrig, 2004: 142. PNG (PNG).
vera Mohrig, 2004: 141. PNG (PNG); Australia (Qld).
zealandica Mohrig in Mohrig & Jaschhof, 1999: 27. NZ (NZ).
DODECASCIARA Edwards, 1928c: 37. Type species: Dodecasciara
debilis Edwards, 1928, mon.
debilis Edwards, 1928c: 37. Western Samoa; American Samoa, Fiji, Tonga, Samoa.
EPIDAPUS Haliday in Walker, 1851b: 7. Type species: Chionea
venatica Haliday in Walker, 1856 [= Tipula atomaria De Geer,
1778], subs. mon. Haliday in Walker, 1856b: 56.
ATOMARIA Bigot, 1854: 454. Type species: Tipula atomaria De
Geer, 1778, taut.
EPIDAPUS Haliday in Walker, 1851b: 7 (as gen.). Type species: Chionea
venatica Haliday in Walker, 1856 [= Tipula atomaria De Geer,
1778], subs. mon. Haliday in Walker, 1856b: 56.
aciculatus Vilkamaa, Hippa & Mohrig, 2014: 430. New Caledonia (New Caledonia).
chaetovenosus Mohrig in Mohrig & Jaschhof, 1999: 30. NZ (NZ).
ctenosciaroides Mohrig in Mohrig & Jaschhof, 1999: 32. NZ (NZ).
espinosalus Mohrig in Mohrig & Jaschhof, 1999: 31. NZ (NZ).
exselcus Mohrig & Kauschke, 2017: 465. Australia (Qld).
formosus Vilkamaa, Hippa & Mohrig, 2014: 433. New Caledonia (New Caledonia).
parvus Mohrig in Mohrig & Jaschhof, 1999: 35. NZ (NZ).
probus Mohrig, 2004: 150. PNG (PNG).
strenuus Mohrig, 2004: 151. PNG (PNG).
triquetrus Vilkamaa, Hippa & Mohrig, 2014: 434. New Caledonia (New Caledonia).
Undescribed sp.—Belau (Steffan, 1969: 685).
Subgenus PSEUDOAPTANOGYNA Vimmer
PSEUDOAPTANOGYNA Vimmer, 1926: 10 (Aptanogyna subg.). Type species: Aptanogyna abscondita Vimmer, 1926, mon.
angulatus Vilkamaa, Hippa & Mohrig, 2014: 431. New Caledonia (New Caledonia).
inversus Mohrig, 2004: 153. PNG (PNG).
iucundus Mohrig, 2004: 152. PNG (PNG).
Subgenus ZUHALIA Koçak & Hüseyinoglu
MACROTARSUS Mohrig, 2004: 154. Type species: Epidapus pellitus Mohrig, 2004, orig. des. [Preoccupied by Link, 1795.]
ZUHALIA Koçak & Hüseyinoglu, 2008: 2. Type species: Epidapus pellitus Mohrig, 2004, automatic.
longicubitalis Mohrig, 2004: 157. PNG (PNG).
pellitus Mohrig, 2004: 155. PNG (PNG).
primus Mohrig, 2004: 156. PNG (PNG); New Caledonia (New Caledonia).
quinquespinus Mohrig, 2004: 150. PNG (PNG).
HYPERLASION Schmitz, 1918: 95, 96. Type species: Hyperlasion
wasmanni Schmitz, 1918, mon.
alienus Mohrig, 2004: 159. PNG (PNG).
magnisenorium Hardy, 1956c: 89 (Scythropochroa). Hawaiian
Is.
politum Mohrig, 2004: 160. PNG (PNG).
Genus KEILBACHIA Mohrig & Martens
KEILBACHIA Mohrig & Martens, 1987: 483. Type species: Keilbachia nepalensis Mohrig & Martens, 1987, orig. des.
adstrictaVilkamaa et al., 2011: 54. New Caledonia (New Caledonia).
adstrictatula Mohrig & Kauschke, 2017: 467. Australia (Qld).
cracens Vilkamaa et al., 2011: 54. New Caledonia (New Caledonia).
constricta Vilkamaa et al., 2011: 57. New Caledonia (New Caledonia).
fratercula Vilkamaa et al., 2011: 60. New Caledonia (New Caledonia).
iiindigena Mohrig, 2004: 165. PNG (PNG).
truncata Vilkamaa et al., 2011: 58. New Caledonia (New Caledonia).
LOBOSCIARA Steffan, 1969: 727. Type species: Bradysia spinipennis
Sasakawa, 1962, orig. des.
trilobata Vilkamaa & Hippa, 1994: 41. Indonesia (Sulawesi); Australia (Qld).
spinipennis Sasakawa, 1962: 130 (Bradysia). Thailand; Guam,
Micronesia.
LYCORIELLA Frey, 1942: 39. Type species: Sciara vivida
Winnertz, 1867 [misid., = Sciara auripila Winnertz, 1867], orig.
des.
agraria Felt, 1898a: 225 [1898b: 225] (Sciara). USA; Australia (NSW, Vic) [immigrant], NZ (NZ)
[immigrant]; England.
agarici Loudon, 1978: 163. Australia (NSW).
hoyti Hardy, 1956c: 80 (Sciara). Hawaiian Is.
ingenua Dufour, 1839: 29 (Sciara). [Not given; = France]; Australia (NSW, Tas) [immigrant], Hawaiian Is [immigrant], Antarctica, Kerguelen Is, Macquarie I; Holarctic.
mali Fitch, 1856: 486 (Molobrus). USA; Hawaiian Is.
pauciseta Felt, 1898: 224. USA.
womersleyi Séguy, 1940: 210. Kerguelen Is.
sativae Johannsen, 1912: 133 (Sciara). USA; Australia (ACT, NSW, Wld, SA, Tas, Vic), Hawaiian Is, Kerguelen Is, Norfolk I; Holarctic.
auberti Séguy, 1940: 211. Kerguelen Is.
solispina Hardy, 1956c: 84 (Sciara). Hawaiian Is.
NEOPHNYXIA Tonnoir in Tonnoir & Edwards, 1927: 800.
Type species: Neophnyxia nelsoniana Tonnoir in Tonnoir &
Edwards, 1927, orig. des.
nelsoniana Tonnoir in Tonnoir & Edwards, 1927: 800. NZ
(NZ).
ODONTONYX Rübsaamen, 1894: 19, 25. Type species: Sciara
nigra Wiedemann, 1821, des. Coquillett, 1910: 578. [Preocc. Stephens,
1827.]
ODONTOSCIARA Rübsaamen, 1908: 450 (n. n. for Odontonyx).
PHORODONTA Coquillett, 1910: 589 (n. n. for Odontonyx).
filipes Walker, 1865a: 102 (Sciara). "New Guinea"
[= Indonesia (Irian Jaya)].
pacifica Edwards, 1924f: 569 (Phorodonta). Fiji; American
Samoa, Vanuatu, Western Samoa.
politicollis Edwards, 1928c: 34 (Phorodonta). Western Samoa.
Genus PELLICIPLANTA Mohrig
PELLICIPLANTA Mohrig, 2004: 168. Type species: Pelliciplanta unica Mohrig, 2004, orig. des.
unica Mohrig, 2004: 169. PNG (PNG).
Genus PHYTOSCIARA Frey
PHYTOSCIARA Frey, 1942: 27. Type species: Sciara halterata Lengersdorf, 1926, orig. des.
Subgenus DOLICHOSCIARATuomikoski
DOLICHOSCIARA Tuomikoski, 1960: 107. Type species: Sciara flavipes Meigen, 1804, orig. des.
atrata Mohrig, 1999: 195. PNG (PNG).
bella Mohrig, 1999: 185. PNG (PNG); Australia Qld).
bellaformis Mohrig, 1999: 186. PNG (PNG).
chaetocoxa Mohrig, 1999: 191. PNG (PNG).
conturbata Mohrig, 1999: 183. PNG (PNG); Australia (Qld).
filispinosa Mohrig, 1999: 188. PNG (PNG).
fumida Mohrig, 1999: 190. PNG (PNG).
gilvifucata Mohrig, 1999: 193. PNG (PNG).
pseudoornata Mohrig, 1999: 187. PNG (PNG); Australia (Qld).
ruficoxa Brunetti, 1912: 146 (Sciara). India; PNG (PNG); Indonesia (Sumatra), Malaysia (Sabah).
subfumida Mohrig, 1999: 192. PNG (PNG).
Subgenus PHYTOSCIARA Frey
PHYTOSCIARA Frey, 1942: 27 (as gen.). Type species: Sciara halterata
Lengersdorf, 1926, orig. des.
brachygaster Steffan, 1969: 709. Belau; Micronesia.
maculosa Mohrig, 1999: 182. PNG (PNG).
vulcanata Steffan, 1973: 358. Hawaiian Is.
PSEUDOSCIARA Kieffer, 1898a: 194. Type species: Pseudosciara
pictiventris Kieffer, 1898, orig. des. [Preocc. Schiner, 1866.]
PLASTOSCIARA Berg, 1899: 78 (n. n. for Pseudosciara).
COSMOSCIARA Frey, 1942: 39. Type species: Plastosciara perniciosa
Edwards, 1922, orig. des.
adrostylata Hardy, 1956c: 72. Hawaiian Is.
flavibasis Edwards, 1928c: 35. American Samoa; Fiji.
jaluitensis Steffan, 1969: 688. Marshall Is.
latipons Hardy, 1956c: 77. Hawaiian Is; Marshall Is, Micronesia.
longicosta Hardy, 1956c: 75. Hawaiian Is.
yapensis Steffan, 1969: 692. Micronesia.
PNYXIA Johannsen, 1912: 114. Type species: Epidapus scabiei
Hopkins, 1895, orig. des.
Undescribed sp.--Hawaiian Is (Steffan, 1981: 140).
Genus PSEUDOLYCORIELLA Menzel & Mohrig
PSEUDOLYCORIELLA Menzel & Mohrig, 1998: 367. Type species: Sciara bruckii Winnertz, 1867, orig. des.
acicula Vilkamaa et al., 2012: 3. New Caledonia (New Caledonia).
attrita Mohrig, 2014: 144. PNG (PNG).
bispina Mohrig in Mohrig & Jaschhof, 1999: 40. NZ (NZ).
bisulca Vilkamaa et al., 2012: 3. New Caledonia (New Caledonia).
bitorquia Mohrig, 2014: 162. PNG (PNG).
breviseta Mohrig in Mohrig & Jaschhof, 1999: 41. NZ (NZ).
cavatiostyla Mohrig, 2014: 152. PNG (PNG).
cavicata Skuse, 1888b: 688 (Sciara). Australia (NSW).
familiaris Skuse, 1888b: 687 (Sciara). Australia (NSW).
festiva Skuse, 1888b: 689 (Sciara). Australia (NSW).
capillosa Vilkamaa et al., 2012: 6. New Caledonia (New Caledonia).
commoda Mohrig, 2014: 149. PNG (PNG).
densispina Mohrig, 2014: 140. PNG (PNG).
espinosa Mohrig, 2014: 146. PNG (PNG).
espinula Mohrig, 2014: 145. PNG (PNG).
forunata Mohrig, 2014: 151. PNG (PNG).
frederickedwardsi Köhler in Köhler & Mohrig, 2016: 11. NZ (NZ).
ignobilis Skuse, 1888b: 717 (Sciara). Australia (NSW).
jejuna Edwards in Tonnoir & Edwards, 1927: 796 (Sciara). NZ (NZ).
latiflagellata Vilkamaa et al., 2012: 8. New Caledonia (New Caledonia).
longicostalis Mohrig, 2014: 139. PNG (PNG).
macrotegmenta Mohrig in Mohrig & Jaschhof, 1999: 41. NZ (NZ).
macrotrichata Mohrig, 2014: 157. PNG (PNG).
microphalli Mohrig, 2014: 161. PNG (PNG).
microtrichata Mohrig, 2014: 159. PNG (PNG).
pallidula Vilkamaa et al., 2012: 9. New Caledonia (New Caledonia).
paucispina Mohrig, 2014: 142. PNG (PNG).
perspicua Mohrig, 2014: 163. PNG (PNG).
praecipua Mohrig, 2014: 155. PNG (PNG).
quadrispinosa Mohrig, 2014: 154. PNG (PNG).
senticosa Vilkamaa et al., 2012: 10. New Caledonia (New Caledonia).
setigera Hardy, 1960: 234 (Spathobdella). Hawaiian Is.
simplex Vilkamaa et al., 2012: 10. New Caledonia (New Caledonia).
sinoupalpa Mohrig, 2014: 143. PNG (PNG).
skusei Mohrig, Kauschke & Broadley, 2016: 141. Norfolk I; Australia (Qld).
snellingi Mohrig, 2014: 156. PNG (PNG).
spicata Vilkamaa et al., 2012: 12. New Caledonia (New Caledonia).
tenebriocoxa Mohrig, 2014: 159. PNG (PNG).
tenebriolata Mohrig, 2014: 137. PNG (PNG).
tenuis Vilkamaa et al., 2012: 14. New Caledonia (New Caledonia).
tonnoiri Köhler in Köhler & Mohrig, 2016: 14. NZ (NZ).
tribulosa Vilkamaa et al., 2012: 15. New Caledonia (New Caledonia).
tricanthula Mohrig, 2014: 147. PNG (PNG).
trispicata Vilkamaa et al., 2012: 16. New Caledonia (New Caledonia).
tumidior Vilkamaa et al., 2012: 17. New Caledonia (New Caledonia).
vestita Mohrig, 2014: 150. PNG (PNG).
villosa Vilkamaa et al., 2012: 20. New Caledonia (New Caledonia).
zealandica Edwards in Tonnoir & Edwards, 1927: 796 (Sciara). NZ
(NZ). [Preocc. Kieffer, 1910; no n. n. proposed at this time.]
Genus PSEUDOZYGOMMA Mohrig
PSEUDOZYGOMMA Mohrig, 2004: 144. Type species: Pseudozygomma contraria Mohrig, 2004, orig. des.
contraria Mohrig, 2004: 145. PNG (PNG).
flavoabdominalis Mohrig, 2004: 147. PNG (PNG).
maculipennis Skuse, 1890c: 411 (Zygoneura). Australia (NSW).
nigroalata Mohrig, 2004: 148. PNG (PNG).
nigrocorporea Mohrig, 2004: 146. PNG (PNG).
PSEUDOZYGONEURA Steffan, 1969: 676. Type species: Pseudozygoneura
musicola Steffan, 1969, orig. des.
musicola Steffan, 1969: 677. Belau; Fiji, Hawaiian Is [introduced; ?established],
Micronesia, Solomon Is.
SCAPTOPSCIARA Edwards in Tonnoir & Edwards, 1927: 798
(Sciara subg.). Type species: Sciara quinquelineata Macquart,
1834 [= Sciara vitripennis Meigen, 1818], orig. des.
SCAPTOPSCIARA Edwards in Tonnoir & Edwards, 1927: 798
(Sciara subg.). Type species: Sciara quinquelineata Macquart,
1834 [= Sciara vitripennis Meigen, 1818], orig. des.
UDMANNIA Frey, 1942: 38. Type species: Sciara pusilla Meigen,
1818, orig. des. [Preocc. Bergroth, 1915.]
UDMANNIELLA Frey, 1948: 86 (n. n. for Udmannia).
crassivena Mohrig, 2004: 167. PNG (PNG).
nigrita Hardy, 1956c: 86 (Scatopsciara). Hawaiian Is.
spiculata Vilkamaa et al., 2012: 69. New Caledonia (New Caledonia).
tenuistyla Vilkamaa et al., 2012: 71. New Caledonia (New Caledonia).
unicalcarata Edwards in Tonnoir & Edwards, 1927: 798 (Sciara).
NZ (NZ).
validovenosa Mohrig & Kauschke, 2017: 477. Australia (Qld).
vicina Vilkamaa et al., 2012: 72. New Caledonia (New Caledonia).
Subgenus XENOPYGINA Frey
XENOPYGINA Frey, 1948: 51 (Bradysia subg.). Type species: Bradysia paradoxa Frey, 1948, orig. des.
ampliocerosa Mohrig, 2004: 167. PNG (PNG).
brevicolla Mohrig & Kauschke, 2017: 473. Australia (Qld).
exigua Vilkamaa et al., 2012: 68. New Caledonia (New Caledonia).
monocerosa Mohrig, 2004: 166. PNG (PNG).
Unplaced to Subgenus of SCAPTOSCIARA
dubiosa Mohrig & Kauschke, 2017: 475. Australia (Qld).
SCHWENKFELDINA Frey, 1942: 32 (Neosciara subg.). Type species:
Sciara carbonaria Meigen, 1820, orig. des.
inopinata Mohrig, 1999: 157. PNG (PNG).
ponderosa Walker, 1864: 230 (Sciara). Indonesia (Maluku).
regia Mohrig, 1999: 156. PNG (PNG).
LYCORIA Meigen, 1800: 17. Suppressed by I.C.Z.N., 1963b: 339.
SCIARA Meigen, 1803: 263. Type species: Tipula thomae Linnaeus,
1767, mon.
adjuncta Skuse, 1890c: 398. Australia (NSW).
aemula Skuse, 1888b: 678. Australia (NSW).
approximata Skuse, 1888b: 697. Australia (NSW).
aquila Skuse, 1888b: 692. Australia (NSW).
atratula Skuse, 1888b: 704. Australia (NSW).
audax Skuse, 1888b: 693. Australia (NSW).
brevifurca Skuse, 1888b: 700. Australia (NSW).
consanguinea Skuse, 1890c: 401. Australia (NSW).
contermina Skuse, 1890c: 395. Australia (NSW).
distigma Edwards, 1924f: 568. Fiji.
diversa Skuse, 1888b: 701. Australia (NSW).
dolosa Skuse, 1888b: 683. Australia (NSW).
erratica Skuse, 1888b: 696. Australia (NSW).
evanescens Skuse, 1888b: 698. Australia (NSW).
exposita Skuse, 1890c: 397. Australia (NSW).
femoralis Doleschall, 1858: 78. Indonesia (Maluku).
festina Skuse, 1888b: 684. Australia (NSW).
finitima Skuse, 1888b: 677. Australia (NSW).
flavicoxis Skuse, 1890c: 396. Australia (NSW).
fumipennis Skuse, 1888b: 706. Australia (NSW).
helmsi Skuse, 1890c: 408. Australia (NSW).
infixa Skuse, 1890c: 400. Australia (NSW).
lucidipennis Skuse, 1888b: 714. Australia (NSW).
luculenta Skuse, 1888b: 705. Australia (NSW).
marcilla Hutton, 1902b: 192. NZ (NZ).
marginata Skuse, 1890c: 399. Australia (NSW).
neorufescens Miller, 1950: 57 (n. n. for rufescens Hutton).
NZ (NZ).
rufescens Hutton, 1881: 13. NZ (NZ). [Preocc. Zetterstedt, 1852.]
nubicula Skuse, 1888b: 715. Australia (NSW).
palliceps Edwards, 1928: 25. Malaysia (Pen); PNG (PNG); Indonesia (Sumatra), Philippines, Thailand.
philpotti Tonnoir in Tonnoir & Edwards, 1927: 796. NZ
(NZ).
praescellens Skuse, 1890c: 404. Australia (NSW).
prominens Hardy, 1956c: 83. Hawaiian Is.
promiscua Skuse, 1890c: 394. Australia (NSW).
reciproca Walker, 1856d: 420. Australia (Tas).
recondita Skuse, 1890c: 390. Australia (NSW).
scitula Skuse, 1888b: 699. Australia (NSW).
sedula Skuse, 1888b: 674. Australia (NSW).
selecta Walker, 1858b: 77. Indonesia (Maluku).
sororia Skuse, 1888b: 716. Australia (NSW).
tepperi Skuse, 1890c: 393. Australia (SA).
tryoni Skuse, 1890c: 391. Australia (Qld).
insulana Vilkamaa, Hippa & Mohrig, 2015: 590. New Caledonia (New Caledonia).
turgidula Vilkamaa, Hippa & Mohrig, 2015: 592. New Caledonia (New Caledonia).
turrida de Meijere, 1913a: 45. Indonesia (Maluku).
varipes Walker, 1865a: 102. "New Guinea" [= Indonesia (Irian
Jaya)].
vecors Skuse, 1888b: 695. Australia (NSW).
zealandica Kieffer, 1910a: 327. NZ (NZ).
SCYTHROPOCHROA Enderlein, 1911c: 138. Type species: Scythropochroa
latefurcata Enderlein, 1911, orig. des.
antarctica Hudson, 1892: pl. 4, fig. 5a,b (Mycetophila). Not
given [NZ (NZ)]. Nomen dubium.
atrichata Mohrig, 2004: 134. PNG (PNG).
atrichoalata Mohrig, 2004: 135. PNG (PNG).
gressitti Steffan, 1969: 680. Micronesia; Guam.
macleayi Skuse, 1888b: 691 (Sciara). Australia (NSW).
micropalpa Mohrig, 2004: 137. PNG (PNG).
nitida Edwards in Tonnoir & Edwards, 1927: 799. NZ (NZ).
parapectinea Mohrig, 2004: 138. PNG (PNG).
paucitrichata Mohrig, 2004: 132. PNG (PNG).
quadrispinosa Steffan, 1969: 682. Belau.
samoana Edwards, 1928c: 36. Western Samoa.
semitrichata Mohrig, 2004: 131. PNG (PNG).
trichoalata Mohrig, 2004: 130. PNG (PNG).
trichovenosa Mohrig, 2004: 133. PNG (PNG).
trispinosa Steffan, 1969: 682. Marshall Is.
VULAGISCIARA Evenhuis in Evenhuis et al., 2007: 4. Type species: Vulagisciara myrmecophila Evenhuis, 2007, orig. des.
myrmecophila Evenhuis in Evenhuis et al., 2007: 4.
Fiji.
PLASTOSCIARA Frey, 1942: 34. Type species: Sciara lignicola
Winnertz, 1867, orig. des. [Preocc. Berg, 1899.]
XYLOSCIARA Tuomikoski, 1957: 10 (n. n. for Plastosciara).
brevipes Steffan, 1964: 293. NZ (Campbell I).
ZYGONEURA Meigen, 1830: 304. Type species: Zygoneura sciarina
Meigen, 1830, mon.
contractans Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ
(NZ).
remota de Meijere, 1913b: 305. Indonesia (Irian Jaya).