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86. Family SCIOMYZIDAE

[Originally] by Jeffrey K. Barnes & Lloyd V. Knutson

The Sciomyzidae, or marsh flies, consist of 4 subfamilies, with 600 species in about 60 genera worldwide. The family is poorly represented in the Australasian/Oceanian Regions, where only 2 subfamilies, 7 genera, and 37 native species are known. The familial placement of the Huttonininae, with 1 genera and 7 species, has been elevated to its own family as has the Phaeomyiinae, with its single genus Pelidnoptera and one species found in the Australasian/Oceanian regions.

Life cycles of about 200 species of sciomyzid flies have been studied. Larvae of all reared species are predators or parasitoids of molluscs, especially aquatic, amphibious, and terrestrial pulmonate snails. The larvae of Australasian and Oceanian species show diverse feeding habits. Larvae of Dichetophora, Sepedomerus, and Sepedon spp. have been observed to feed on aquatic pulmonate snails. Larvae of Eulimnia feed on sphaeriid clams, and larvae of Neolimnia feed on aquatic or terrestrial, pulmonate, or operculate prosobranch snails (Lynch 1965; Neff & Berg 1966; Fontana 1972; Bhuangprakone & Areekul 1973; Beaver et al. 1977; Barnes 1979c, 1980a). Several species of Sciomyzidae were introduced into the Hawaiian Islands as biological control agents for aquatic snails that are intermediate hosts of Fasciola, but only Sepedomerus macropus and Sepedon aenescens are believed to be established (Davis 1960, Chock et al. 1961, Davis et al. 1961, Davis & Krauss 1962, 1967, Davis 1972, Steyskal in Hardy & Delfinado 1980).
Refs.: Harrison (1959, key NZ spp), Berg & Knutson (1978, bibliogr.),Barnes (1979b, 1980a, key NZ spp.), Steyskal in Hardy & Delfinado (1980, key Hawaiian spp.).


Tribe PROSOCHAETINI

Genus PROSOCHAETA Malloch

PROSOCHAETA Malloch, 1935b: 95. Type species: Prosochaeta prima Malloch, 1935, mon.
prima Malloch, 1935b: 95. NZ (NZ).

Subfamily SCIOMYZINAE
Tribe SCIOMYZINI

Genus ATRICHOMELINA Cresson

ATRICHOMELINA Cresson, 1920: 40. Type species: Sciomyza pubera Loew, 1862, orig. des.
ACHAETOMELINA. Incorr. orig. spell. of Atrichomelina (Cresson, 1920: 30).
pubera Loew, 1862a: 106 (Sciomyza). USA; Hawaiian Is [introduced; ?established]; Canada, USA, Mexico.

Genus PHERBELLIA Robineau-Desvoidy

MELINA Robineau-Desvoidy, 1830: 695. Type species: Melina riparia Robineau-Desvoidy, 1830 [= Sciomyza dubia Fallén, 1820], mon. [Preocc. Retzius, 1788.]
PHERBELLIA Robineau-Desvoidy, 1830: 695. Type species: Pherbellia vernalis Robineau-Desvoidy, 1830 [= Sciomyza schoenherri Fallén, 1826], mon.

Subgenus CHETOCERA Robineau-Desvoidy

CHETOCERA Robineau-Desvoidy, 1830: 697 (as gen.). Type species: Chetocera claripennis Robineau-Desvoidy, 1830 [= Sciomyza albocostata Fallén, 1820], mon.
CHAETOCERA Agassiz, 1846b: 54 (unjust. emend. of Chetocera).
dorsata Zetterstedt, 1846: 2096 (Sciomyza). Sweden; Hawaiian Is [introduced; ?established]; England, E to Kazakh SSR, S to SE Spain, Turkey & Israel.

Subgenus DITAENIELLA Sack

DITAENIELLA Sack, 1939: 37 (as gen.). Type species: Sciomyza grisescens Meigen, 1830, orig. des.
grisescens Meigen--not Australasian/Oceanian.
parallela Walker, 1853: 401 (Sciomyza). USA; Hawaiian Is [introduced; ?established]; S Canada to Costa Rica.

Subgenus GRAPHOMYZINA Macquart

GRAPHOMYZINA Macquart, 1835: 558 (as gen.). Type species: Graphomyzina elegans Macquart, 1835 [= Sciomyza limbata Meigen, 1830], mon.
CORMOPTERA Schiner, 1862a: 149 (unnec. n. n. for Graphomyzina).
dives Bezzi, 1928: 85 (Sciomyza). Fiji; Tonga.
javana de Meijere, 1911b: 349 (Sciomyza). Indonesia (Java); widesp. Australia (excl. Tas); Indonesia (Sumatra).
juxtajavana Knutson, Manguin & Orth, 1990: 281. Australia (NT); widesp. Australia.

Tribe TETANOCERINI

Genus DICHETOPHORA Rondani

DICHETOPHORA Rondani, 1868a: 206 [1868b: 8]. Type species: Scatophaga obliterata Fabricius, 1805, orig. des.

Subgenus DICHETOPHORA Rondani

DICHETOPHORA Rondani, 1868a: 206 [1868b: 8] (as gen.). Type species: Scatophaga obliterata Fabricius, 1805, orig. des.
DICHAETOPHORA, error for Dichetophora.
australis Walker, 1853: 402 (Tetanocera). Australia (Tas).

Subgenus NEOSEPEDON Malloch

NEOSEPEDON Malloch, 1928f: 323. Type species: Dichaetophora punctipennis Malloch, 1928, orig. des.
biroi Kertész, 1901c: 418 (Cylindria). Australia (NSW); Australia (Qld, SA, Vic).
boyesi Steyskal, 1972b: 362. Australia (WA).
conjuncta Malloch, 1928f: 324. Australia (NSW); Australia (Vic).
hendeli Kertész, 1901c: 417 (Cylindria). Australia (NSW).
punctipennis Malloch, 1928f: 323. Australia (NSW).

Genus DICTYA Meigen

DICTYA Meigen, 1803: 277. Type species: Musca umbrarum Linnaeus, 1758 [designation pending submission of application to I.C.Z.N. for fixation of type species].
MONOCHAETOPHORA Hendel, 1900: 355. Type species: Musca umbrarum Linnaeus, 1758, mon.
abnormis Steyskal, 1954: 518. Mexico; Hawaiian Is [introduced; ?established].

Genus EULIMNIA Tonnoir & Malloch

EULIMNIA Tonnoir & Malloch, 1928: 172. Type species: Eulimnia milleri Tonnoir & Malloch, 1928, orig. des.
milleri Tonnoir & Malloch, 1928: 172. NZ (NZ).
philpotti Tonnoir & Malloch, 1928: 173. NZ (NZ).

Genus NEOLIMNIA Tonnoir & Malloch

NEOLIMNIA Tonnoir & Malloch, 1928: 163. Type species: Limnia obscura Hutton, 1901, orig. des.

Subgenus NEOLIMNIA Tonnoir & Malloch

NEOLIMNIA Tonnoir & Malloch, 1928: 165. Type species: Limnia obscura Hutton, 1901, orig. des.
castanea Hutton, 1904c: 153 (Limnia). NZ (NZ).
    ocellata Tonnoir & Malloch, 1928: 166. NZ (NZ).
diversa Tonnoir & Malloch, 1928: 167. NZ (NZ).
irrorata Tonnoir & Malloch, 1928: 167. NZ (NZ).
    dubiosa Tonnoir & Malloch, 1928: 168. NZ (NZ).
minuta Tonnoir & Malloch, 1928: 170. NZ (NZ).
obscura Hutton, 1901: 85. (Limnia). NZ (NZ).
pepekeiti Barnes, 1979b: 260. NZ (NZ).
raiti Barnes, 1979b: 261. NZ (NZ).
striata Hutton, 1904c: 154 (Limnia). NZ (NZ); NZ (Chatham Is).
  ssp. brunneifrons Tonnoir & Malloch, 1928: 169 (striata var.). NZ (NZ).
  ssp. striata s. str. NZ (NZ); NZ (Chatham Is).

Subgenus PSEUDOLIMNIA Tonnoir & Malloch

PSEUDOLIMNIA Tonnoir & Malloch, 1928: 171. Type species: Tetanocera sigma Walker, 1849, orig. des.
repo Barnes, 1979b: 245. NZ (NZ).
sigma Walker, 1849: 1084 (Tetanocera). NZ (NZ).
tranquilla Hutton, 1901: 85 (Limnia). NZ (NZ).
ura Barnes, 1979b: 251. NZ (NZ).

Subgenus SUBLIMNIA Harrison

SUBLIMNIA Harrison, 1959: 72. Type species: Neolimnia nitidiventris Tonnoir & Malloch, 1928, orig. des.
nitidiventris Tonnoir & Malloch, 1928: 170. NZ (NZ).
vittata Harrison, 1959: 72. NZ (NZ).

Genus SEPEDOMERUS Steyskal

SEPEDOMERUS Steyskal, 1973: 144. Type species: Sepedon macropus Walker, 1849, orig. des.
macropus Walker, 1849: 1078 (Sepedon). Jamaica; Hawaiian Is [introduced]; S USA & Mexico E to Puerto Rico & Trinidad, S to Peru.

Genus SEPEDON Latreille

SEPEDON Latreille, 1804: 196. Type species: Syrphus sphegeus Fabricius, 1775, mon.
SEPODON, error for Sepedon.
aenescens Wiedemann, 1830a: 579. China; Hawaiian Is [introduced]; widesp. Oriental Reg.
    violacea Hendel, 1909c: 86. Hong Kong.
    sauteri Hendel, 1911b: 270. Taiwan.
    sinensis Mayer, 1953: 217. China.
batjanensis Kertész, 1899b: 180 (n. n. for costalis Walker). Indonesia (Maluku); Indonesia (Sulawesi).
     costalis Walker, 1861d: 291. Indonesia (Maluku). [Preocc. Walker, 1858.]
    batjanensis Brunetti, 1907: 214 (n. n. for costalis Walker). [Preocc. Kertész, 1899.]
costalis Walker, 1858b: 110. Indonesia (Maluku); PNG (PNG).
lata Bezzi, 1928: 84 (costalis var.). Fiji; PNG (PNG). N. stat.
noteoi Steyskal--not Australasian/Oceanian.
oriens Steyskal in Hardy & Delfinado 1980: 119. Hawaiian Is [introduced; ?established].
pacifica Cresson, 1914b: 457. USA, Hawaiian Is [introduced; ?established]; Canada, Mexico.
plumbella Wiedemann, 1830a: 577. "China"; Australia (NT, Qld, WA), Hawaiian Is [introduced; ?established], Indonesia (Irian Jaya), New Caledonia (New Caledonia), PNG (Bougainville I, PNG), Solomon Is; widesp. Oriental Reg.
    javanensis Robineau-Desvoidy, 1830: 677. Indonesia (Java).
    fuscinervis Brunetti, 1907: 215. Indonesia (Java).
    sanguinipes Brunetti, 1907: 215. Indonesia (Java).
    javana, error for javanensis.
praemiosa Giglio-Tos, 1893: 8. Mexico; Hawaiian Is [introduced; ?established]; USA.
senex Wiedemann, 1830a: 578. Not given [unknown]; Hawaiian Is [introduced; ?established]; widesp. Oriental Reg.

Unplaced species of Sciomyzidae

transmarina Schiner, 1868: 234 (Limnia). NZ (NZ).



Last revised 9 December 2017