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Australasian/Oceanian Diptera Catalog -- Web Version

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11. Family SCIARIDAE

[Originally] by Wallace A. Steffan

Sciaridae are common faunal elements in nearly all regions of the world but are poorly known because their small size and superficial homogeneity have not attracted collectors or taxonomists. Immature stages can be found in rotten wood, under bark of fallen trees, in other kinds of decaying plant material, feeding in fungi or animal excrement or, in Austrosciara termitophila, as inquilines in termite nests. One genus (Vulagisciara) and new species has been found as a guest in rubiaceous ant-plants in Fiji (Evenhuis et al., 2007.). Some species feed on living plant tissue and can cause economically important damage, especially in greenhouses and mushroom houses.

Recent studies indicate that the Australasian/Oceanian fauna is rich in taxa. The regional literature, with the exception of the faunal studies of Hawaii (Hardy 1960) and Micronesia (Steffan 1969), consists primarily of descriptions of new species. Skuse (1888b, 1890d) described 61 Sciaridae from Australia, but most of the collections were from New South Wales only. The Papua New Guinean fauna has been studied by Mohrig (1999, 2004). The New Zealand fauna was treated by Tonnoir & Edwards (1927) and most recently revised by Mohrig & Jaschhof (1999), and thirteen new species of Bradysia were described by Vilkamaa et al. (2012) from New Caledonia.

This catalog is essentially a list of published names, although some necessarily obvious taxonomic changes have been made. Many older generic placements undoubtedly are incorrect, and types need to be examined before correct generic assignments can be made. Most of the species listed under Sciara should be transferred to other genera after examination of the types.
Refs.: Hardy (1960, Hawaiian spp.), Freeman (1983), Frey (1942, 1948), Steffan (1966b, rev. Nearctic gen.; 1969, Micronesian spp.), Tuomikoski (1960), Evenhuis (2007 et al., Fiji sciarids), Vilkamaa et al. (2012, New Caledonian Bradysia).

Genus AERUMNOSA Mohrig

AERUMNOSA Mohrig, 1999: 158. Type species: Aerumnosa noctinigra Mohrig, 1999, orig. des.
furcillata Mohrig, 1999: 163. PNG (PNG).
noctinigra Mohrig, 1999: 159. PNG (PNG).
nocturna Mohrig, 1999: 161. PNG (PNG).
perfida Mohrig, 1999: 160. PNG (PNG).
rhinocerata Mohrig, 1999: 165. PNG (PNG).
trichovenosa Mohrig, 1999: 164. PNG (PNG).

Genus AUSTROSCIARA Schmitz & Mjöberg

AUSTROSCIARA Schmitz & Mjöberg, 1924: 1. Type species: Austrosciara termitophila Schmitz & Mjöberg, 1924, mon.
CTENOSCIARA Tuomikoski, 1960: 110. Type species: Sciara hyalipennis Meigen, 1804, orig. des.
abdita Mohrig, 2014: 134 (Ctenosciara). PNG (PNG).
certa Mohrig, 2014: 124 (Ctenosciara). PNG (PNG).
constrictans Edwards in Tonnoir & Edwards, 1927: 792 (Sciara). NZ (NZ).
     nubeculosa Edwards in Tonnoir & Edwards, 1927: 793 (Sciara). NZ (NZ).
cracens
Vilkamaa et al., 2012: 38 (Ctenosciara). New Caledonia (New Caledonia).
crinita
Vilkamaa et al., 2012: 41 (Ctenosciara). New Caledonia (New Caledonia).
depilis Vilkamaa et al., 2012: 41 (Ctenosciara). New Caledonia (New Caledonia).
depressa Vilkamaa et al., 2012: 44 (Ctenosciara). New Caledonia (New Caledonia).
etorutao Köhler in Köhler & Mohrig, 2016: 9 (Ctenosciara). NZ (NZ).
exilis
Vilkamaa et al., 2012: 45 (Ctenosciara). New Caledonia (New Caledonia).
griseinervis Edwards in Tonnoir & Edwards, 1927: 793 (Sciara). NZ (NZ).
hawaiiensis Hardy, 1956c: 78 (Sciara). Hawaiian Is.
hyalipennis Meigen, 1804: 99 (Sciara). Not given [Central Europe; see Gerbachevskaya-Pavluchenko, 1986: 43]; NZ (Campbell I, NZ) [immigrant]; Taiwan, widesp. Palearctic Reg.
     annulata Meigen, 1818: 284 (Sciara). Not given [Germany].
inferma Mohrig, 2014: 130 (Ctenosciara). PNG (PNG).
inflata
Vilkamaa et al., 2012: 45 (Ctenosciara). New Caledonia (New Caledonia).
infrequens Skuse, 1888b: 719 (Sciara). Australia (NSW).
lobigera Vilkamaa et al., 2012: 47 (Ctenosciara). New Caledonia (New Caledonia).
lubrica
Mohrig, 2014: 133 (Ctenosciara). PNG (PNG).
montivaga
Skuse, 1888b: 710 (Sciara). Australia (NSW).
multiformis Mohrig, 2014: 129 (Ctenosciara). PNG (PNG).
multispinosa
Steffan, 1969: 712 (Ctenosciara). Micronesia.
nigrostyla Mohrig in Mohrig & Jaschhof, 1999: 1 (Ctenosciara)7. NZ (NZ).
nudopteryx Mohrig in Mohrig & Jaschhof, 1999: 19 (Ctenosciara). NZ (NZ).
obesa Vilkamaa et al., 2012: 49. New Caledonia (New Caledonia).
ovalis
Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ (NZ).
pedibusa Mohrig, Kauscke & Keller, 2017: 362. Australia (Vic).
pullata Mohrig, 2014: 127 (Ctenosciara). PNG (PNG).
semipilosa
Mohrig, 2014: 131 (Ctenosciara). PNG (PNG).
spectabilis
Skuse, 1888b: 716 (Sciara). Australia (NSW).
subabdita Mohrig, 2014: 136 (Ctenosciara). PNG (PNG).
subpullata
Mohrig, 2014: 128 (Ctenosciara). PNG (PNG).
termitophila
Schmitz & Mjöberg, 1924: 2. Australia (NSW).
uniocolorata Mohrig, 2014: 126 (Ctenosciara). PNG (PNG).
winnertzi
Skuse, 1888b: 709 (Sciara). Australia (NSW); NZ(NZ); China.
     rufulenta Edwards in Tonnoir & Edwards, 1927: 794 (Sciara). NZ (NZ).
     vicarians Edwards in Tonnoir & Edwards, 1927: 793 (Sciara). NZ (NZ).
xanthonota
Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ (NZ).

Genus BRADYSIA Winnertz

BRADYSIA Winnertz, 1867: 180. Type species: Bradysia angustipennis Winnertz, 1867, des. Enderlein, 1911c: 127.
aequispina Vilkamaa et al., 2012: 26. New Caledonia (New Caledonia).
amabilis Skuse, 1888b: 712 (Sciara). Australia (NSW).
angustopalpa Mohrig, 2016: 29. PNG (PNG).
aspera
Mohrig, 2016: 6. PNG (PNG).
aspercerca
Mohrig, 2016: 5. PNG (PNG).
beata Mohrig, 2016: 25. PNG (PNG).
bishopi Steffan, 1973: 353. Hawaiian Is; American Samoa, Fiji, French Polynesia, New Caledonia (New Caledonia); Puerto Rico, Seychelles, Thailand.
boninensis Steffan, 1969: 717. Bonin Is.
bullata Mohrig, 2016: 21. PNG (PNG).
campbellensis Steffan, 1964: 296. NZ (Campbell I).
centidens Vilkamaa et al., 2012: 28. New Caledonia (New Caledonia).
circumfilata Mohrig, 2016: 26. PNG (PNG).
clavistylis Vilkamaa et al., 2012: 28. New Caledonia (New Caledonia).
conjuncta
Skuse, 1890c: 403 (Sciara). Australia (NSW).
     serenipennis Skuse, 1890c: 402 (Sciara). Australia (NSW).
crassicornis
Skuse, 1890c: 407 (Sciara). Australia (NSW).
credula Mohrig, 2016: 29. PNG (PNG).
cristata
Mohrig, 2016: 17. PNG (PNG).
exsequialis
Skuse, 1890c: 409 (Sciara). Australia (NSW).
fantula Mohrig, 2016: 11. PNG (PNG).
filigera
Vilkamaa et al., 2012: 30. New Caledonia (New Caledonia).
flaveola
Mohrig, 2016: 14. PNG (PNG).
flavocristata
Mohrig, 2016: 15. PNG (PNG).
frequens Skuse, 1888b: 682 (Sciara). Australia (NSW).
froggatti Skuse, 1888b: 681 (Sciara). Australia (NSW).
gemellata Vilkamaa et al., 2012: 32. New Caledonia (New Caledonia).
gibbosa
Vilkamaa et al., 2012: 33. New Caledonia (New Caledonia); PNG (PNG).
impatiens Johannsen, 1912: 136 (Sciara). USA; Austrralia (ACT, NSW, Qld, SA, Tas, Vic, WA); Hawaiian I, Norfolk I; Brazil, China, Japan.
     hardyi Shaw, 1952: 493 (Sciara). Hawaiian Is.
inaequispina Mohrig, 2016: 10. PNG (PNG).
incohata Mohrig, 2016: 6. PNG (PNG).
irwini
Köhler & Menzel, 2013: 66. New Caledonia (New Caledonia).
kraussi
Steffan, 1969: 718. Guam.
luctifica Skuse, 1888b: 680 (Sciara). Australia (SA); New Caledonia (New Caledonia), PNG (PNG)..
     planistylata
Vilkamaa et al., 2012: 38. New Caledonia (New Caledonia).
maesta
Skuse, 1888b: 691 (Sciara). Australia (NSW).
mastersi Skuse, 1888b: 723 (Trichosia). Australia (NSW).
megahypopygialis Mohrig, 2016: 23. PNG (PNG).
melina Vilkamaa et al., 2012: 34. New Caledonia (New Caledonia).
microdentata Mohrig, 2016: 24. PNG (PNG).
molokaiensis Grimshaw, 1901: 2 (Sciara). Hawaiian Is.
     stonei Shaw, 1952: 495 (Sciara). Hawaiian Is.
mutuata Mohrig, 2016: 12. PNG (PNG).
novaeseelandiae
Köhler & Mohrig, 2016: 2. NZ (NZ).
nudilobata Vilkamaa et al., 2012: 36. New Caledonia (New Caledonia).
obnoxia Mohrig, 2016: 11. PNG (PNG).
ocellaris
Comstock, 1882: 202 (Sciara). USA; Australia (ACT, NSW, Qld, SA, Tas, Vic, WA), Hawaiian Is, Micronesia; cosmopol.
     tritici Coquillett, 1895a: 408 (Sciara). Not given [USA.
     johannseni Shaw, 1952: 493 (Sciara). Hawaiian Is. [Preocc. Enderlein, 1912.]
     garreti Shaw, 1952: 494 (Sciara). Hawaiian Is.
     laffooni Shaw, 1952: 494 (Sciara). Hawaiian Is.
ornatula Skuse, 1888b: 711 (Sciara). Australia (NSW).
pallipes Fabricius, 1787: 326 (Tipula). Germany; NZ (NZ) [immigrant]; Europe, Algeria.
paudicens Vilkamaa et al., 2012: 37. New Caledonia (New Caledonia).
pellistyla
Mohrig, 2016: 4. PNG (PNG).
perfrigida
Mohrig, 2016: 9. PNG (PNG).
pernitida Skuse, 1888b: 685 (Sciara). Australia (NSW).
pictipes Skuse, 1888b: 721 (Sciara). Australia (NSW).
     notata
Skuse, 1888b: 720 (Sciara). Australia (NSW).
     seticornis
Vilkamaa et al., 2012: 39. New Caledonia (New Caledonia).
praebullata Mohrig, 2016: 19. PNG (PNG).
pseudocampestris Mohrig--not Australasian/Oceanian.
quinquespina
Mohrig, 2016: 7. PNG (PNG).
radicum
Brunetti, 1912b: 139 (Sciara). India; American Samoa, Fiji, French Polynesia (Marquesas, ?Society Is), Hawaiian Is, Western Samoa; England.
robusta Mohrig, 2016: 27. PNG (PNG).
rubra Harrison, 1955: 213 (Sciara). NZ (Campbell I).
saetibullata Mohrig, 2016: 21. PNG (PNG).
semicristata
Mohrig, 2016: 18. PNG (PNG).
similigibbosa
Köhler & Menzel, 2013: 64. New Caledonia (New Caledonia).
snyderi Steffan, 1969: 723. Bonin Is.
spatitergum Hardy, 1956c: 85 (Sciara). Hawaiian Is; PNG (PNG); Brazil, India, Madagascar, Panama, Zimbabwe.
     radicum, Hardy, 1960, not Brunetti, 1912, misid.
spicotegmata Mohrig, 2016: 28. PNG (PNG).
spissa Vilkamaa et al., 2012: 41. New Caledonia (New Caledonia).
stricta Vilkamaa et al., 2012: 42. New Caledonia (New Caledonia).
subbullata Mohrig, 2016: 20. PNG (PNG).
subcristata
Mohrig, 2016: 18. PNG (PNG).
suspecta
Mohrig, 2016: 8. PNG (PNG).
tenuitegmata
Mohrig, 2016: 15. PNG (PNG).
trispinea
Mohrig, 2016: 8. PNG (PNG).
unica
Skuse, 1888b: 708 (Sciara). Australia (NSW).
validospina Mohrig, 2016: 13. PNG (PNG).
vevnusta
Mohrig, 2016: 22. PNG (PNG).
violenta
Mohrig, 2016: 16. PNG (PNG).
watsoni
Colless, 1962b: 955. Macquarie I.


Genus CHAETOSCIARA Frey

CHAETOSCIARA Frey, 1942:33 (as Neosciara Pettey subg.). Type species: Sciara fenestralis Zetterstedt, 1838 [misidentification, = Sciara estlandica Lengersdorf, 1929], by orig. des.
NEOSCIARA, authors, not Pettey, 1918, misid.
ingrata Mohrig, 1999: 199. PNG (PNG).
recondita Mohrig & Kausckhke, 2017: 453, Australia (Qld).
recta Mohrig, 1999: 198. PNG (PNG).

Genus CHIASMATA Mohrig

CHIASMATA Mohrig, 2016: 30. Type species: Chiasmata duubioproterva Mohrig, 2016, orig. des.
dubioproterva Mohrig, 2016: 30. PNG (PNG).



Genus CORYNOPTERA Winnertz

CORYNOPTERA Winnertz, 1867: 177. Type species: Corynoptera perpusilla Winnertz, 1867, des. Enderlein, 1911c: 127.
alpina Mohrig—not Australasian/Oceanian.
aggregata Köhler & Mohrig, 2016: 4. NZ (NZ).
ancylospina Mohrig in Mohrig & Jaschhof, 1999: 77. NZ (NZ).
basisetosa Mohrig in Mohrig & Jaschhof, 1999: 53. NZ (NZ).
brevipalpis
Steffan, 1969: 695. Belau; Hawaiian Is.
catrinjaschhofae Köhler & Mohrig, 2016: 7. NZ (NZ).
coronospina Mohrig in Mohrig & Jaschhof, 1999: 84. NZ (NZ).
cowanorum Mohrig in Mohrig & Jaschhof, 1999: 81. NZ (NZ).
denospica Mohrig in Mohrig & Jaschhof, 1999: 74. NZ (NZ).
densisetosa Mohrig in Mohrig & Jaschhof, 1999: 54. NZ (NZ).
didymistyla Mohrig in Mohrig & Jaschhof, 1999: 67. NZ (NZ).
diversicalcaria Mohrig, 2004: 163. PNG (PNG).
dividospica Mohrig in Mohrig & Jaschhof, 1999: 70. NZ (NZ).
expressospina Mohrig in Mohrig & Jaschhof, 1999: 79. NZ (NZ).
facticia Mohrig in Mohrig & Jaschhof, 1999: 49. NZ (NZ).
fatigans
Johannsen, 1912: 132 (Sciara). USA; New Caledonia (New Caledonia), NZ (NZ) [immigrant]; Holarctic.
filisetosa Mohrig in Mohrig & Jaschhof, 1999: 63. NZ (NZ).
filispica Mohrig in Mohrig & Jaschhof, 1999: 75. NZ (NZ).
fuscispica Mohrig in Mohrig & Jaschhof, 1999: 71. NZ (NZ).
gladiota Mohrig, 2004: 162. PNG (PNG).
harrisi
Edwards in Tonnoir & Edwards, 1927: 797 (Sciara). NZ (NZ).
hemisetosa Mohrig in Mohrig & Jaschhof, 1999: 64. NZ (NZ).
heterochela
Steffan, 1969: 697. Micronesia; Belau.
latistylata Hardy, 1956c: 82 (Sciara). Hawaiian Is; Marshall Is, Northern Marianas.
microsetosa Mohrig in Mohrig & Jaschhof, 1999: 62. NZ (NZ).
minutela Skuse, 1888b: 702 (Sciara). Australia (NSW).
nigrospina
Mohrig in Mohrig & Jaschhof, 1999: 45. NZ (NZ).
nigrotegminis Mohrig in Mohrig & Jaschhof, 1999: 56. NZ (NZ).
oririclausa Mohrig in Mohrig & Jaschhof, 1999: 65. NZ (NZ).
parasetosa Mohrig in Mohrig & Jaschhof, 1999: 58. NZ (NZ).
pentaspina Mohrig in Mohrig & Jaschhof, 1999: 47. NZ (NZ).
plasiosetosa Mohrig in Mohrig & Jaschhof, 1999: 59. NZ (NZ).
ponapensis
Steffan, 1969: 703. Micronesia.
praegladiota Mohrig, 2004: 162. PNG (PNG).
prinospina Mohrig in Mohrig & Jaschhof, 1999: 80. NZ (NZ).
priscospina Mohrig in Mohrig & Jaschhof, 1999: 86. NZ (NZ).
pronospica Mohrig in Mohrig & Jaschhof, 1999: 72. NZ (NZ).
propriospina Mohrig in Mohrig & Jaschhof, 1999: 87. NZ (NZ).
prosopspina Mohrig in Mohrig & Jaschhof, 1999: 82. NZ (NZ).
psilospina Mohrig in Mohrig & Jaschhof, 1999: 83. NZ (NZ).
quasisetosa Mohrig in Mohrig & Jaschhof, 1999: 57. NZ (NZ).
sabroskyi
Steffan, 1969: 703. Belau.
semiaggregata Mohrig in Mohrig & Jaschhof, 1999: 61. NZ (NZ).
subantarctica
Steffan, 1964: 295. NZ (Campbell I).
sylviae Steffan, 1969: 705. Belau; Micronesia.
tapleyi Edwards in Tonnoir & Edwards, 1927: 797 (Sciara). NZ (NZ).
variospina Mohrig in Mohrig & Jaschhof, 1999: 50. NZ (NZ).

Genus COSMOSCIARA Frey

COSMOSCIARA Frey, 1942: 39. Type species: Plastosciara perniciosa Edwards, 1922 [= Sciara hartii Johannsen, 1912], orig. des.
hartii Johannsen, 1912: 144 (Sciara). USA; Australia (WA), Fiji, French Polynesia (Marquesas), Hawaiian Is, Western Samoa; cosmopol..
     perniciosa Edwards, 1922c: 160 (Plastosciara). England.
     brevicalcarata Hardy, 1956c: 73 (Plastosciara). Hawaiian Is.

Genus CRATYNA Winnerz

CRATYNA Winnertz, 1867: 167. Type species: Cratyna atra Winnertz, 1867, orig. des.
adulterina Mohrig & Kauschke, 2017: 456. Australia ( Qld).
compta Mohrig, 1999: 173. PNG (PNG).
consensa Mohrig, 1999: 172. PNG (PNG).
consentanea Mohrig, 1999: 169. PNG (PNG).
distorta Mohrig, 1999: 168. PNG (PNG).
ficta Mohrig, 1999: 159. PNG (PNG).
flagria Mohrig, 1999: 179. PNG (PNG); Australia (Qld).
flagriola Mohrig & Kauschke, 2017: 457. Australia (Qld).
flagriantennata Mohrig, 1999: 178. PNG (PNG).
flagritissima Mohrig, 1999: 180. PNG (PNG).
flavivaria Mohrig, 1999: 170. PNG (PNG).
flavothoracica Mohrig & Kauschke, 2017: 460. Australia (Qld).
interflagria Mohrig, 1999: 177. PNG (PNG).
livida Mohrig & Kauschke, 2017: 461. Australia (Qld).
longipeda Mohrig & Kauschke, 2017: 462. Australia (Qld).
longitegmentaMohrig, 1999: 167. PNG (PNG).
pilosostyla Mohrig, 1999: 176. PNG (PNG).
praecompta Mohrig, 1999: 171. PNG (PNG).
pullata Mohrig & Kauschke, 2017: 462. Australia (Qld).
subcompta Mohrig, 1999: 174. PNG (PNG).
subvera Mohrig, 2004: 143. PNG (PNG).
variovera Mohrig, 2004: 142. PNG (PNG).
vera Mohrig, 2004: 141. PNG (PNG); Australia (Qld).
zealandica Mohrig in Mohrig & Jaschhof, 1999: 27. NZ (NZ).

Genus DODECASCIARA Edwards

DODECASCIARA Edwards, 1928c: 37. Type species: Dodecasciara debilis Edwards, 1928, mon.
debilis Edwards, 1928c: 37. Western Samoa; American Samoa, Fiji, Tonga, Samoa.


Genus EPIDAPUS Haliday

EPIDAPUS Haliday in Walker, 1851b: 7. Type species: Chionea venatica Haliday in Walker, 1856 [= Tipula atomaria De Geer, 1778], subs. mon. Haliday in Walker, 1856b: 56.
ATOMARIA Bigot, 1854: 454. Type species: Tipula atomaria De Geer, 1778, taut.

Subgenus EPIDAPUS Haliday

EPIDAPUS Haliday in Walker, 1851b: 7 (as gen.). Type species: Chionea venatica Haliday in Walker, 1856 [= Tipula atomaria De Geer, 1778], subs. mon. Haliday in Walker, 1856b: 56.
aciculatus Vilkamaa, Hippa & Mohrig, 2014: 430. New Caledonia (New Caledonia).
chaetovenosus
Mohrig in Mohrig & Jaschhof, 1999: 30. NZ (NZ).
ctenosciaroides Mohrig in Mohrig & Jaschhof, 1999: 32. NZ (NZ).
espinosalus Mohrig in Mohrig & Jaschhof, 1999: 31. NZ (NZ).
exselcus Mohrig & Kauschke, 2017: 465. Australia (Qld).
formosus Vilkamaa, Hippa & Mohrig, 2014: 433. New Caledonia (New Caledonia).
parvus Mohrig in Mohrig & Jaschhof, 1999: 35. NZ (NZ).
probus Mohrig, 2004: 150. PNG (PNG).
strenuus Mohrig, 2004: 151. PNG (PNG).
triquetrus Vilkamaa, Hippa & Mohrig, 2014: 434. New Caledonia (New Caledonia).
Undescribed sp.—Belau (Steffan, 1969: 685).

Subgenus PSEUDOAPTANOGYNA Vimmer

PSEUDOAPTANOGYNA Vimmer, 1926: 10 (Aptanogyna subg.). Type species: Aptanogyna abscondita Vimmer, 1926, mon.
angulatus Vilkamaa, Hippa & Mohrig, 2014: 431. New Caledonia (New Caledonia).
inversus Mohrig, 2004: 153. PNG (PNG).
iucundus Mohrig, 2004: 152. PNG (PNG).

Subgenus ZUHALIA Koçak & Hüseyinoglu

MACROTARSUS Mohrig, 2004: 154. Type species: Epidapus pellitus Mohrig, 2004, orig. des. [Preoccupied by Link, 1795.]
ZUHALIA Koçak & Hüseyinoglu, 2008: 2. Type species: Epidapus pellitus Mohrig, 2004, automatic.
longicubitalis Mohrig, 2004: 157. PNG (PNG).
pellitus Mohrig, 2004: 155. PNG (PNG).
primus Mohrig, 2004: 156. PNG (PNG); New Caledonia (New Caledonia).
quinquespinus Mohrig, 2004: 150. PNG (PNG).

Genus HYPERLASION Schmitz

HYPERLASION Schmitz, 1918: 95, 96. Type species: Hyperlasion wasmanni Schmitz, 1918, mon.
alienus Mohrig, 2004: 159. PNG (PNG).
magnisenorium
Hardy, 1956c: 89 (Scythropochroa). Hawaiian Is.
politum Mohrig, 2004: 160. PNG (PNG).


Genus KEILBACHIA Mohrig & Martens

KEILBACHIA Mohrig & Martens, 1987: 483. Type species: Keilbachia nepalensis Mohrig & Martens, 1987, orig. des.
adstrictaVilkamaa et al., 2011: 54. New Caledonia (New Caledonia).
adstrictatula Mohrig & Kauschke, 2017: 467. Australia (Qld).
cracens Vilkamaa et al., 2011: 54. New Caledonia (New Caledonia).
constricta Vilkamaa et al., 2011: 57. New Caledonia (New Caledonia).
fratercula Vilkamaa et al., 2011: 60. New Caledonia (New Caledonia).
iiindigena Mohrig, 2004: 165. PNG (PNG).
truncata Vilkamaa et al., 2011: 58. New Caledonia (New Caledonia).



Genus LOBOSCIARA Steffan

LOBOSCIARA Steffan, 1969: 727. Type species: Bradysia spinipennis Sasakawa, 1962, orig. des.
trilobata Vilkamaa & Hippa, 1994: 41. Indonesia (Sulawesi); Australia (Qld).
spinipennis Sasakawa, 1962: 130 (Bradysia). Thailand; Guam, Micronesia.


Genus LYCORIELLA Frey

LYCORIELLA Frey, 1942: 39. Type species: Sciara vivida Winnertz, 1867 [misid., = Sciara auripila Winnertz, 1867], orig. des.
agraria Felt, 1898a: 225 [1898b: 225] (Sciara). USA; Australia (NSW, Vic) [immigrant], NZ (NZ) [immigrant]; England.
agarici Loudon, 1978: 163. Australia (NSW).
hoyti Hardy, 1956c: 80 (Sciara). Hawaiian Is.
ingenua Dufour, 1839: 29 (Sciara). [Not given; = France]; Australia (NSW, Tas) [immigrant], Hawaiian Is [immigrant], Antarctica, Kerguelen Is, Macquarie I; Holarctic.
     mali Fitch, 1856: 486 (Molobrus). USA; Hawaiian Is.
     pauciseta Felt, 1898: 224. USA.
     womersleyi
Séguy, 1940: 210. Kerguelen Is.
sativae Johannsen, 1912: 133 (Sciara). USA; Australia (ACT, NSW, Wld, SA, Tas, Vic), Hawaiian Is, Kerguelen Is, Norfolk I; Holarctic.
     auberti Séguy, 1940: 211. Kerguelen Is.
     solispina Hardy, 1956c: 84 (Sciara). Hawaiian Is.


Genus NEOPHNYXIA Tonnoir

NEOPHNYXIA Tonnoir in Tonnoir & Edwards, 1927: 800. Type species: Neophnyxia nelsoniana Tonnoir in Tonnoir & Edwards, 1927, orig. des.
nelsoniana Tonnoir in Tonnoir & Edwards, 1927: 800. NZ (NZ).


Genus ODONTOSCIARA Rübsaamen

ODONTONYX Rübsaamen, 1894: 19, 25. Type species: Sciara nigra Wiedemann, 1821, des. Coquillett, 1910: 578. [Preocc. Stephens, 1827.]
ODONTOSCIARA Rübsaamen, 1908: 450 (n. n. for Odontonyx).
PHORODONTA Coquillett, 1910: 589 (n. n. for Odontonyx).
filipes Walker, 1865a: 102 (Sciara). "New Guinea" [= Indonesia (Irian Jaya)].
pacifica Edwards, 1924f: 569 (Phorodonta). Fiji; American Samoa, Vanuatu, Western Samoa.
politicollis Edwards, 1928c: 34 (Phorodonta). Western Samoa.


Genus PELLICIPLANTA Mohrig

PELLICIPLANTA Mohrig, 2004: 168. Type species: Pelliciplanta unica Mohrig, 2004, orig. des.
unica Mohrig, 2004: 169. PNG (PNG).


Genus PHYTOSCIARA Frey

PHYTOSCIARA Frey, 1942: 27. Type species: Sciara halterata Lengersdorf, 1926, orig. des.

Subgenus DOLICHOSCIARATuomikoski

DOLICHOSCIARA Tuomikoski, 1960: 107. Type species: Sciara flavipes Meigen, 1804, orig. des.
atrata Mohrig, 1999: 195. PNG (PNG).
bella Mohrig, 1999: 185. PNG (PNG); Australia Qld).
bellaformis Mohrig, 1999: 186. PNG (PNG).
chaetocoxa Mohrig, 1999: 191. PNG (PNG).
conturbata Mohrig, 1999: 183. PNG (PNG); Australia (Qld).
filispinosa Mohrig, 1999: 188. PNG (PNG).
fumida Mohrig, 1999: 190. PNG (PNG).
gilvifucata Mohrig, 1999: 193. PNG (PNG).
pseudoornata Mohrig, 1999: 187. PNG (PNG); Australia (Qld).
ruficoxa Brunetti, 1912: 146 (Sciara). India; PNG (PNG); Indonesia (Sumatra), Malaysia (Sabah).
subfumida Mohrig, 1999: 192. PNG (PNG).

Subgenus PHYTOSCIARA Frey

PHYTOSCIARA Frey, 1942: 27 (as gen.). Type species: Sciara halterata Lengersdorf, 1926, orig. des.
brachygaster Steffan, 1969: 709. Belau; Micronesia.
maculosa Mohrig, 1999: 182. PNG (PNG).
vulcanata
Steffan, 1973: 358. Hawaiian Is.

Genus PLASTOSCIARA Berg

PSEUDOSCIARA Kieffer, 1898a: 194. Type species: Pseudosciara pictiventris Kieffer, 1898, orig. des. [Preocc. Schiner, 1866.]
PLASTOSCIARA Berg, 1899: 78 (n. n. for Pseudosciara).
COSMOSCIARA Frey, 1942: 39. Type species: Plastosciara perniciosa Edwards, 1922, orig. des.
adrostylata Hardy, 1956c: 72. Hawaiian Is.
flavibasis Edwards, 1928c: 35. American Samoa; Fiji.
jaluitensis Steffan, 1969: 688. Marshall Is.
latipons Hardy, 1956c: 77. Hawaiian Is; Marshall Is, Micronesia.
longicosta Hardy, 1956c: 75. Hawaiian Is.
yapensis Steffan, 1969: 692. Micronesia.

Genus PNYXIA Johannsen

PNYXIA Johannsen, 1912: 114. Type species: Epidapus scabiei Hopkins, 1895, orig. des.
Undescribed sp.--Hawaiian Is (Steffan, 1981: 140).


Genus PSEUDOLYCORIELLA Menzel & Mohrig

PSEUDOLYCORIELLA Menzel & Mohrig, 1998: 367. Type species: Sciara bruckii Winnertz, 1867, orig. des.
acicula Vilkamaa et al., 2012: 3. New Caledonia (New Caledonia).
attrita
Mohrig, 2014: 144. PNG (PNG).
bispina Mohrig in Mohrig & Jaschhof, 1999: 40. NZ (NZ).
bisulca Vilkamaa et al., 2012: 3. New Caledonia (New Caledonia).
bitorquia Mohrig, 2014: 162. PNG (PNG).
breviseta Mohrig in Mohrig & Jaschhof, 1999: 41. NZ (NZ).
cavatiostyla
Mohrig, 2014: 152. PNG (PNG).
cavicata
Skuse, 1888b: 688 (Sciara). Australia (NSW).
     familiaris
Skuse, 1888b: 687 (Sciara). Australia (NSW).
     festiva Skuse, 1888b: 689 (Sciara). Australia (NSW).
capillosa Vilkamaa et al., 2012: 6. New Caledonia (New Caledonia).
commoda
Mohrig, 2014: 149. PNG (PNG).
densispina
Mohrig, 2014: 140. PNG (PNG).
espinosa
Mohrig, 2014: 146. PNG (PNG).
espinula
Mohrig, 2014: 145. PNG (PNG).
forunata
Mohrig, 2014: 151. PNG (PNG).
frederickedwardsi
Köhler in Köhler & Mohrig, 2016: 11. NZ (NZ).
ignobilis
Skuse, 1888b: 717 (Sciara). Australia (NSW).
jejuna Edwards in Tonnoir & Edwards, 1927: 796 (Sciara). NZ (NZ).
latiflagellata Vilkamaa et al., 2012: 8. New Caledonia (New Caledonia).
longicostalis Mohrig, 2014: 139. PNG (PNG).
macrotegmenta Mohrig in Mohrig & Jaschhof, 1999: 41. NZ (NZ).
macrotrichata Mohrig, 2014: 157. PNG (PNG).
microphalli Mohrig, 2014: 161. PNG (PNG).
microtrichata Mohrig, 2014: 159. PNG (PNG).
pallidula Vilkamaa et al., 2012: 9. New Caledonia (New Caledonia).
paucispina Mohrig, 2014: 142. PNG (PNG).
perspicua Mohrig, 2014: 163. PNG (PNG).
praecipua Mohrig, 2014: 155. PNG (PNG).
quadrispinosa Mohrig, 2014: 154. PNG (PNG).
senticosa Vilkamaa et al., 2012: 10. New Caledonia (New Caledonia).
setigera
Hardy, 1960: 234 (Spathobdella). Hawaiian Is.
simplex Vilkamaa et al., 2012: 10. New Caledonia (New Caledonia).
sinoupalpa Mohrig, 2014: 143. PNG (PNG).
skusei Mohrig, Kauschke & Broadley, 2016: 141. Norfolk I; Australia (Qld).
snellingi
Mohrig, 2014: 156. PNG (PNG).
spicata Vilkamaa et al., 2012: 12. New Caledonia (New Caledonia).
tenebriocoxa Mohrig, 2014: 159. PNG (PNG).
tenebriolata Mohrig, 2014: 137. PNG (PNG).
tenuis Vilkamaa et al., 2012: 14. New Caledonia (New Caledonia).
tonnoiri
Köhler in Köhler & Mohrig, 2016: 14. NZ (NZ).
tribulosa
Vilkamaa et al., 2012: 15. New Caledonia (New Caledonia).
tricanthula Mohrig, 2014: 147. PNG (PNG).
trispicata Vilkamaa et al., 2012: 16. New Caledonia (New Caledonia).
tumidior Vilkamaa et al., 2012: 17. New Caledonia (New Caledonia).
vestita Mohrig, 2014: 150. PNG (PNG).
villosa Vilkamaa et al., 2012: 20. New Caledonia (New Caledonia).
zealandica Edwards in Tonnoir & Edwards, 1927: 796 (Sciara). NZ (NZ). [Preocc. Kieffer, 1910; no n. n. proposed at this time.]


Genus PSEUDOZYGOMMA Mohrig

PSEUDOZYGOMMA Mohrig, 2004: 144. Type species: Pseudozygomma contraria Mohrig, 2004, orig. des.
contraria Mohrig, 2004: 145. PNG (PNG).
flavoabdominalis Mohrig, 2004: 147. PNG (PNG).
maculipennis Skuse, 1890c: 411 (Zygoneura). Australia (NSW).
nigroalata
Mohrig, 2004: 148. PNG (PNG).
nigrocorporea Mohrig, 2004: 146. PNG (PNG).


Genus PSEUDOZYGONEURA Steffan

PSEUDOZYGONEURA Steffan, 1969: 676. Type species: Pseudozygoneura musicola Steffan, 1969, orig. des.
musicola Steffan, 1969: 677. Belau; Fiji, Hawaiian Is [introduced; ?established], Micronesia, Solomon Is.


Genus SCATOPSCIARA Edwards

SCAPTOPSCIARA Edwards in Tonnoir & Edwards, 1927: 798 (Sciara subg.). Type species: Sciara quinquelineata Macquart, 1834 [= Sciara vitripennis Meigen, 1818], orig. des.

Subgenus SCATOPSCIARA Edwards

SCAPTOPSCIARA Edwards in Tonnoir & Edwards, 1927: 798 (Sciara subg.). Type species: Sciara quinquelineata Macquart, 1834 [= Sciara vitripennis Meigen, 1818], orig. des.
UDMANNIA Frey, 1942: 38. Type species: Sciara pusilla Meigen, 1818, orig. des. [Preocc. Bergroth, 1915.]
UDMANNIELLA Frey, 1948: 86 (n. n. for Udmannia).
crassivena Mohrig, 2004: 167. PNG (PNG).
nigrita
Hardy, 1956c: 86 (Scatopsciara). Hawaiian Is.
spiculata Vilkamaa et al., 2012: 69. New Caledonia (New Caledonia).
tenuistyla Vilkamaa et al., 2012: 71. New Caledonia (New Caledonia).
unicalcarata Edwards in Tonnoir & Edwards, 1927: 798 (Sciara). NZ (NZ).
validovenosa Mohrig & Kauschke, 2017: 477. Australia (Qld).
vicina
Vilkamaa et al., 2012: 72. New Caledonia (New Caledonia).

Subgenus XENOPYGINA Frey

XENOPYGINA Frey, 1948: 51 (Bradysia subg.). Type species: Bradysia paradoxa Frey, 1948, orig. des.
ampliocerosa Mohrig, 2004: 167. PNG (PNG).
brevicolla Mohrig & Kauschke, 2017: 473. Australia (Qld).
exigua
Vilkamaa et al., 2012: 68. New Caledonia (New Caledonia).
monocerosa
Mohrig, 2004: 166. PNG (PNG).

Unplaced to Subgenus of SCAPTOSCIARA

dubiosa Mohrig & Kauschke, 2017: 475. Australia (Qld).



Genus SCHWENKFELDINA Frey

SCHWENKFELDINA Frey, 1942: 32 (Neosciara subg.). Type species: Sciara carbonaria Meigen, 1820, orig. des.
inopinata Mohrig, 1999: 157. PNG (PNG).
ponderosa Walker, 1864: 230 (Sciara). Indonesia (Maluku).
regia Mohrig, 1999: 156. PNG (PNG).

Genus SCIARA Meigen

LYCORIA Meigen, 1800: 17. Suppressed by I.C.Z.N., 1963b: 339.
SCIARA Meigen, 1803: 263. Type species: Tipula thomae Linnaeus, 1767, mon.
adjuncta Skuse, 1890c: 398. Australia (NSW).
aemula Skuse, 1888b: 678. Australia (NSW).
approximata Skuse, 1888b: 697. Australia (NSW).
aquila Skuse, 1888b: 692. Australia (NSW).
atratula Skuse, 1888b: 704. Australia (NSW).
audax Skuse, 1888b: 693. Australia (NSW).
brevifurca Skuse, 1888b: 700. Australia (NSW).
consanguinea Skuse, 1890c: 401. Australia (NSW).
contermina Skuse, 1890c: 395. Australia (NSW).
distigma Edwards, 1924f: 568. Fiji.
diversa Skuse, 1888b: 701. Australia (NSW).
dolosa Skuse, 1888b: 683. Australia (NSW).
erratica Skuse, 1888b: 696. Australia (NSW).
evanescens Skuse, 1888b: 698. Australia (NSW).
exposita Skuse, 1890c: 397. Australia (NSW).
femoralis Doleschall, 1858: 78. Indonesia (Maluku).
festina Skuse, 1888b: 684. Australia (NSW).
finitima Skuse, 1888b: 677. Australia (NSW).
flavicoxis Skuse, 1890c: 396. Australia (NSW).
fumipennis Skuse, 1888b: 706. Australia (NSW).
helmsi Skuse, 1890c: 408. Australia (NSW).
infixa Skuse, 1890c: 400. Australia (NSW).
lucidipennis Skuse, 1888b: 714. Australia (NSW).
luculenta Skuse, 1888b: 705. Australia (NSW).
marcilla Hutton, 1902b: 192. NZ (NZ).
marginata Skuse, 1890c: 399. Australia (NSW).
neorufescens Miller, 1950: 57 (n. n. for rufescens Hutton). NZ (NZ).
     rufescens Hutton, 1881: 13. NZ (NZ). [Preocc. Zetterstedt, 1852.]
nubicula Skuse, 1888b: 715. Australia (NSW).
palliceps Edwards, 1928: 25. Malaysia (Pen); PNG (PNG); Indonesia (Sumatra), Philippines, Thailand.
philpotti Tonnoir in Tonnoir & Edwards, 1927: 796. NZ (NZ).
praescellens Skuse, 1890c: 404. Australia (NSW).
prominens Hardy, 1956c: 83. Hawaiian Is.
promiscua Skuse, 1890c: 394. Australia (NSW).
reciproca Walker, 1856d: 420. Australia (Tas).
recondita Skuse, 1890c: 390. Australia (NSW).
scitula Skuse, 1888b: 699. Australia (NSW).
sedula Skuse, 1888b: 674. Australia (NSW).
selecta Walker, 1858b: 77. Indonesia (Maluku).
sororia Skuse, 1888b: 716. Australia (NSW).
tepperi Skuse, 1890c: 393. Australia (SA).
tryoni Skuse, 1890c: 391. Australia (Qld).
     insulana Vilkamaa, Hippa & Mohrig, 2015: 590. New Caledonia (New Caledonia).
turgidula
Vilkamaa, Hippa & Mohrig, 2015: 592. New Caledonia (New Caledonia).
turrida de Meijere, 1913a: 45. Indonesia (Maluku).
varipes Walker, 1865a: 102. "New Guinea" [= Indonesia (Irian Jaya)].
vecors Skuse, 1888b: 695. Australia (NSW).
zealandica Kieffer, 1910a: 327. NZ (NZ).

Genus SCYTHROPOCHROA Edwards

SCYTHROPOCHROA Enderlein, 1911c: 138. Type species: Scythropochroa latefurcata Enderlein, 1911, orig. des.
antarctica Hudson, 1892: pl. 4, fig. 5a,b (Mycetophila). Not given [NZ (NZ)]. Nomen dubium.
atrichata Mohrig, 2004: 134. PNG (PNG).
atrichoalata Mohrig, 2004: 135. PNG (PNG).
gressitti
Steffan, 1969: 680. Micronesia; Guam.
macleayi Skuse, 1888b: 691 (Sciara). Australia (NSW).
micropalpa
Mohrig, 2004: 137. PNG (PNG).
nitida
Edwards in Tonnoir & Edwards, 1927: 799. NZ (NZ).
parapectinea Mohrig, 2004: 138. PNG (PNG).
paucitrichata Mohrig, 2004: 132. PNG (PNG).
quadrispinosa
Steffan, 1969: 682. Belau.
samoana Edwards, 1928c: 36. Western Samoa.
semitrichata Mohrig, 2004: 131. PNG (PNG).
trichoalata Mohrig, 2004: 130. PNG (PNG).
trichovenosa Mohrig, 2004: 133. PNG (PNG).
trispinosa
Steffan, 1969: 682. Marshall Is.

 

Genus VULAGISCIARA Evenhuis

VULAGISCIARA Evenhuis in Evenhuis et al., 2007: 4. Type species: Vulagisciara myrmecophila Evenhuis, 2007, orig. des.
myrmecophila Evenhuis in Evenhuis et al., 2007: 4. Fiji.

Genus XYLOSCIARA Tuomikoski

PLASTOSCIARA Frey, 1942: 34. Type species: Sciara lignicola Winnertz, 1867, orig. des. [Preocc. Berg, 1899.]
XYLOSCIARA Tuomikoski, 1957: 10 (n. n. for Plastosciara).
brevipes Steffan, 1964: 293. NZ (Campbell I).

Genus ZYGONEURA Meigen

ZYGONEURA Meigen, 1830: 304. Type species: Zygoneura sciarina Meigen, 1830, mon.
contractans Edwards in Tonnoir & Edwards, 1927: 795 (Sciara). NZ (NZ).
remota de Meijere, 1913b: 305. Indonesia (Irian Jaya).


Last revised 17 June 2018