Tephritidae -- a/o cat.

Australasian/Oceanian Diptera Catalog -- Web Version

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[Originally] by D. Elmo Hardy & Richard H. Foote

Tephritidae is a large family of predominantly picture-winged, usually highly ornamented flies with brightly contrasting color patterns on the bodies. Over 6,200 species are known worldwide. This catalog lists 829 species in 150 genera. Fruit flies of most of the world's tropics are still poorly known, and comparatively little biological data or host information are available for most of the fauna. There is no universal agreement on the higher classification of the family on a worldwide basis. For this catalog we prefer to arrange the genera in 3 subfamilies: Dacinae, Trypetinae, and Tephritinae.

Although members of this family are commonly called fruit flies, the vast majority of the world's species are not frugivorous. Species of the subfamily Dacinae are exclusively fruit breeders and attack all kinds of fleshy fruits, including those of cucurbitaceous and solanaceous crops, and members of the genus Dacus s. l. comprise the major economic pests of this region. Except for some genera of fruit-breeding Trypetinae, the non-dacine Tephritidae breed predominantly in various tissues of living plants. Tephritinae breed predominantly in flower heads of Compositae (Asteraceae), and some are gall formers and stem borers in Compositae, Goodenaceae, and some other plant families. The Trypetinae have a wide range of biological associations: some are fruit breeders (largely in native and endemic hosts) and are seldom economically important in these regions, some are leaf miners; some infest flower heads (feeding on seeds and ovaries); some breed in buds of large flowers such as Bauhinia, Bombax, and Hibiscus; many of the Acanthonevrini, especially Gastrozonina, are associated with bamboo, with some boring in the young shoots, others in the terminal growths. Some Trypetinae show a marked departure from the usual breeding habits of the family: Termitorioxa Hendel has been reared from galleries of Mastotermes in northern Australia (Hill, 1921); Phytalmiini apparently breed in rotting bark or wood; species of Dirioxa have been reared from decaying fruit and rotting bark of mahogany trees (Dysoxylum) in northern Queensland (1 species of Dirioxa has been reared from decaying wood); Dacopsis breeds in thick stringy bark of fallen Dysoxylum trees in Papua New Guinea; and some species of Themara have been collected commonly on trunks of fallen trees and may be bark breeders. The strangest association known for a "fruit fly" is that of Alloeomyia flavida from Papua New Guinea, which has been collected only in traps baited with human feces; no doubt it is a scavenger.

Refs.: Hering (1947b, key world Tephritidae subfams., tribes), Hardy (1956a, classif. world Dacinae; 1977b, cat. Oriental spp.; 1983b, rev. Indonesian, New Guinean, adjacent Is Euphrantini; 1985, rev. Indonesian, New Guinean Schistopterinae; 1986a, rev. Indonesian, New Guinean, adjacent Is Adramini; 1986b, rev. Indonesian, New Guinean, Bismarck Arch, Solomon Is Acanthonevrina; 1987, rev. Indonesian, New Guinean, Bismarck Arch, Solomon Is Trypetini, Aciurini, Ceratitini; 1988, rev. Indonesian, New Guinean, Bismarck Arch, Solomon Is, Tephritinae), Drew (1972a, gen., subg. classif. S Pacific Dacinae; 1975, zoogeogr. S Pacific Dacinae; 1978, econ. import. S Pacific Dacinae; 1989, rev. A/O Dacinae), Hardy & Delfinado (1980, rev. Hawaiian spp.).

Subfamily DACINAE



ACIDOXANTHA Hendel, 1914c: 83. Type species: Acidoxantha punctiventris Hendel, 1914, orig. des.
ACIDIOXANTHA Hendel, 1915: 450 (unjust. emend. of Acidoxantha).
quinaria Permkam & Hancock, 1995a: 1326. Australia (WA).
Undetermined sp.—Indonesia (Maluku), PNG (PNG) (Hardy, 1987: 272).


CARPOPHTHORELLA Hendel, 1914c: 80 [1915: 448]. Type species: Carpophthorella magnifica Hendel, 1914 [= Trypeta nigrifascia Walker, 1860], mon.
nigrifascia Walker, 1860a: 158 (Trypeta). Indonesia (Sulawesi); Australia (Qld), Indonesia (Maluku), PNG (PNG), Solomon Is; Indonesia (Sumatra), Malaysia (Sarawak).
     retorta Walker, 1861e: 16 (Trypeta). Indonesia (Maluku).
     bifasciata de Meijere, 1916a: 48 (Gastrozona). Indonesia (Sumatra).
     albiscutellata Enderlein, 1920a: 354 (Gastrozona). Indonesia (Sumatra).
     setifrons Malloch, 1939d: 263. Solomon Is.


CERATITELLA Malloch, 1939f: 452. Type species: Ceratitis loranthi Froggatt, 1911, orig. des.
amyemae Permkam & Hancock, 1995a: 1330. Australia (NT); Australia (Qld).
bifasciata Hardy, 1967: 133. Australia (Qld); Australia (NSW), Indonesia (Irian Jaya), PNG (PNG).
loranthi Froggatt, 1911: 863 (Ceratitis). Australia (WA); Australia (ACT, NSW, NT, SA, Vic).
recondita Permkam & Hancock, 1995a: 1334. Australia (Qld); Australia (NSW).
schlingeri Norrbom & Hancock, 2004: 70. New Caledonia (New Caledonia).
solomonensis Hancock & Drew, 2003: 11. Solomon Is.
unifasciata Hardy, 1967: 137. Australia (Qld); Australia (NSW).

Genus CERATITIS Macleay

CERATITIS Macleay, 1829: 482. Type species: Ceratitis citriperda Macleay, 1829 [= Tephritis capitata Wiedemann, 1824], mon.
CERATITES Agassiz, 1846: (unjust. emend. of Ceratitis).
HALTEROPHORA Rondani, 1861: 10 (unnec. n.n. for Ceratitis). [Preocc. Enlicher, 1836.]
capitata Wiedemann, 1824: 55 (Tephritis). "India Orient"; widesp. W Australia, Hawaiian Is [immigrant]; USA [immigrant], pantrop.


PARACERATITELLA Hardy, 1967: 138. Type species: Paraceratitella oblonga Hardy, 1967, orig. des.
compta Hardy, 1987: 342. PNG (PNG); Australia (Qld).
connexa Hardy, 1987: 344. Solomon Is.
eurycephala Hardy, 1967: 140. Australia (Qld); Australia (NSW, NT, WA).
oblonga Hardy, 1967: 143. Australia (Qld).

Genus BACTROCERA Macquart

BACTROCERA Macquart, 1835: 452. Type species: Bactrocera longicornis Macquart, 1835, mon.

Subgenus AFRODACUS Bezzi

AFRODACUS Bezzi, 1924b: 469 (as gen.). Type species: Chaetodacus biguttulus Bezzi, 1922, mon.
brunnea Perkins & May, 1949: 18 (Afrodacus). Australia (Qld).
grandistylus Drew & Hancock, 1995: 7. New Caledonia (New Caledonia).
hypomelaina Drew, 1989: 20. PNG (PNG).
jarvisi Tryon, 1927: 201 (Chaetodacus). Australia (Qld); Australia (NSW, NT, WA), Lord Howe I.
     careya Tryon, 1927: 202 (jarvisi var.). Nomen nudum.
     australis Hendel, 1928b: 341 (Dacus). Australia (NT).
     halterata Hendel, 1928b: 342 (Dacus australis var.). Australia (Qld).
minuta Drew, 1971: 29 (Dacus). Vanuatu.
ochracea Drew, 1989: 23. PNG (PNG).

Subgenus ASIADACUS Perkins

ASIADACUS Perkins, 1937: 57 (as gen.). Type species: Chaetodacus bakeri Bezzi, 1919, orig. des.
absolutus Walker, 1861e: 22 (Dacus). Indonesia (Maluku).
atypica White & Evenhuis, 1999: 490. Indonesia (Irian Jaya).

Subgenus AUSTRODACUS Perkins

AUSTRODACUS Perkins, 1937: 56 (as gen.). Type species: Dacus cucumis French, 1907, orig. des.
cucumis French, 1907: 307 (Dacus tryoni var.). Australia (Qld); Australia (NSW, NT).

Subgenus BACTROCERA Macquart

BACTROCERA Macquart, 1835: 452 (as gen.). Type species: Bactrocera longicornis Macquart, 1835, mon.
BACTROCERA Guérin-Méneville, 1838: 300. Type species: Bactrocera longicornis Guérin-Méneville, 1838 [preocc.], mon. [Preocc. Macquart, 1835.]
DASYNEURA Saunders, 1842: 60 (as gen.). Type species: Dasyneura zonata Saunders, 1842, mon.
STRUMETA Walker, 1856a: 33 (as gen.). Type species: Strumeta conformis Walker, 1856 [= Dacus umbrosus Fabricius, 1805], mon.
CHAETODACUS Bezzi, 1913b: 93 (as gen.). Type species: Musca ferruginea Fabricius, 1784 [preocc., = Dacus dorsalis Hendel, 1912], orig. des.
MARQUESADACUS Malloch, 1932m: 145. Type species: Chaetodacus perfuscus Aubertin, 1929, orig. des.
APODACUS Perkins, 1939: 26 (as gen.). Type species: Apodacus cheesmanae Perkins, 1939 (as cheesemani), orig. des.
abdofuscata Drew, 1971: 48 (Dacus). PNG (PNG).
abdolonginqua Drew, 1971: 50 (Dacus). PNG (Bismarck Arch).
abdonigella Drew, 1971: 52 (Dacus). PNG (PNG); Australia (Qld).
abscondita Drew & Hancock in Drew et al., 1981: 54 (Dacus). Australia (Qld).
absidata Drew, 1989: 101. PNG (PNG).
abundans Drew, 1989: 106. PNG (PNG); Australia (Qld).
aemula Drew, 1989: 26. PNG (PNG).
aeroginosa Drew & Hancock in Drew et al., 1981: 57 (Dacus). Australia (Qld).
aithogaster Drew, 1989: 117. Solomon Is.
allwoodi Drew, 1979: 79 (Dacus). Australia (NT).
alyxiae May, 1953: 335 (Strumeta). Australia (Qld); PNG (PNG).
amplexiseta May, 1962: 66 (Strumeta). Australia (Qld).
ampla Drew, 1971: 55 (Dacus). PNG (Bismarck Arch).
anfracta Drew, 1989: 102. PNG (PNG); Australia (Qld).
angustifasciata Drew, 1989: 119. PNG (Bismarck Arch).
anomala Drew, 1971: 57 (Dacus). Vanuatu.
anthracina Drew, 1971: 59 (Dacus). PNG (Bismarck Arch).
antigone Drew & Hancock in Drew et al., 1981: 59 (Dacus). Australia (Qld).
aquilonis May, 1965: 62 (Strumeta). Australia (NT); Australia (WA).
assita Drew, 1989: 36. PNG (PNG).
atra Malloch, 1938a: 113 (Dacus). French Polynesia (Austral Is).
aterrima Drew, 1972b: 204 (Dacus). PNG (Bougainville I); Solomon Is.
atribifasciata Drew & Romig, 2001: 118. Solomon Is.
atriliniellata Drew, 1989: 52. PNG (PNG).
aurantiaca Drew & Hancock in Drew et al., 1981: 62 (Dacus). Australia (Qld); PNG (PNG).
bancroftii Tryon, 1927: 199 (Chaetodacus). Australia (Qld).
barringtoniae Tryon, 1927: 196 (Chaetodacus). Australia (Qld).
batemani Drew, 1989: 94. Australia (Qld).
biarcuata Walker, 1865b: 122 (Dacus). "New Guinea" [= Indonesia (Irian Jaya)]; PNG (Bougainville I, PNG), Solomon Is.
bidentata May, 1963: 527 (Strumeta). Australia (Qld).
breviaculea Hardy, 1951b: 145 (Dacus). Australia (Qld); PNG (PNG).
brevistriata Drew, 1968b: 77 (Strumeta). PNG (PNG).
bryoniae Tryon, 1927: 192 (Chaetodacus). Australia (Qld); Australia (NSW, NT, WA), PNG (Bismarck Arch, PNG).
     costalis Shiraki, 1933: 66 (Chaetodacus). Japan.
buloloensis Drew, 1989: 125. PNG (PNG).
cacuminatus Hering, 1941c: 46 (Strumeta). Australia (Qld); Australia (NSW, Vic), Lord Howe I.
     solani Jarvis, 1922a: 247 [1922b: 309] (Dacus tryoni var.). Nomen nudum.
     major Tryon, 1927: 195 (dorsalis var.). Nomen nudum.
     solani Perkins & May, 1949: 14 (Strumeta). Australia (Qld).
     dorsalis, Tryon, 1927, not Hendel, 1912, misid.
caledonensis Drew, 1989: 76. New Caledonia (New Caledonia).
caliginosa Hardy, 1970: 116 (Dacus). PNG (Bismarck Arch).
carbonaria Hendel, 1927c: 61 (Dacus). PNG (Bismarck Arch).
     atramentata Hering, 1941e: 9 (Strumeta). PNG (Bismarck Arch).
cheesmanae Perkins, 1939: 27 (Apodacus; as cheesemani). PNG (PNG); Australia (Qld).
cinnamea Drew, 1989: 108. PNG (PNG).
circamusae Drew, 1989: 37. PNG (PNG).
commina Drew, 1989: 38. PNG (PNG).
confluens Drew, 1971: 35 (Dacus). PNG (Bougainville I).
     honiarae Drew, 1989: 137. Solomon Is.
Drew, 1989: 127. PNG (PNG).
consectorata Drew, 1989: 27. PNG (PNG); PNG (Bismarck Arch).
contermina Drew, 1989: 39. PNG (PNG).
contigua Drew, 1989: 40. PNG (PNG).
curreyi Drew, 1989: 53. PNG (PNG); Australia (Qld), Indonesia (Irian Jaya).
curvifera Walker, 1864: 229 (Dacus). Indonesia (Irian Jaya); PNG (Bismarck Arch, PNG).
     speculifer Walker, 1865b: 122 (Dacus). "New Guinea" [= Indonesia (Irian Jaya)].
curvipennis Froggatt, 1909: 93 (Dacus). "Fiji" [error, = New Caledonia (New Caledonia); see Drew & Hancock, 2000: 29]; Vanuatu.
dapsiles Drew, 1989: 64. PNG (PNG).
daruensis Drew, 1989: 129. PNG (PNG); Australia (Qld).
decumana Drew, 1972b: 205 (Dacus). PNG (Bougainville I); Solomon Is.
diallagma Drew, 1989: 66. PNG (PNG).
diaphana Hering, 1953: 508 (Strumeta). Indonesia (Irian Jaya).
diospyri Drew, 1989: 130. Australia (NT); Australia (Qld).
distincta Malloch, 1931k: 259 (Dacus). Western Samoa; American Samoa, Fiji, Niue, Tonga, Wallis & Futuna.
dorsalis Hendel, 1912b: 18 (Dacus). Taiwan; Hawaiian Is [immigrant], Nauru [immigrant], Northern Marianas [immigrant], Solomon Is [introduced; not established]; USA [introduced; not established], widesp. Oriental Reg.
     ferruginea Fabricius, 1794: 342 (Musca). "India orientali." [Preocc. Scopoli, 1763.]
dyscrita Drew, 1971: 63 (Dacus). PNG (Bismarck Arch).
ebenea Drew, 1971: 65 (Dacus). New Caledonia (New Caledonia).
endiandrae Perkins & May, 1949: 9 (Strumeta). Australia (Qld); Australia (NSW), PNG (PNG).
enochra Drew, 1972b: 207 (Dacus). PNG (Bougainville I); Solomon Is.
epicharis Hardy, 1970: 119 (Dacus). PNG (Bismarck Arch); Solomon Is.
erubescentis Drew & Hancock in Drew et al., 1981: 64 (Dacus). Australia (Qld); PNG (PNG).
exspoliata Hering, 1941d: 54 (Strumeta). PNG (PNG).
facialis Coquillett, 1909b: 12 (Dacus). Tonga.
     tongensis Froggatt, 1911: 870 (Dacus). Tonga.
     fascialis, error for facialis.
fagraea Tryon, 1927: 188 (Chaetodacus). Australia (Qld).
fergussonensis Drew, 1989: 56. PNG (PNG).
finitima Drew, 1989: 41. PNG (PNG).
frauenfeldi Schiner, 1868: 262 (Dacus). Solomon Is; Australia (Qld), Belau, Kiribati (Gilbert Is), Marshall Is, Micronesia, Nauru, Northern Marianas, PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Indonesia, Malaysia (Pen), Singapore.
     albistrigatus de Meijere, 1911b: 377 (Dacus). Indonesia (Java).
froggatti Bezzi, 1928: 101 (Chaetodacus). Not given [Solomon Is]; ?Indonesia (Maluku), PNG (Bougainville I, PNG);.
     moluccensis, Hardy & Adachi, 1954, not Perkins, 1939, misid.
     zonatus, Froggatt, 1909, not Saunders, 1842, misid.
fuliginis Drew & Hancock in Drew et al., 1981: 66 (Dacus). Australia (Qld); PNG (PNG).
fulvicauda Perkins, 1939: 13 (Strumeta). PNG (PNG).
furfurosa Drew, 1989: 82. PNG (PNG).
furvescens Drew, 1989: 135. PNG (PNG); Solomon Is.
furvilineata Drew, 1989: 57. PNG (PNG); Australia (Qld).
fuscohumeralis White & Evenhuis, 1999: 492. Indonesia (Irian Jaya).
grandifascia White & Evenhuis, 1999: 493. Indonesia (Irian Jaya).
halfordiae Tryon, 1927: 190 (Chaetodacus). Australia (Qld); Australia (NSW).
     gurneyi Perkins, 1934: 41 (Chaetodacus). Australia (NSW & Qld).
hispidula May, 1958: 301 (Strumeta). Australia (Qld).
hollingsworthi Drew & Romig, 2001: 124. Solomon Is.
humilis Drew & Hancock in Drew et al., 1981: 68 (Dacus). Australia (Qld).
incisus Walker--not Australasian/Oceanian.
inconstans Drew, 1989: 28. PNG (PNG).
indecora Drew, 1971: 37 (Dacus). PNG (Bismarck Arch).
ismayi Drew, 1989: 138. PNG (Bismarck Arch).
kelaena Drew, 1989: 95. PNG (PNG).
kirki Froggatt, 1911: 871 (Dacus). Not given [?Fiji]; American Samoa, French Polynesia (Austral Is, Society Is), Niue, Tonga, Wallis & Futuna, Western Samoa.
kraussi Hardy, 1951b: 156 (Dacus). Australia (Qld).
lampabilis Drew, 1971: 67( Dacus). PNG (Bismarck Arch).
laticauda Hardy, 1950b: 87 (Dacus). Australia (Qld).
     fuscatus Perkins & May, 1949: 5 (Strumeta). Australia (Qld). [Preocc. Wiedemann, 1819.]
laticosta Drew, 1989: 30. PNG (PNG); PNG (Bismarch Arch).
latifrons Hendel, 1915: 425 (Chaetodacus). Taiwan; Hawaiian Is [immigrant]; Oriental Reg.
latilineata Drew, 1989: 79. PNG (PNG).
latissima Drew, 1989: 47. PNG (PNG).
lineata Perkins, 1939: 16 (Strumeta). PNG (PNG); Australia (Qld).
longicornis Macquart, 1835: 452, 453 (Bactrocera). Solomon Is; PNG (Bismarck Arch, Bougainville I).
     denigratus Drew, 1971: 61 (Dacus). PNG (Bismarck Arch).
longicornis Guérin-Méneville, 1838: 300. Australia (NSW). [Preocc. Macquart, 1835; no n.n. proposed at this time.]
luteola Malloch, 1931k: 262 (Dacus). French Polynesia (Society Is); French Polynesia (Tuamotu Arch).
     incertus Malloch, 1938a: 113 (Dacus). French Polynesia (Tuamotu Arch).
maculigera Doleschall, 1858: 122. Indonesia (Maluku).
manskii Perkins & May, 1949: 3 (Strumeta). Australia (Qld).
mayi Hardy, 1951b: 161 (Dacus; n. n. for bilineata Perkins & May). Australia (Qld).
     bilineata Perkins & May, 1949: 7 (Strumeta). Australia (Qld). [Previously considered a junior homonym in Dacus; mayi valid under I.C.Z.N. Code Art. 59b.]
melanogaster Drew, 1989: 143. Solomon Is; PNG (Bougainville I).
melanota Coquillett, 1909b: 13 (Dacus). Cook Is.
     rarotongae Froggatt, 1911: 872 (Dacus). Cook Is.
melas Perkins & May, 1949: 12 (Strumeta). Australia (Qld).
mendosa May, 1958: 303 (Strumeta). Australia (Qld); Australia (NT).
mimulus Drew, 1989: 68. PNG (PNG).
moluccensis Perkins, 1939: 17 (Strumeta). Indonesia (Maluku); PNG (Bismarck Arch, Bougainville I, PNG).
morobiensis Drew, 1989: 58. PNG (PNG).
morula Drew, 1989: 146. Solomon Is.
mucronis Drew, 1971: 70 (Dacus). New Caledonia (New Caledonia).
musae Tryon, 1927: 197 (Chaetodacus). Australia (Qld); PNG (?Bismarck Arch, PNG), ?Solomon Is.
     musa Tryon, 1927: 187 (Chaetodacus tryoni var.). Nomen nudum.
     dorsopicta Tryon, 1927: 198 (Chaetodacus musae var.). Australia (Qld).
mutabilis May, 1952: 6 (Strumeta). Australia (Qld).
naucleae Drew & Romig, 2001: 126. Solomon Is.
neocheesmanae Drew, 1989: 147. PNG (PNG).
neohumeralis Hardy, 1951b: 169 (Dacus tryoni var.; n.n. for humeralis Perkins). Australia (Qld); Australia (NSW), PNG (PNG).
     humeralis Perkins, 1934: 42 (Chaetodacus). Australia (Qld). [Previously considered a junior secondary homonym in Dacus; neohumeralis valid under I.C.Z.N. Code Art. 59b.]
neonigrita Drew, 1989: 148 (n.n. for nigritus Drew). PNG (Bismarck Arch); PNG (Bougainville I, PNG), Solomon Is.
     nigritus Drew, 1971: 75 (Dacus). PNG (Bismarck Arch). [Preocc. Hardy, 1955.]
     neonigritus Hardy in Hardy & Foote, 1989: 505 (n.n. for nigritus Drew). PNG (Bismarck Arch). [Preocc. Drew, 1989.]
nigella Drew, 1968b: 78 (Strumeta). PNG (PNG).
nigrescens Drew, 1968a: 23 (Asiadacus). PNG (Bismarck Arch); PNG (Bougainville I).
nigrescentis Drew, 1971: 72 (Dacus). PNG (Bismarck Arch); PNG (Bougainville I, PNG).
nigroscutata White & Evenhuis, 1999: 499. Indonesia (Irian Jaya).
nigrovittata Drew, 1989: 83. PNG (PNG); Australia (Qld).
notatagena May, 1953: 337 (Strumeta). Australia (Qld).
obfuscata Drew, 1989: 84. PNG (PNG).
oblineata Drew, 1989: 59. PNG (PNG).
obliqua Malloch, 1939d: 238 (Dacus). PNG (Bismarck Arch); PNG (Bougainville I).
obliquevenosa Drew & Romig, 2001: 129. Solomon Is.
obscura Malloch, 1931: 264 (Dacus). Western Samoa; American Samoa, Niue, Tonga.
occipitalis Bezzi--not Australasian/Oceanian.
ochromarginis Drew, 1971: 40 (Dacus). PNG (Bismarck Arch).
ochrosiae Malloch, 1942: 201 (Dacus). Guam; Hawaiian Is [introduced; ?established], Northern Marianas.
opiliae Drew & Hardy, 1981: 131 (Dacus). Australia (NT); Australia (Qld, WA).
pallida Perkins & May, 1949: 10 (Strumeta). Australia (Qld); Australia (NT).
papayae Drew & Hancock, 1994: 48. Malaysia (Pen); Australia (Qld) [introduced], Indonesia (Irian Jaya, Maluku), PNG (PNG); Indonesia (Nusa Tenggara, Sulawesi), Christmas I (Indian Ocean), Thailand.
     conformis Doleschall, 1858: 122. Indonesia (Maluku). [Preocc. Walker, 1856.]
parabarringtoniae Drew & Hancock in Drew et al., 1999: 2. Australia (Qld).
parafrauenfeldi Drew, 1989: 77. Australia (NT).
parafroggatti Drew & Romig, 2001: 130. Solomon Is.
paramusae Drew, 1989: 48. PNG (PNG).
passiflorae Froggatt, 1911: 870 (Dacus). Fiji; Niue, Tuvalu, Wallis & Futuna.
pectoralis Walker, 1859a: 114 (Dacus). Indonesia (Maluku); Indonesia (Java).
pedestris Bezzi, 1914: 322 (Chaetodacus). Philippines; ?Indonesia (Irian Jaya); SE Oriental Reg.
peneobscura Drew & Romig, 2001: 131. Vanuatu.
peninsularis Drew & Hancock in Drew et al., 1981: 70 (Dacus). Australia (Qld); PNG (PNG).
pepisalae Froggatt, 1911: 869 (Dacus). Solomon Is; PNG (Bougainville I).
perfusca Aubertin in Aubertin & Cheesman, 1929: 173 (Strumeta). French Polynesia (Marquesas).
perkinsi Drew & Hancock in Drew et al., 1981: 72 (Dacus). Australia (Qld).
phaea Drew, 1971: 77 (Dacus). PNG (Bismarck Arch); PNG (PNG).
phaleriae May, 1956: 159 (Strumeta). Australia (Qld).
picea Drew, 1972b: 208 (Dacus). PNG (Bougainville I); Solomon Is.
pisinna Drew, 1989: 155. PNG (PNG).
popondettiensis Drew, 1989: 85. PNG (PNG).
prolixa Drew, 1989: 93. PNG (PNG).
pseudodistincta Drew, 1971: 79 (Dacus). PNG (Bismarck Arch); Australia (Qld), PNG (PNG).
psidii Froggatt, 1899: 500 (Tephritis). New Caledonia (New Caledonia).
     virgatus Coquillett, 1909b: 13 (Dacus). Tonga.
     ornatissimus Froggatt, 1909: 93 (Dacus). New Caledonia (New Caledonia).
pulchra Tryon, 1927: 206. Australia (Qld).
quadrata May, 1963: 530 (Strumeta). Australia (Qld); PNG (Bismarck Arch, PNG).
quadrisetosa Bezzi, 1928: 102 (Chaetodacus). Vanuatu.
     varipes Malloch, 1939d: 240 (Dacus). Solomon Is.
Drew, 1989: 109. PNG (PNG).
reclinata Drew, 1989: 157. PNG (Bougainville I).
recurrens Hering, 1941c: 47 (Strumeta). PNG (PNG).
redunca Drew, 1971: 82 (Dacus). Vanuatu; Australia (Qld), PNG (Bougainville I, PNG), Solomon Is.
repanda Drew, 1989: 33. PNG (PNG); Australia (Qld).
resima Drew, 1971: 85 (Dacus). PNG (PNG); Australia (Qld).
retrorsa Drew, 1989: 158. PNG (PNG).
rhabdota Drew, 1989: 61. PNG (PNG).
robertsi Drew, 1989: 42. PNG (PNG).
robiginosa May, 1958: 305 (Strumeta). Australia (Qld).
romigae Drew & Hancock in Drew et al., 1981: 75 (Dacus). Australia (Qld).
rufescens May, 1967: 81 (Strumeta). Australia (Qld).
rufofuscula Drew & Hancock in Drew et al., 1981: 77 (Dacus). Australia (Qld).
russeola Drew & Hancock in Drew et al., 1981: 80 (Dacus). Australia (Qld).
rutila Hering, 1941c: 45 (Strumeta). PNG (PNG); Indonesia (Irian Jaya).
samoae Drew, 1989: 147. Western Samoa.
seguyi Hering, 1939a: 165 (Strumeta). Indonesia (Irian Jaya); PNG (Bismarck Arch, PNG).
     peculiaris Malloch, 1939d: 235 (Dacus). PNG (Bismarck Arch).
setinervis Malloch, 1938a: 112 (Dacus). Pitcairn Is.
silvicola May, 1962: 68 (Strumeta). Australia (Qld).
simulata Malloch, 1939d: 241 (Dacus). Solomon Is; PNG (Bougainville I).
speewahensis Fay & Hancock in Huxam et al., 2006: 34. Australia (Qld).
strigata Perkins, 1934: 43 (Chaetodacus). Australia (Qld); Australia (Vic).
tenuifascia May, 1965: 64 (Strumeta). Australia (NT); Australia (WA).
terminaliae Drew, 1989: 35. PNG (PNG).
thistletoni Drew, 1989: 163. PNG (PNG).
tinomiscii Drew, 1989: 43. PNG (PNG).
torresae Huxham & Hancock in Huxam et al., 2006: 35. PNG (PNG); Australia (Qld).
tortuosa White & Evenhuis, 1999: 503. Indonesia (Irian Jaya).
trifaria Drew, 1971: 87 (Dacus). PNG (Bismarck Arch); Australia (Qld).
trilineola Drew, 1989: 147 (n.n. for triseriatus Drew). Vanuatu; New Caledonia (Loyalty Is, New Caledonia).
     triseriatus Drew, 1971: 90 (Dacus). Vanuatu. [Preocc. Curran, 1927.]
     distotriseriata Hardy in Hardy & Foote, 1989: 504 (n.n. for triseriatus Drew).
trivialis Drew, 1971: 93 (Dacus). PNG (PNG); Australia (Qld), Indonesia (Irian Jaya).
tryoni Froggatt, 1897: 410 (Tephritis). Australia (Qld); Australia (NSW, Vic), Easter I, French Polynesia (Austral Is, Society Is), Lord Howe I, New Caledonia (New Caledonia), PNG (PNG).
     juglandis Tryon, 1927: 188 (Chaetodacus tryoni var.). Australia (Qld).
     sarcocephali Tryon, 1927: 188 (Chaetodacus tryoni var.). Australia (Qld).
turneri Drew, 1989: 112. PNG (PNG); Australia (Qld), Solomon Is.
umbrosa Fabricius, 1805: 274 (Dacus). Indonesia (Sumatra); New Caledonia (New Caledonia), PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is, Vanuatu; Christmas I (Indian Ocean), widesp. E Oriental Reg.
     frenchi Froggatt, 1909: 22 (Dacus). New Caledonia (New Caledonia).
unifasciata Malloch, 1939d: 233 (Dacus). Solomon Is.
unilineata Drew, 1989: 166. PNG (PNG).
unipunctata Malloch, 1939d: 245 (Dacus). Solomon Is.
unistriata Drew, 1971: 96 (Dacus). PNG (Bismarck Arch); PNG (PNG).
unitaeniola Drew & Romig, 2001: 137. Solomon Is.
ustulata Drew, 1989: 86. PNG (PNG).
vulgaris Drew, 1971: 99 (Dacus). PNG (PNG); Australia (Qld).
yorkensis Drew & Hancock in Drew et al., 1999: 5. Australia (Qld).
zonata Saunders, 1842: 61 (Dasyneura). India; Indonesia (Maluku); Reunion I, widesp. Oriental Reg.

Subgenus BULLADACUS Drew & Hancock

BULLADACUS Drew & Hancock, 1995: 9. Type species: Bactrocera gnetum Drew & Hancock, 1995, orig. des.
aceraglans White & Evenhuis, 1999: 506. PNG (PNG).
aceromata White & Evenhuis, 1999: 509. PNG (PNG).
aenigmatica Malloch, 1931k: 261 (Dacus). Western Samoa.
bullata Drew, 1989: 124. PNG (PNG).
eximia Drew, 1989: 132. PNG (PNG).
flavinotus May, 1958: 293 (Afrodacus). Australia (Qld).
     furvus May, 1958: 294 (Afrodacus). Australia (Qld).
     neotigrina Drew & Hancock in Drew et al., 1999: 7. Australia (Qld).
Drew & Hancock, 1995: 9. Fiji.
obtrullata White & Evenhuis, 1999: 510. Indonesia (Irian Jaya).
pacificae Drew & Romig, 2001: 138. Solomon Is.
penefurva Drew, 1989: 151. PNG (PNG); Solomon Is.
tigrina May, 1953: 339 (Afrodacus). Australia (Qld).
warisensis White & Evenhuis, 1999: 512. Indonesia (Irian Jaya).


DIPLODACUS May, 1952: 12 (as gen.). Type species: Dacus signatifer Tryon, 1927, orig. des.
signatifera Tryon, 1927: 210 (Dacus). Australia (Qld); Australia (NT).

Subgenus GYMNODACUS Munro

GYMNODACUS Munro, 1938: 117 (Dacus subg.). Type species: Dacus mesomelas Bezzi, 1908, orig. des.
calophylli Perkins & May, 1949: 16 (Asiadacus). Australia (Qld); Belau; Malaysia (Pen), Singapore.
hastigerina Hardy, 1954b: 19 (Dacus). PNG (Bismarck Arch).
petila Drew, 1989: 169. PNG (PNG).


HEMINOTODACUS Drew, 1989: 15, 187. Type species: Bactrocera dissidens Drew, 1989, orig. des.
dissidens Drew, 1989: 187. PNG (PNG).


HEMIPARATRIDACUS Drew, 1989: 15, 188. Type species: Bactrocera abdoaurantiaca Drew, 1989, orig. des.
abdoaurantiaca Drew, 1989: 188. PNG (PNG).


HEMISURSTYLUS Drew, 1989: 13, 178. Type species: Bactrocera melanoscuta Drew, 1989, orig. des.
melanoscuta Drew, 1989: 178. PNG (PNG).


HEMIZEUGODACUS Hardy, 1951b: 131 (Dacus subg.). Type species: Dacus aglaiae Hardy, 1951, orig. des.
NEOZEUGODACUS May, 1952: 10 (as gen.). Type species: Neozeugodacus aureus May, 1952, orig. des.
abdominigra Drew, 1989: 179. PNG (PNG).
aglaiae Hardy, 1951b: 132 (Dacus). Australia (Qld).
aurea May, 1952: 10 (Neozeugodacus). Australia (Qld).
buinensis Drew, 1989: 123. PNG (Bougainville I).
ektoalangiae Drew & Hancock in Drew et al., 1999: 9. Australia (Qld).


HETERODACULUS Hardy, 1951b: 134 (Dacus subg.). Type species: Dacus visendus Hardy, 1951, orig. des.
Drew, 1989: 172. PNG (PNG).
mesonotochra Drew, 1989: 172. PNG (PNG).
toxopeusi Hering, 1953: 510 (Daculus). Indonesia (Irian Jaya).
visenda Hardy, 1951b: 135 (Dacus). Australia (Qld); PNG (PNG).

Subgenus JAVADACUS Hardy

JAVADACUS Hardy, 1983a: 26. Type species: Dacus montanus Hardy, 1983, orig. des.
aberrans Hardy, 1951b: 118 (Afrodacus). Australia (Qld).
     mesoniger May, 1952: 8 (Afrodacus). Australia (Qld).
melanothoracica Drew, 1989: 190. Australia (Qld).
unirufa Drew, 1989: 191. PNG (PNG); Australia (Qld).

Subgenus MELANODACUS Perkins

MELANODACUS Perkins, 1937: 57 (as gen.). Type species: Dacus niger Tryon, 1927, orig. des.
nigra Tryon, 1927: 211 (Dacus). Australia (Qld).
satanellus Hering, 1941c: 48 (Melanodacus). PNG (PNG).


NIUGINIDACUS Drew, 1989: 15, 192. Type species: Bactrocera singularis Drew, 1989, orig. des.
singularis Drew, 1989: 192. PNG (PNG).

Subgenus NOTODACUS Perkins

NOTODACUS Perkins, 1937: 56 (as gen.). Type species: Tephritis xanthodes Broun, 1904, orig. des.
neoxanthodes Drew & Romig, 2001: 142. Vanuatu.
paraxanthodes Drew & Hancock, 1995: 10. New Caledonia (New Caledonia); ?Vanuatu, Western Samoa.
xanthodes Broun, 1904a: 306 [1905a: 3] (Tephrites) [1905b: 327 (Dacus)]. Cook Is; American Samoa, Fiji, Nauru, Niue, Tonga, Wallis & Futuna, Western Samoa.

Subgenus PAPUODACUS Drew

PAPUODACUS Drew, 1972a: 13 (Dacus subg.). Type species: Dacus pallescentis Drew, 1971 [preocc., = Bactrocera neopallescentis Drew, 1989], orig. des.
neopallescentis Drew, 1989: 193 (n.n. for pallescentis Drew). PNG (PNG)
     pallescentis Drew, 1971: 44. PNG (PNG) (Dacus). [Preocc. Hardy, 1955.]
     neopallescentis Hardy, 1989: 507 (n.n. for pallescentis Drew). [Preocc. Drew, 1989.]

Subgenus PARADACUS Perkins

PARADACUS Perkins, 1938: 143 (as gen.). Type species: Paradacus fulvipes Perkins, 1938, orig. des.
areolata Walker, 1861d: 295 (Dacus). Indonesia (Maluku).
aurantiventer Drew, 1989: 194. PNG (PNG).
citroides Drew, 1989: 195. PNG (PNG).
decipiens Drew, 1972a: 130 (Dacus). PNG (Bismarck Arch).
magnicauda White & Evenhuis, 1999: 517. Indonesia (Irian Jaya).
perplexa Walker, 1861e: 14 (Dacus). Indonesia (Maluku).
     implexus Perkins, 1939: 34 (Dacus). Nomen nudum.
urens White in White & Evenhuis, 1999: 518. Indonesia (Maluku).

Subgenus PARATRIDACUS Shiraki

PARATRIDACUS Shiraki, 1933: 109 (as gen.). Type species: Dacus yayeyamanus Matsumura, 1916 [= Dacus expandens Walker, 1859], orig. des.
alampeta Drew, 1989: 196. PNG (PNG).
atrisetosa Perkins, 1939: 29 (Zeugodacus). PNG (PNG).
     papuaensis Malloch, 1939f: 412 (Dacus). PNG (PNG).
     rubidus May, 1958: 297 (Melanodacus). PNG (PNG).
banneri White in White & Evenhuis, 1999: 520. Indonesia (Maluku).
coracina Drew, 1971: 46 (Dacus). PNG (PNG).
expandens Walker, 1859a: 114 (Dacus). Indonesia (Maluku); Australia (Qld), PNG (PNG); India, Indonesia, Japan, Malaysia (Pen), Philippines, Ryukyu Is, Sri Lanka.
     yayeyamanus Matsumura, 1916: 413 (Dacus). Japan.
     yayemanus, error for yayeyamanus.
mesonotaitha Drew, 1989: 199. PNG (PNG).
unichromata Drew, 1989: 200. PNG (PNG).


PARAZEUGODACUS Shiraki, 1933: 107 (as gen.). Type species: Parazeugodacus matsumurai Shiraki, 1933, orig. des.
matsumurai Shiraki, 1933: 107 (Parazeugodacus). Bonin Is.
     boninensis Hardy & Adachi, 1956: 12 (Dacus). Bonin Is.


QUEENSLADACUS Drew, 1989: 14, 184. Type species: Psilodacus exigua May, 1958, orig. des.
exigua May, 1958: 300 (Psilodacus). Australia (Qld).


SEMICALLANTRA Drew, 1989: 17, 264 (Dacus subg.). Type species: Dacus aquilus Drew, 1989, orig. des.
aquilus Drew, 1989: 264 (Dacus). PNG (PNG).
memnonius Drew, 1989: 265 (Dacus). PNG (PNG).
nigriculus Drew, 1989: 266 (Dacus). PNG (PNG).

Subgenus SINODACUS Zia

SINODACUS Zia, 1936: 157 (as gen.). Type species: Sinodacus hochii Zia 1936, orig. des.
PACIFODACUS Drew, 1972a: 12 (Dacus subg.). Type species: Asiadacus triangularis Drew, 1968, orig. des.
abdopallescens Drew, 1971: 31 (Dacus). PNG (PNG).
aneuvittata Drew, 1971: 33 (Dacus). New Caledonia (New Caledonia).
angusticostata Drew, 1989: 202. PNG (PNG).
buvittata Drew, 1989: 203. PNG (PNG).
emarginata Perkins, 1939: 24 (Neodacus). PNG (PNG).
hamaceki Drew & Romig, 2001: 143. Solomon Is.
paulula Drew, 1989: 204. PNG (PNG).
perpusilla Drew, 1971: 42 (Dacus). New Caledonia (New Caledonia).
salamander Drew & Hancock in Drew et al., 1981: 51 (Dacus). Australia (Qld).
sepikae Drew, 1989: 206. PNG (PNG).
strigifinis Walker, 1861d: 295 (Dacus). Indonesia (Maluku); Australia (Qld), PNG (PNG), Solomon Is.
     lanceolatus Perkins, 1939: 22 (Neodacus). PNG (PNG).
     albolateralis Malloch, 1939f: 413 (Dacus). PNG (PNG).
     atritus May, 1962: 65 (Asiadacus strigifinis ssp.). PNG (PNG).
surrufula Drew, 1989: 208. PNG (PNG).
triangularis Drew, 1968a: 21 (Asiadacus). PNG (Bismarck Arch); PNG (Bougainville I).
univittata Drew, 1972b: 189 (Dacus). PNG (Bougainville I).

Subgenus TETRADACUS Miyake

TETRADACUS Miyake, 1919: 95 (Dacus subg.). Type species: Dacus tsunoenis Miyake, 1919, orig. des.
POLISTOMIMETES Enderlein, 1920a: 358 (as gen.). Type species: Polistomimetes minax Enderlein, 1920, orig. des.
decurtans May, 1965: 61 (Daculus). Australia (NT); Australia (Qld, WA).
discipennis Walker, 1861d: 294 (Dacus). Indonesia (Maluku).
murrayi Perkins, 1939: 25 (Daculus). Australia (Qld).
neopagdeni Drew, 1989: 175. PNG (PNG).
pagdeni Malloch, 1939d: 243 (Dacus). Solomon Is.


TRIPETIDACUS Drew, 1989: 13, 177. Type species: Bactrocera invisitata Drew, 1989, orig. des.
invisitata Drew, 1989: 177. PNG (PNG).

Subgenus ZEUGODACUS Hendel

ZEUGODACUS Hendel, 1927a: 26 (Dacus subg.). Type species: Dacus caudatus Fabricius, 1805, orig. des.
abdoangusta Drew, 1972b: 191 (Dacus). PNG (Bougainville I); Indonesia (Sulawesi).
amoena Drew, 1972b: 192 (Dacus). PNG (Bougainville I).
anchitrichota Drew, 1989: 211. PNG (PNG).
buruensis White in White & Evenhuis, 1999: 524. Indonesia (Maluku).
brachus Drew, 1972b: 194 (Dacus). PNG (PNG).
chorista May, 1962: 72 (Zeugodacus). Australia (Qld); PNG (PNG).
curcubitae Coquillett, 1899b: 129 (Dacus). Hawaiian Is [introduced]; Australia (NT), Guam, Indonesia (Irian Jaya), Nauru, Northern Marianas, PNG (Bismarck Arch, Bougainville I, PNG), Solomon Is; Mauritius, widesp. Oriental Reg.
curta Drew, 1972b: 195 (Dacus). PNG (Bismarck Arch).
daula Drew, 1989: 214. PNG (PNG).
emittens Walker, 1860a: 152 (Dacus). Indonesia (Sulawesi); Indonesia (Maluku); Indonesia (Nusa Tenggara).
chrysotoxus Hendel, 1912b: 24 (Dacus). Indonesia (Maluku).
     chrysotaxus, error for chrysotoxus.
exornata Hering, 1941d: 55 (Zeugodacus). Indonesia (Maluku).
fallacis Drew, 1972b: 196 (Dacus). Australia (Qld).
fulvifacies Perkins, 1939: 32 (Zeugodacus). New Caledonia (New Caledonia).
fulvoabdominalis White & Evenhuis, 1999: 528. Indonesia (Irian Jaya).
fuscipennula Drew & Romig, 2001: 146. Solomon Is.
gracilis Drew, 1972b: 198 (Dacus). Vanuatu.
hoedi White in White & Evenhuis, 1999: 529. Indonesia (Irian Jaya).
macrovittata Drew, 1989: 217. PNG (PNG); Australia (Qld).
pura White in White & Evenhuis, 1999: 533. Indonesia (Irian Jaya).
reflexa Drew, 1971: 101 (Dacus). PNG (Bismarck Arch).
sandaracina Drew, 1989: 218. PNG (PNG).
trichota May, 1962: 74 (Zeugodacus). PNG (PNG); PNG (Bismarck Arch).
ubiquita Hardy, 1973: 71 (Dacus). Philippines; PNG (Bismarck Arch).
unilateralis Drew, 1989: 221. PNG (PNG).

Nomen nudum in Bactrocera Macquart

nigrofasciata Tryon, 1927: 199 (Dacus).

Genus DACUS Fabricius

DACUS Fabricius, 1805: 272. Type species: Dacus armatus Fabricius, 1805, des. Hendel, 1927a: 24.

Subgenus CALLANTRA Walker

CALLANTRA Walker, 1860a: 153 (as gen.). Type species: Callantra smieroides Walker, 1860, orig. des.
ambonensis Drew & Hancock in Drew, Hancock & White, 1998: 572. Indonesia (Maluku).
axanus Hering, 1938: 410 (Callantra). Indonesia (Maluku); Australia (NT, Qld, WA), PNG (Bismarck Arch, PNG); Indonesia (Nusa Tenggara).
axanthinus White & Evenhuis, 1999: 536. PNG (PNG).
auricoma May, 1956: 153 (Callantra). Australia (Qld).
capillaris Drew, 1972b: 185 (Callantra). PNG (Bougainville I).
discors Drew, 1989: 242. PNG (PNG).
impar Drew, 1989: 243. PNG (PNG).
longicornis Wiedemann, 1830a: 524. Indonesia (Java); Indonesia (Maluku).
mayi Drew, 1972b: 187 (Callantra). PNG (PNG).
melanohumeralis Drew, 1989: 244. PNG (PNG).
petioliformus May, 1956: 151 (Callantra). Australia (Qld).
pusillus May, 1965: 58 (Callantra). Australia (Qld).
smieroides Walker--not Australasian/Oceanian.
solomonensis Malloch, 1939d: 236. Solomon Is; PNG (Bougainville I).
unicolor Hendel, 1927c: 58 (Callantra). PNG (Bismarck Arch).
vespoides Doleschall, 1858: 123 (Bactrocera). Indonesia (Maluku).
wallacei White in Drew, Hancock & White, 1998: 639. Indonesia (Maluku).

Subgenus DACUS Fabricius

DACUS Fabricius, 1805: 272 (as gen.). Type species: Dacus armatus Fabricius, 1805, des. Hendel, 1927a: 24.
TRIDACUS Bezzi, 1915: 86 (as gen.). Type species: Dacus armatus Fabricius, 1805, des. Collart, 1935: 9.
NEODACUS Perkins, 1937: 58 (as gen.). Type species: Neodacus newmani Perkins, 1937, orig. des.
absonifacies May, 1956: 156 (Polistomimetes). Australia (Qld).
alarifumidus Drew, 1989: 252. PNG (PNG).
alulapictus Drew, 1989: 252. PNG (PNG).
badius Drew, 1989: 253. PNG (PNG).
bellulus Drew & Hancock in Drew et al., 1981: 50. Australia (Qld); Australia (NT), PNG (PNG).
concolor Drew, 1989: 255. Australia (Qld).
newmani Perkins, 1937: 58 (Neodacus). Australia (WA); widesp. Australia.
secamonae Drew, 1989: 257. Australia (NT); Australia (Qld).
signatifrons May, 1956: 155 (Neodacus). Australia (Qld).
taui Drew & Romig, 2001: 148. Solomon Is.

Subgenus DIDACUS Collart

DIDACUS Collart, 1935: 33 . Type species: Dacus ciliatus Loew, 1862, orig. des.
aequalis Coquillett, 1909a: 794. Australia (NSW); Australia (Qld).
dissimilis Drew, 1989: 260. PNG (PNG).
hardyi Drew, 1979: 74. Australia (NT); Australia (Qld).
maprikensis Drew, 1989: 262. PNG (PNG).
palmerensis Drew, 1989: 263. Australia (Qld).


PARACALLANTRA Hendel, 1927c: 59. Type species: Paracallantra vespiformis Hendel, 1927, orig. des.
vespiformis Hendel, 1927c: 60. PNG (Bismarck Arch).

Unplaced species of Dacinae

ornatissimus Froggatt, 1909: 101 (Dacus). New Caledonia (New Caledonia).



ACANTHONEVRA Macquart, 1843: 377(220). Type species: Acanthonevra fuscipennis Macquart, 1843, orig. des.
RIOXOPTILONA Hendel, 1914c: 78. Type species: Trypeta vaga Wiedemann, 1830, orig. des.
ACANTHONEURA, error for Acanthonevra.
ceramensis de Meijere, 1913a: 61 (dunlopi Wulp var.). Indonesia (Maluku); Malaysia (Sabah).

Genus ACANTHONEVROIDES Permkam & Hancock

ACANTHONEVROIDES Permkam & Hancock, 1995b: 1053. Type species: Urophora bicolor Macquart, 1855, orig. des.
basalis Walker, 1853: 380 (Trypeta). "Brazil' [error, not Neotropical; see Norrbom et al., 1998: 67]; Australia (SA).
     bicolor Macquart, 1855: 144(124) (Urophora). Australia (SA).
jarvisi Tryon, 1927: 221 (Rioxa). Australia (Qld).
mayi Permkam & Hancock, 1995b: 1056. Australia (Qld).
nigriventris Malloch, 1939f: 432 (Acanthonevra). Australia (NSW).
variegatus Permkam & Hancock, 1995b: 1061. Australia (WA); Australia (NT).


ALLOEOMYIA Hardy, 1986b: 29. Type species: Alloeomyia flavida Hardy, 1986, orig. des.
flavida Hardy, 1986b: 30. PNG (PNG).


ANCHIACANTHONEVRA Hardy, 1986b: 31. Type species: Anchiacanthonevra maculipennis Hardy, 1986, orig. des.
maculipennis Hardy, 1986b: 31. PNG (Bismarck Arch).

Genus ARIDONEVRA Permkam & Hancock

ARIDONEVRA Permkam & Hancock, 1995b: 1063. Type species: Aridonevra cunnamullae Permakam & Hancock, 1995, orig. des.
cunnamullae Permkam & Hancock, 1995b: 1064. Australia (Qld).

Genus AUSTRONEVRA Permkam & Hancock

AUSTRONEVRA Permkam & Hancock, 1995b: 1065. Type species: Acanthoneura australina Hendel, 1928, orig. des.
australina Hendel, 1928b: 359 (Acanthoneura). Australia (Qld).
bimaculata Permkam & Hancock, 1995b: 1067. Australia (Qld).
irwini Norrbom & Hancock, 2004: 68. New Caledonia (New Caledonia).

Genus AUSTRORIOXA Permkam & Hancock

AUSTRORIOXA Permkam & Hancock, 1995b: 1069. Type species: Acanthoneura acidiomorpha Hendel, 1928, orig. des.
acidiomorpha Hendel, 1928b: 360 (Acanthoneura). Australia (NSW); Australia (Qld).
     acidomorpha, error for acidiomorpha.

Genus BULOLOA Hardy

BULOLOA Hardy, 1986b: 32. Type species: Buloloa spinicosta Hardy, 1986, orig. des.
spinicosta Hardy, 1986b: 33. PNG (PNG).


CHEESMANOMYIA Malloch, 1939f: 419. Type species: Cheesmanomyia unica Malloch, 1939 [= Rioxa nigra de Meijere, 1906], orig. des.
nigra de Meijere, 1906b: 95 (Rioxa). Indonesia (Irian Jaya); PNG (PNG).
     unica Malloch, 1939f: 420. Indonesia (Irian Jaya).

Genus CLUSIOSOMA Malloch

CLUSIOSOMA Malloch, 1926d: 547. Type species: Clusiosoma semifusca Malloch, 1926 [= Acanthoneura minuta de Meijere, 1913], orig. des.

Subgenus CLUSIOSOMA Malloch

CLUSIOSOMA Malloch, 1926d: 547 (as gen.). Type species: Clusiosoma semifusca Malloch, 1926 [= Acanthoneura minuta de Meijere, 1913], orig. des.
centrale Malloch, 1939f: 426. PNG (PNG).
dami Hardy, 1986b: 40. PNG (Bismarck Arch); ?PNG (PNG).
daruense Hardy, 1986b: 41. PNG (PNG).
laterale Walker, 1865b: 123 (Dacus). "New Guinea" [= Indonesia (Irian Jaya)]; Australia (Qld), PNG (PNG).
     biseriata Malloch, 1939f: 426. PNG (PNG).
mcalpinei Permkam & Hancock, 1995b: 1074. Australia Qld).
melanthe Hering, 1947a: 2. PNG (Bismarck Arch).
minutum de Meijere, 1913a: 62 (Acanthoneura). Indonesia (Irian Jaya); PNG (PNG).
nigricorne Hardy, 1986b: 45. PNG (PNG).
nigripenne Hardy, 1986b: 46. PNG (PNG).
partitum Malloch, 1939f: 425. PNG (PNG).
pleurale Malloch, 1939d: 259. Solomon Is.
pullatum Hering, 1941d: 60. PNG (PNG).
semifuscum Malloch, 1926d: 548. Australia (Qld); Australia (NT).
subpullatum Hardy, 1986b: 50. PNG (PNG).
vittiferum Walker, 1865b: 126 (Psila). "New Guinea" [= Indonesia (Irian Jaya)].


PARACLUSIOSOMA Hardy, 1986b: 52. Type species: Clusiosoma papuaense Hardy, 1986, orig. des.
papuaense Hardy, 1986b: 52. PNG (PNG); Australia (Qld).


CLUSIOSOMINA Malloch, 1939f: 426 (Clusiosoma subg.). Type species: Clusiosoma puncticeps Malloch, 1939, mon.
puncticeps Malloch, 1939f: 426 (Clusiosoma). Australia (NSW); Australia (Qld).

Genus COPIOLEPIS Enderlein

COPIOLEPIS Enderlein, 1920a: 341. Type species: Copiolepis quadrisquamosa Enerlein, 1920, orig. des.
colpopteris Permkam & Hancock, 1995b: 1081. Australia (Qld).
quadrisquamosa Enderlein, 1920a: 341. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG).

Genus DACOPSIS Hering

DACOPSIS Hering, 1944a: 2. Type species: Dacopsis dacina Hering, 1944 [= Seraca signata Walker, 1860], orig. des.
SOPHIRA, authors, not Walker, 1856, misid.
flava Edwards, 1915b: 421 (Rioxa). Indonesia (Irian Jaya); Australia (Qld), PNG (Bismarck Arch, PNG).
     apicalis Hardy, 1980b: 151. PNG (Bismarck Arch).
     picturara Hardy, 1980b: 155. PNG (PNG).
holoxantha Hering, 1941e: 21 (Sophira). PNG (Bismarck Arch); PNG (PNG); Malaysia (Sarawak).
quadripunctata Malloch, 1939d: 255 (Sophira). Solomon Is.
signata Walker, 1860a: 165 (Seraca). Indonesia (Sulawesi); Indonesia (Maluku); Philippines.
     dacina Hering, 1944a: 3. Indonesia (Maluku).


DIARRHEGMA Bezzi, 1913b: 76, 108. Type species: Dacus modestus Fabricius, 1805, orig. des.
modestum Fabricius, 1805: 278 (Dacus). India; Indonesia (Maluku); widesp. Oriental Reg.
paritii Doleschall, 1856: 412 (Tephritis). Indonesia (Maluku).


DIARRHEGMOIDES Malloch, 1939f: 437. Type species: Diarrhegmoides hastata Malloch, 1939, orig. des.
hastatus Malloch, 1939f: 437. PNG (PNG); ?Indonesia (Irian Jaya).

Genus DIRIOXA Hendel

DIRIOXA Hendel, 1928b: 353 (Rioxa subg.). Type species: Trypeta musae Froggatt, 1899 [= Trypeta pornia Walker, 1849], orig. des.
fuscipennis Hancock & Drew, 2003: 66. Vanuatu.
incerta Hardy, 1986b: 18 (Acanthonevra). Indonesia (Irian Jaya).
pornia Walker, 1849: 1039 (Trypeta). Australia (NSW); American Samoa [intercepted; not established], Australia (Qld), Fiji [intercepted; not established], French Polynesia (Society Is) [intercepted; not established], New Caledonia (New Caledonia), NZ (NZ) [intercepted; not established], Vanuatu [intercepted; not established].
     psidii Froggatt, 1899: 500 (Tephritis). New Caledonia (New Caledonia).
     musae Froggatt, 1899: 501 (Trypeta). Vanuatu [error, = Australia (Qld)].
     confusa Hardy, 1951b: 183 (Rioxa). Australia (Qld).


HERINGOMYIA Hardy, 1986b: 67. Type species: Acanthoneura longiplaga Hering, 1939, orig. des. [Preocc. Hardy, 1963.]
EMHERINGIA Hardy, 1989: 512 (n. n. for Heringomyia Hardy, 1986).
longiplaga Hering, 1939a: 174 (Acanthoneura). Indonesia (Maluku).


ENICOPTERINA Malloch, 1939g: 240. Type species: Enicopterina bivittata Malloch, 1939, orig. des.
bivittata Malloch, 1939g: 241. Fiji.

Genus ENOPLOPTERON de Meijere

ENOPLOPTERON de Meijere, 1913c: xl [1915b: 127]. Type species: Enoplopteron hieroglyphicum de Meijere, 1913, mon.
hieroglyphicum de Meijere, 1913c: xl [1915b: 127]. PNG (PNG); Indonesia (Irian Jaya), PNG (Bismarck Arch).
occulatum Hardy, 1986b: 62. PNG (PNG).
reticulatum Hardy, 1986b: 62. PNG (PNG).


GRESSITTIDIUM Hardy, 1986b: 64. Type species: Gressittidium flavicoxa Hardy, 1986, orig. des.
flavicoxa Hardy, 1986b: 64. PNG (PNG); Indonesia (Irian Jaya).


HEMICLUSIOSOMA Hardy, 1986b: 66. Type species: Hemiclusiosoma trivittatum Hardy, 1986, orig. des.
trivittatum Hardy, 1986b: 66. PNG (PNG).

Genus HEXACINIA Hendel

HEXACINIA Hendel, 1914c: 82 [1915: 459]. Type species: Acinia stellata Macquart, 1851 [preocc., = Acinia radiosa Rondani, 1868], orig. des.
punctifera Walker, 1861e: 15 (Sophira). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).
     multipunctata Malloch, 1939f: 438. PNG (PNG).
     flavipunctata Hering, 1940: 8. Indonesia (Maluku).


HEXARESTRA Hering, 1941: 18. Type species: Hexarestra juanita Hering, 1941 [= Trypeta multistriga Walker, 1858], orig. des.
HYPONEOTHEMARA Hardy, 1986b: 71. Type species: Trypeta multistriga Walker, 1859, orig. des.
formosa Malloch, 1939d: 255 (Neothemara). Solomon Is.
multistriga Walker, 1859a: 119 (Trypeta). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG); Indonesia (Sulawesi).
     juanita Hering, 1941: 19. Surinam [error, not Neotropical].
setinervis Hering, 1953: 520 (Dirioxa). PNG (PNG).

Genus LUMIRIOXA Permkam & Hancock

LUMIRIOXA Permkam & Hancock, 1995b: 1087. Type species: Rioxa araucariae Tryon, 1927, orig. des.
araucariae Tryon, 1927: 219 (Rioxa). Australia (Qld); Australia (NSW).

Genus LYRONOTUM Hering

LYRONOTUM Hering, 1941f: 112. Type species: Acanthoneura seriata de Meijere, 1915, orig. des.
seriatum de Meijere, 1915b: 125 (Acanthoneura). PNG (PNG).

Genus MICRONEVRINA Permkam & Hancock

MICRONEVRINA Permkam & Hancock, 1995b: 1089. Type species: Micronevrina apicalis Permkam & Hancock, 1995, orig. des.
apicalis Permkam & Hancock, 1995b: 1090. Australia (NSW); Australia (Qld).
breviseta Permkam & Hancock, 1995b: 1093. Australia (Qld).
gloriosa Permkam & Hancock, 1995b: 1094. Australia (Qld).
hyalina Permkam & Hancock, 1995b: 1095. Australia (Qld); Australia (NSW).
mediivitta Permkam & Hancock, 1995b: 1097. Australia (Qld); Australia (NSW).
montana Permkam & Hancock, 1995b: 1098. Australia (Qld).
setosa Permkam & Hancock, 1995b: 1102. Australia (NSW); Australia (Qld).


MIMOEUPHRANTA Hardy, 1986b: 79. Type species: Mimoeuphranta diaspora Hardy, 1986, orig. des.
diaspora Hardy, 1986b: 80. PNG (PNG); Indonesia (Irian Jaya).

Genus NEOTHEMARA Malloch

NEOTHEMARA Malloch, 1939d: 253. Type species: Rioxa formosipennis Walker, 1861, orig. des.
NESTHEMARA, error for Neothemara.
digressa Hardy, 1986b: 82. PNG (PNG).
formosipennis Walker, 1861b: 252 (Rioxa). Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).
     trigonifera Hering, 1951a: 7 (formosipennis f.). "New Guinea."


NOTHOCLUSIOSOMA Hardy, 1986b: 85. Type species: Trypanocentra vittithorax Malloch, 1939, orig. des.
vittithorax Malloch, 1939f: 429 (Trypanocentra). PNG (PNG); Indonesia (Irian Jaya).


OTHNIOCERA Hardy, 1986b: 87. Type species: Othniocera pictipennis Hardy, 1986, orig. des.
aberrans Hardy, 1986b: 88. PNG (PNG).
pallida Hardy, 1986b: 89. PNG (PNG).
pictipennis Hardy, 1986b: 90. PNG (PNG).


PAEDOHEXACINIA Hardy, 1986b: 92. Type species: Paedohexacinia flavithorax Hardy, 1986, orig. des.
clusiosomopsis Hardy, 1986b: 92. PNG (PNG); Australia (Qld).
flavithorax Hardy, 1986b: 93. PNG (PNG); Australia (Qld).


PARACHLAENA Hering, 1944a: 3. Type species: Rhacochlaena greenwoodi Bezzi, 1928, orig. des.
greenwoodi Bezzi, 1928: 110 (Rhacochlaena). Fiji.


PARAPHASCA Hardy, 1986b: 94. Type species: Paraphasca taenifera Hardy, 1986, orig. des.
taenifera Hardy, 1986b: 94. PNG (PNG).

Genus PHASCA Hering

PHASCA Hering, 1953: 518. Type species: Phasca bidens Hering, 1953 [= Helomyza ortaloides Walker, 1865], orig. des.
bicunea Hardy, 1986b: 97. PNG (PNG).
connexa Hardy, 1986b: 99. PNG (PNG); Indonesia (Irian Jaya).
maculifacies Hardy, 1986b: 100. PNG (PNG).
ortaloides Walker, 1865b: 116 (Helomyza). "New Guinea" [= Indonesia (Irian Jaya)]; PNG (PNG).
     bidens Hering, 1953: 519. Indonesia (Irian Jaya).
sedlaceki Hardy, 1986b: 102. PNG (PNG).
trifasciata Hardy, 1986b: 103. PNG (PNG); Indonesia (Irian Jaya).

Genus POLYARA Walker

POLYARA Walker, 1859a: 122. Type species: Polyara insolita Walker, 1859, orig. des.
bambusae Hardy, 1986b: 105. PNG (PNG).
insolita Walker, 1859a: 123. Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).
leptotrichosa Hardy, 1986b: 107. PNG (PNG).


PSEUDACANTHONEURA Malloch, 1939f: 434. Type species: Pseudacanthoneura septemnotata Malloch, 1939 [= Acanthoneura sexguttata de Meijere, 1913], orig. des.
aberrans Hardy, 1986b: 109. PNG (PNG).
sexguttata de Meijere, 1913b: 364 (Acanthoneura). PNG (PNG); Australia (Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch).
     septemnotata Malloch, 1939f: 434. PNG (PNG).


PSEUDONEOTHEMARA Hardy, 1986b: 111. Type species: Themarohystrix exul Curran, 1936, orig. des.
exul Curran, 1936: 27 (Themarohystrix). Solomon Is; PNG (Bismarck Arch).
repleta Walker, 1861d: 296 (Strumeta). Indonesia (Maluku).


QUASIRHABDOCHAETA Hardy, 1986b: 114. Type species: Quasirhabdochaeta singularis Hardy, 1986, orig. des.
singularis Hardy, 1986b: 114. PNG (Bismarck Arch).

Genus RABAULIA Malloch

RABAULIA Malloch, 1939d: 257. Type species: Rabaulia fascifacies Malloch, 1939, orig. des.
fascifacies Malloch, 1939d: 258. Solomon Is; PNG (Bismarck Arch).
invittata Hering, 1951a: 6. Indonesia (Irian Jaya).
nigrotibia Hering, 1941d: 61. PNG (PNG); Australia (Qld).


RABAULIOMORPHA Hardy, 1970: 122. Type species: Rabauliomorpha gibbosa Hardy, 1970, orig. des.
gibbosa Hardy, 1970: 124. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG).

Genus RIOXA Walker

RIOXA Walker, 1856a: 35. Type species: Rioxa lanceolata Walker, 1856, mon.
discalis Walker, 1861: 321 (Tetanocera). Myanmar; Soloon Is; SE Asia, Indonesia (Java, Sumatra), Malaysia (Sarawak).
     sumatrana Enderlein, 1911b: 449. Indonesia (Sumatra).
Wulp, 1880: 185 [1881: 51] (Ptilona). Indonesia (Sumatra); Indonesia (Irian Jaya); widesp. Oriental Reg.


SOPHIROPSIS Hardy, 1986b: 126. Type species: Sophiropsis improbata Hering, 1941, orig. des.
calcarata Hardy, 1986b: 128. PNG (PNG).
improbata Hering, 1941e: 22 (Termitorioxa). PNG (PNG); Indonesia (Irian Jaya).


STIGMATOMYIA Hardy, 1986b: 129. Type species: Stigmatomyia arcuata Hardy, 1986, orig. des.
arcuata Hardy, 1986b: 129. PNG (PNG).

Genus TAENIORIOXA Permkam & Hancock

TAENIORIOXA Permkam & Hancock, 1995b: 1115. Type species: Taeniorioxa quinaria Permkam & Hancock, 1995, orig. des.
quinaria Permkam & Hancock, 1995b: 1115. Australia (Qld).


TERMITORIOXA Hendel, 1928b: 351 (Rioxa subg.). Type species: Rioxa termitoxena Bezzi, 1919, mon.
KERTESZIOLA Hering, 1941c: 50. Type species: Ptilona lateralis Kertész, 1901 [= Helomyza meritoria Walker, 1864], orig. des.
RIOXENA, error for Termitorioxa.
acanthoneurides Hering, 1953: 516 (Rioxa). Indonesia (Irian Jaya).
bicalcarata Hering, 1944a: 4 (Diarrhegmoides). Australia (Qld).
cobourgensis Hancock, 1996: 88. Australia (NT).
exleyae Permkam & Hancock, 1995b: 1120. Australia (WA); Australia (NT, Qld).
flava Hardy, 1986b: 76 (Kertesziola). PNG (PNG).
inconnexa Permkam & Hancock, 1995b: 1122. Australia (NT).
laurae Permkam & Hancock, 1995b: 1124. Australia (Qld); Australia (NT, WA).
meritoria Walker, 1864: 218 (Helomyza). Indonesia (Irian Jaya); Indonesia (Maluku), PNG (PNG).
     lateralis Kertész, 1901c: 428 (Ptilona). PNG (PNG).
termitoxena Bezzi, 1919b: 1, 2. Australia (NT); Australia (Qld, WA).
testacea Hendel, 1928b: 352 (Rioxa). Australia (Qld).

Genus THEMARA Walker

THEMARA Walker, 1856a: 33. Type species: Themara ampla Walker, 1856 [= Achias maculipennis Westwood, 1848], mon.
lunifera Hering, 1938: 408. Philippines; Solomon Is.


THEMAROHYSTRIX Hendel, 1914c: 432. Type species: Themarohystrix erinaceus Hendel, 1914 [= Strumeta helomyzoides Walker, 1864], orig. des.
alpina Hardy, 1986b: 143. PNG (PNG).
bivittata Hardy, 1986b: 145. Indonesia (Irian Jaya).
flaviceps Malloch, 1939f: 422. PNG (PNG).
helomyzoides Walker, 1864: 220 (Strumeta). Indonesia (Maluku); Indonesia (Irian Jaya).
     helemyzoides, error for helomyzoides.
hyalina Hardy, 1986b: 147. PNG (PNG).
nigrifacies Hardy, 1986b: 149. Indonesia (Irian Jaya).
perkinsi Hardy, 1986b: 149. Indonesia (Irian Jaya).
suttoni Malloch, 1939f: 423. PNG (PNG); Indonesia (Irian Jaya).
variabilis Hardy, 1986b: 151. PNG (PNG); Indonesia (Irian Jaya).


THEMAROIDES Hendel, 1914c: 77. Type species: Helomyza quadrifera Walker, 1861, orig. des.
RIOXINA Hering, 1941e: 16. Type species: Rioxa debeauforti de Meijere, 1906 [= Seraca abbreviata Walker, 1865], orig. des.
abbreviata Walker, 1865b: 117 (Seraca). "New Guinea" [= Indonesia (Irian Jaya)]; PNG (PNG).
     debeauforti de Meijere, 1906b: 94 (Rioxa). Indonesia (Irian Jaya).
bicolor Hancock & Drew, 2003: 72. PNG (Bismarck Arch).
quadrifera Walker, 1861b: 246 (Helomyza). PNG (PNG); Indonesia (Irian Jaya, Maluku).
     optatura Walker, 1865b: 116 (Helomyza). "New Guinea" [= Indonesia (Irian Jaya)].
     ampla, Doleschall, 1858, not Walker, 1856, misid.
robertsi Hardy, 1986b: 156. PNG (PNG).
vittata Hardy, 1986b: 158. PNG (PNG).
xanthosoma Hardy, 1986b: 160. PNG (PNG).


THEMAROIDOPSIS Hering, 1941e: 17. Type species: Acanthoneura insignis de Meijere, 1913, orig. des.
insignis de Meijere, 1913b: 366 (Acanthoneura). Indonesia (Irian Jaya).
quinquevittata Hardy, 1986b: 162. PNG (PNG).
rufescens Hardy, 1986b: 163. Solomon Is.
tetraspilota Hardy, 1986b: 164. PNG (PNG).


TRYPANOCENTRA Hendel, 1914c: 33 [1915: 433]. Type species: Trypanocentra nigripennis Hendel, 1914 [preocc., = Trypanocentra atripennis Malloch, 1939], orig. des.

Subgenus CLUSIOMORPHA Hering

CLUSIOMORPHA Hering, 1941f: 111 (as gen.). Type species: Clusiosoma funebre Hering, 1941, orig. des.
atrifacies Hardy, 1986b: 173. PNG (PNG).
bipectinata Hardy, 1986b: 174. PNG (PNG).
funebris Hering, 1941d: 61 (Clusiosoma). PNG (PNG).
gressitti Hardy, 1986b: 176. Indonesia (Irian Jaya).
nigripennis de Meijere, 1913b: 366 (Acanthoneura). Indonesia (Irian Jaya); PNG (Bismarck Arch, PNG).
tenuifemorale Hering, 1941d: 58 (Clusiosoma). PNG (PNG).
     adustata Hering, 1947a: 3. PNG (Bismarck Arch).
     adjusta, error for adustata.


TRYPANOCENTRA Hendel, 1914c: 33 [1915: 433] (as gen.). Type species: Trypanocentra nigripennis Hendel, 1914 [preocc., = Trypanocentra atripennis Malloch, 1939], orig. des.
atripennis Malloch, 1939f: 430 (n. n. for nigripennis Hendel). ?PNG (PNG).
     nigripennis Hendel, 1914c: 33 [1915: 434]. "Indischen-Archipel" [prob. = New Guinea]. [Preocc. de Meijere, 1914.]
longicornis Hardy, 1986b: 168. PNG (PNG).
mallochi Hardy, 1986b: 169. PNG (PNG).
nigridorsalis Hering, 1941d: 59 (Clusiosoma). PNG (PNG).
nigrithorax Malloch, 1939f: 428. PNG (PNG); Australia (Qld), PNG (PNG).
tricuneata Hardy, 1986b: 172. PNG (PNG).


WALKERAITIA Hardy, 1986b: 177. Type species: Helomyza nivistriga Walker, 1861, orig. des.
nivistriga Walker, 1861b: 246 (Helomyza). "New Guinea."

Genus XARNUTA Walker

XARNUTA Walker, 1856a: 28. Type species: Xarnuta leucotelis Walker, 1856, mon.
confusa Malloch, 1939d: 261. Solomon Is; Australia (Qld), PNG (PNG).
cribralis Hering, 1941d: 57. PNG (PNG); Australia (Qld), PNG (Bismarck Arch).
leucotela Walker, 1856a: 28. Singapore; Indonesia (Irian Jaya, Maluku); widesp. Oriental Reg.
stellaris Hardy, 1970: 85. Philippines; PNG (Bismarck Arch, PNG), Solomon Is.



AGNOSTOPHANA Hering, 1953: 510. Type species: Agnostophana veterrima Hering, 1953, orig. des.
AGNOSTOPHARA, error for Agnostophana.
veterrima Hering, 1953: 512. PNG (PNG); Indonesia (Irian Jaya).


EPACROCERUS Hardy, 1982b: 80. Epacrocerus splendens Hardy, 1982, orig. des.
apiculatus Hardy, 1982b: 80. PNG (PNG).
maculatus Hardy, 1982b: 82. PNG (PNG); Indonesia (Irian Jaya).
quadrivittatus Hardy, 1982b: 82. PNG (PNG).
splendens Hardy, 1982b: 85. PNG (PNG); Indonesia (Irian Jaya).


EXALLOSOPHIRA Hardy, 1980b: 149. Type species: Exallosophira elegans Hardy, 1980, orig. des.
elegans Hardy, 1980b: 149. Solomon Is.


ORTALOPTERA Edwards, 1915b: 419. Type species: Ortaloptera cleitamina Edwards, 1915, mon.
callistomyia Hering, 1941e: 11. PNG (PNG); Indonesia (Irian Jaya).
cleitamina Edwards, 1915b: 420. Indonesia (Irian Jaya).


POLYAROIDEA Hardy, 1988a: 105. Type species: Polyaroidea distincta Hardy, 1988, orig. des.
distincta Hardy, 1988a: 106. PNG (PNG).
opposita Hardy, 1988a: 106. PNG (PNG).
univittata Hardy, 1988a: 107. PNG (PNG).


PROEPACROCERUS Hardy, 1988a: 110. Type species: Proepacrocerus pallidoviridus Hardy, 1988, orig. des.
pallidoviridus Hardy, 1988a: 110. Indonesia (Irian Jaya).


PSEUDACROTOXA Hering, 1941c: 51. Type species: Pseudacrotoxa appendicigera Hering, 1941, orig. des.
appendicigera Hering, 1941c: 52. PNG (PNG).

Genus SOPHIRA Walker

SOPHIRA Walker, 1856a: 34. Type species: Sophira venusta Walker, 1856, mon.
venusta Walker, 1856a: 35. Singapore; Indonesia (Maluku); "Borneo," Indonesia (Sumatra).

Genus STYMBARA Walker

STYMBARA Walker, 1865b: 127. Type species: Stymbara vagaria Walker, 1865, mon.
CURVINERVUS Hardy, 1959: 203. Type species: Strumeta concisa Walker, 1864, orig. des.
concisa Walker, 1864: 227 (Strumeta). Indonesia (Maluku); Indonesia (Irian Jaya), PNG (PNG).
     vagaria Walker, 1865b: 127. "New Guinea" [= Indonesia (Irian Jaya)].
     walkeri Hardy, 1959: 203 (Curvinervus; unnec. n. n. for concisa Walker). [The names concisa Walker, 1861 and concisa Walker, 1864 were neither primary nor secondary homonyms at the time of the new replacement name by Hardy.]


TAENIOSTOLA Bezzi, 1913b: 119. Type species: Taeniostola vittigera Bezzi, 1913, orig. des.
limbata Hendel, 1915: 435. Taiwan; PNG (PNG); India, Nepal, Thailand.


TANYMETOPUS Hardy, 1982b: 88. Type species: Tanymetopus claripennis Hardy, 1982, orig. des.
claripennis Hardy, 1982b: 88. PNG (PNG); PNG (Bismarck Arch).


UDAMOLOBIUM Hardy, 1982b: 90. Type species: Udamolobium pictulum Hardy, 1982, orig. des.
pictulum Hardy, 1982b: 90. PNG (PNG).


XENOSOPHIRA Hardy, 1980b: 157. Type species: Xenosophira invibrissata Hardy, 1980, orig. des.
invibrissata Hardy, 1980b: 158. PNG (PNG).
vibrissata Hardy, 1980b: 159. PNG (PNG).


Genus ADRAMA Walker

ADRAMA Walker, 1859a: 117. Type species: Adrama selecta Walker, 1859, mon.
biseta Malloch, 1939e: 332. Australia (Qld); Australia (NT).
fuscoapicata Malloch, 1939d: 249. Solomon Is.
ismayi Hardy, 1986a: 64. PNG (Bismarck Arch).
rufithorax Malloch, 1939d: 249. Solomon Is; PNG (Bismarck Arch).
rufiventris Walker, 1860b: 163 (Enicoptera). Indonesia (Maluku); Philippines, Thailand.
     ceramensis de Meijere, 1914b: 192. Indonesia (Maluku).
     selecta, de Meijere, 1913, not Walker, 1859, misid.
selecta Walker, 1859a: 118. Indonesia (Maluku); Australia (NT, Qld), Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG), Solomon Is; Philippines.
     cruciata Walker, 1865b: 126 (Psila). "New Guinea" [= Indonesia (Irian Jaya)].
     papuaensis Malloch, 1939e: 333. PNG (PNG).
     spinata Enderlein, 1920a: 360. PNG (Bismarck Arch).
     centralis Malloch, 1939d: 247. Solomon Is.


BRANDTOMYIA Hardy, 1986a: 68. Type species: Brandtomyia spuria Hardy, 1986, orig. des.
spuria Hardy, 1986a: 69. PNG (Bismarck Arch).


ICHNEUMONOSOMA de Meijere, 1914b: 195. Type species: Lagarosia imitans de Meijere, 1911, orig. des.
consors Walker, 1861d: 296 (Adrama). Indonesia (Maluku); Australia (Qld), PNG (PNG).

Genus PAPUADRAMA Hancock & Drew

PAPUADRAMA Hancock & Drew, 2005: 10. Type species: Papuadrama buna Hancock & Drew, 2005, orig. des.
buna Hancock & Drew, 2005: 11. PNG (PNG).

Genus SOITA Walker

SOITA Walker, 1865c: 135. Type species: Soita psiloides Walker, 1865, mon.
psiloides Walker, 1865c: 136. Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).


TERASTIOMYIA Bigot, 1859b: 311. Type species: Terastiomyia lobifera Bigot, 1859, mon.
NEOSOPHIRA Hendel, 1914d: 8, 138 [1914e: 15, 76]. Type species: Sophira distorta Walker, 1857, orig. des.
TERASTOMYIA, error for Terastiomyia.
clavigera Hardy, 1958a: 79 (Neosophira). Indonesia (Maluku).
lobifera Bigot, 1859b: 311. Indonesia (Sulawesi); Indonesia (Maluku).
     ferruginea Hendel, 1914d: 138 (Neosophira). Indonesia (Maluku).



CALOSPHENISCA Hendel, 1914: 88. Type species: Calosphenisca voluscris Hendel, 1914, orig. des.
ndomae Hancock & Drew, 2003: 102. Solomon Is.


COELOTRYPES Bezzi, 1924b: 494. Type species: Coelotrypes vittatus Bezzi, 1924, orig. des.
circumscriptus Hering, 1941b: 28 (Staurella). Indonesia (Flores I); Australia (NT, Qld).
flavina Hering, 1941d: 56 (Staurella). PNG (PNG); ?Niue, Solomon Is.
punctilabris Bezzi, 1928: 107 (Ocneros). Fiji; Solomon Is, Tonga, Western Samoa.

Genus CYCLOPSIA Malloch

CYCLOPSIA Malloch, 1939f: 444. Type species: Cyclopsia inaequalis Malloch, 1939 [= Dacus inscripta Walker, 1861], orig. des.
CRINITISOPHIRA Hardy, 1986a: 69. Type species: Crinitisophira bicolor Hardy, 1986, orig. des.
inscripta Walker, 1860b: 162 (Dacus). "Amboyna," error [= "Ambong"; Malaysia (Sabah)]; Indonesia (Irian Jaya, Maluku), PNG (PNG).
     inaequalis Malloch, 1939f: 445. Indonesia (Irian Jaya).
     bicolor Hardy, 1986a: 70 (Crinitisophira). Indonesia (Maluku).
Hardy, 1970: 87. Philippines; PNG (PNG); Malaysia (Pen).


EUPHRANTA Loew, 1862d: 28. Type species: Musca connexa Fabricius, 1794, mon.
RHACOCHLAENA Loew, 1862d: 50 (as gen.). Type species: Trypeta toxoneura Loew, 1846, orig. des.
STAURELLA Bezzi, 1913b: 121 (as gen.). Type species: Musca crux Fabricius, 1794, orig. des.
XANTHOTRYPETA Malloch, 1939d: 250. Type species: Xanthotrypeta bimaculata Malloch, 1939, orig. des.
PARAEUPHRANTA Hardy, 1959: 173. Type species: Dacus furcifer Walker, 1861, orig. des.
athertonia Permkam & Hancock, 1995b: 1150. Australia (Qld).
basalis Walker, 1865b: 126 (Psila). "New Guinea" [= Indonesia (Irian Jaya)].
betikamae Hancock & Drew, 2004: 152. Solomon Is.
bilineata Hardy, 1983b: 162. PNG (PNG).
bimaculata Malloch, 1939d: 250 (Xanthotrypeta). Solomon Is.
bischofi Kertész, 1901c: 427 (Ptilona). Indonesia (Irian Jaya); Philippines.
brunneifemur Hardy, 1983b: 181. Indonesia (Irian Jaya).
furcifera Walker, 1861e: 14 (Dacus). Indonesia (Maluku).
Hancock & Drew, 2004: 155. Solomon Is.
Norrbom & Hancock, 2004: 73. New Caledonia (New Caledonia).
isabellae Hancock & Drew, 2004: 155. Solomon Is.
Hardy, 1983b: 165. PNG (PNG).
leichhardtiae Permkam & Hancock, 1995b: 1151. Australia (Qld); Australia (NSW), New Caledonia (New Caledonia).
lemniscata Enderlein, 1911b: 426 (Trypeta). Taiwan; Australia (Qld), Fiji, New Caledonia (New Caleodina), Northern Marianas, Tonga, PNG (PNG); Burma, India.
     rivulosa Bezzi, 1928: 109. Fiji.
lemniscoides Hancock & Drew, 2003: 94. Solomon Is; PNG (Bismarck Arch).
linocierae Hardy, 1951b: 176. Australia (Qld).
marginata Hardy, 1983b: 191. PNG (PNG).
marina Permkam & Hancock, 1995b: 1157. Australia (Qld); Australia (NSW, NT), PNG (PNG).
mediofusca Hering, 1941e: 14 (Staurella). PNG (Bismarck Arch); Australia (Qld).
meringae Permkam & Hancock, 1995b: 1161. Australia (Qld); Solomon Is.
minor Hendel, 1928b: 362. Australia (NT); Australia (Qld).
moluccensis Hardy, 1983b: 192. Indonesia (Maluku).
mulgravea Permkam & Hancock, 1995b: 1164. Australia (Qld).
nigroapicalis Hardy, 1983b: 194. PNG (PNG).
numeralis Permkam & Hancock, 1995b: 1165. Australia (NSW).
ochrosiae Hancock & Drew, 2003: 97. PNG (PNG).
pallida Hardy, 1983b: 169. Indonesia (Irian Jaya).
perkinsi Hardy, 1983b: 195. Indonesia (Irian Jaya).
quadrimaculata Hardy, 1983b: 170. Indonesia (Irian Jaya).
quatei Hardy, 1983b: 195. Indonesia (Irian Jaya).
scutellata Malloch, 1939d: 252. Solomon Is.
sedlaceki Hardy, 1983b: 196. PNG (PNG); PNG (Bismarck Arch).
simonthomasi Hardy, 1983b: 172. Indonesia (Irian Jaya).
solitaria Hardy, 1983b: 196. PNG (Bismarck Arch); Solomon Is.
ternaria Permkam & Hancock, 1995b: 1166. Australia (Qld).
transiens Walker, 1860b: 164 (Trypeta). Indonesia (Maluku).
tricolor Hardy, 1983b: 174. PNG (PNG).
variabilis Kertész, 1901c: 426. Indonesia (Irian Jaya); Australia (Qld), PNG (PNG).
vitabilis Hardy, 1970: 125. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG).


HARDYADRAMA Lee, 1991: 106. Type species: Hardyadrama excoecariae Lee, orig. des.
alyta Permkam & Hancock, 1995b: 1170. Australia (Qld).
excoecariae Lee, 1991: 107. Singapore; Australia (Qld, WA), PNG (PNG).
magister Lee, 1991: 111 (Adrama). Singapore; Australia (Qld), PNG (PNG).
presignis Hardy, 1973: 147 (Euphranta). Thailand; Australia (Qld); Philippines.


PIESTOMETOPON Meijere, 1914: 213. Type species: Piestometopon luteiceps Meijere, 1914, orig. des.
ELLEIPSA Hardy, 1970: 90. Type species: Elleipsa quadrifasciata Hardy, 1970, orig. des.
distinctumPermkam & Hancock, 1995b: 1146 (Elleipsa). Australia (Qld); New Caledonia (New Caledonia).
luteiceps Meijere, 1914: 213. Indonesia (Java); Australia (Qld), Tonga; Philippines, Singapore.
     quadrifasciata Hardy, 1970: 90 (Elleipsa). Philippines.

Genus PTILONA Wulp

PTILONA Wulp, 1880: 183. Type species: Ptilona brevicornis Wulp, 1880 [= Rioxa confinis Walker, 1856], des. Bezzi, 1913b: 68.
PTIOLINA, error for Ptilona.
confinis Walker, 1856c: 132 (Rioxa). Malaysia (Sarawak); Indonesia (Irian Jaya, Maluku); widesp. Oriental Reg.
     alboguttata Doleschall, 1858: 124 (Themara). Indonesia (Maluku).
     basifascia Walker, 1860b: 158 (Trypeta). Indonesia (Maluku).
     bimaculata Walker, 1860b: 164 (Rioxa). Indonesia (Maluku).


Genus DIPLOCHORDA Osten Sacken

DIPLOCHORDA Osten Sacken, 1881a: 484. Type species: Dacus turgidus Walker, 1865, des. Hendel, 1908d: 4.
NESADRAMA Perkins, 1939: 2. Type species: Nesadrama longistigma Perkins, 1939 [= Dacus turgidus Walker, 1865], mon.
aneura Malloch, 1939c: 175. PNG (PNG).
australis Permkam & Hancock, 1995b: 1178. Australia (Qld).
brevicornis Saunders, 1861: 414, 415 (Elaphomyia). Indonesia (Irian Jaya).
concisa Walker, 1861b: 252 (Dacus). Indonesia (Irian Jaya).
minor Malloch, 1939c: 178. PNG (PNG).
myrmex Osten Sacken, 1881a: 488. Indonesia (Irian Jaya); Philippines.
     petiolata Hardy, 1974: 105 (Nesadrama). Philippines.
ophion Osten Sacken, 1881a: 488. Indonesia (Irian Jaya).
trineata de Meijere, 1913c: xl [1915b: 124 (as trilineata)]. Indonesia (Irian Jaya).
     myrmex, Malloch, 1939, not Osten Sacken, 1881, misid.
     trilineata, error for trineata.
turgida Walker, 1865c: 134 (Dacus). Indonesia (Irian Jaya); PNG (PNG).
     longistigma Perkins, 1939: 3 (Nesadrama). PNG (PNG).
     concisa, Malloch, 1939, not Walker, 1861, misid.
unistriata Malloch, 1939c: 179. PNG (PNG).

Genus PHYTALMIA Gerstaecker

PHYTALMIA Gerstaecker, 1860: 169. Type species: Phytalmia cervicornis Gerstaecker, 1860, des. Enderlein, 1936a: 229.
ELAPHOMYIA Saunders, 1861: 413. Type species: Elaphomyia alcicornis Saunders, 1861, des. Enderlein, 1936a: 230.
ARCHIPHYTALMIA Enderlein, 1936a: 230. Type species: Archiphytalmia prisca Enderlein, 1936 [= Phytalmia cervicornis Gerstaecker, 1860], orig. des.
alcicornis Saunders, 1861: 415 (Elaphomyia). Indonesia (Irian Jaya).
antilocapra McAlpine & Schneider, 1978: 170. PNG (PNG).
biarmata Malloch, 1939c: 174. PNG (PNG); Indonesia (Irian Jaya).
cervicornis Gerstaecker, 1860: 173. "Nova Guinea"; Indonesia (Irian Jaya), PNG (PNG).
     cervicornis Saunders, 1861: 414 (Elaphomyia). Indonesia (Irian Jaya). [Preocc. Gerstaecker, 1960.]
     prisca Enderlein, 1936a: 230 (Archiphytalmia). Indonesia (Irian Jaya).
megalotis Gerstaecker, 1860: 172. "Nova Guinea"; Indonesia (Irian Jaya), PNG (PNG).
     wallacei Saunders, 1861: 413, 414 (Elaphomyia). Indonesia (Irian Jaya).
mouldsi McAlpine & Schneider, 1978: 166. Australia (Qld).


ROBERTSOMYIA Hardy, 1983a: 228. Type species: Robertsomyia paradoxa Hardy, 1983, orig. des.
paradoxa Hardy, 1983a: 230. PNG (PNG).

Genus SESSILINA McAlpine & Schneider

SESSILINA McAlpine & Schneider, 1978: 171. Type species: Dacus nigrilinea Walker, 1861, orig. des.
horrida McAlpine & Schneider, 1978: 173. Indonesia (Irian Jaya).
literata McAlpine & Schneider, 1978: 173. PNG (PNG).
nigrilinea Walker, 1861b: 251 (Dacus). Indonesia (Irian Jaya); PNG (PNG).
wollastoni Edwards, 1915b: 418 (Phytalmia). Indonesia (Irian Jaya).


Genus ORNITHOSCHEMA de Meijere

ORNITHOSCHEMA de Meijere, 1914b: 221. Type species: Ornithoschema oculatum de Meijere, 1914, orig. des.
CYCASIA Malloch, 1942: 202. Type species: Cycasia oculata Malloch, 1942 [preocc., = Ornithoschema mallochi Hardy, 1993], orig. des.
oculatum de Meijere, 1914b: 221. Indonesia (Java); Australia (NT), PNG (Bismarck Arch), Solomon Is; Malaysia (Sabah), Thailand.
     flava Hardy, 1973: 168. Thailand.
mallochi Hardy, 1993: 1 (n.n. for oculata Malloch). Guam; New Caledonia (New Caledonia).
     oculata Malloch, 1942: 203 (Cycasia). Guam.
     pacifica Hancock & Drew, 1994: 26 (n.n. for oculata Malloch).
queenslandense Permkam & Hancock, 1995b: 1183. Australia (Qld).


Genus ACIDIELLA Hendel

ACIDIELLA Hendel, 1914: 83. Type species: Acidiella longipennis Hendel, 1914, orig. des.
bimaculata Hardy, 1987: 318 (Myoleja). PNG (PNG).


ACIUROPSIS Hardy, 1974: 96. Type species: Aciuropsis pusio Hardy, 1974, orig. des.
pusio Hardy, 1974: 96. Philippines; PNG (PNG).


AGARISTINA Hering, 1953: 514. Type species: Agaristina picea Hering, 1953, orig. des.
picea Hering, 1953: 515. Indonesia (Irian Jaya); PNG (PNG).

Genus ANOMOIA Walker

ANOMOIA Walker, 1835b: 80. Type species: Trypeta gaedii Meigen, 1830 (as goedii) [= Musca purmundus Harris, 1780], mon.
PHAGOCARPUS Rondani, 1871a: 171 [1871b: 31]. Type species: Musca purmundus Harris, 1780, mon.
NEANOMOEA Hendel, 1914c: 84. Type species: Neanomoea approximata Hendel, 1914, orig. des.
klossi Edwards, 1919: 51 (Phagocarpus). Indonesia (Sumatra); PNG (PNG); China, Philippines.
     occultus Hering, 1939c: 144. China.
melanopoda Hering, 1953: 522. Indonesia (Irian Jaya); PNG (PNG).
modica Hardy, 1987: 282. PNG (PNG); Indonesia (Irian Jaya).
nigrithorax Malloch, 1939f: 449. PNG (PNG); Indonesia (Irian Jaya).
     curvivenis Hering, 1953: 522. Indonesia (Irian Jaya).


CALLISTOMYIA Bezzi, 1913b: 124. Type species: Callistomyia pavonina Bezzi, 1913, orig. des.
flavilabris Hering, 1953: 513. Indonesia (Irian Jaya); Malaysia (Pen).
horni Hendel, 1928b: 361. Australia ("Palmerston, WA") [= (NT)]; Australia (Qld), PNG (PNG).


CRASPEDOXANTHITEA Hardy, 1987: 285. Type species: Craspedoxanthitea flaviseta Hardy, 1987, orig. des.
flaviseta Hardy, 1987: 286. PNG (PNG).
indistincta de Meijere, 1913b: 364 (Trypeta). Indonesia (Irian Jaya).


CRISTOBALIA Malloch, 1939d: 265. Type species: Cristobalia lutea Malloch, 1939, orig. des.
lutea Malloch, 1939d: 265. Solomon Is.

Genus EPINETTYRA Permkam & Hancock

EPINETTYRA Permkam & Hancock, 1995b: 1188. Type species: Epinettyra setosa Permkam & Hancock, 1995, orig. des.
setosa Permkam & Hancock, 1995b: 1189. Australia (Qld).


FUSCILUDIA Ito, 1984: 182. Type species: Fusciludia aliquantula Ito, 1984, orig. des.
bicuneata Hardy, 1987: 317 (Myoleja). PNG (PNG).
mesopleuralis Malloch, 1939g: 242 (Pseudospheniscus). Fiji.
unicuneata Hardy, 1987: 338 (Myoleja). PNG (PNG); Australia (Qld, NSW).


GRIPHOMYIA Hardy, 1987: 290. Type species: Griphomyia spilota Hardy, 1987, orig. des.
argentifrons Hardy, 1987: 292. PNG (PNG).
brunnipennis Hardy, 1987: 292. PNG (PNG).
spilota Hardy, 1987: 293. PNG (PNG).
vittata Hardy, 1987: 295. PNG (Bougainville I).
vittifrons Hardy, 1987: 296. PNG (PNG).

Genus HEMILEA Loew

HEMILEA Loew, 1862d: 32. Type species: Trypeta dimidiata Costa, 1844, orig. des.
araliae Malloch, 1939d: 271. PNG (Bismarck Arch).
atrata Hardy, 1987: 302. PNG (PNG).
bipars Walker, 1861e: 23 (Sophira). Indonesia (Maluku); PNG (PNG); Laos, Philippines.
flavoscutellata Malloch, 1939d: 270. Solomon Is.
lineomaculata Hardy, 1987: 306. PNG (PNG).

Genus HEMIRISTINA Permkam & Hancock

HEMIRISTINA Permkam & Hancock, 1995b: 1192. Type species: Hemiristina pleomeles Permkam & Hancock ,1995, orig. des.
pleomeles Permkam & Hancock, 1995b: 1193. Australia (Qld).

Genus LALOKIA Hardy

LALOKIA Hardy, 1987: 309. Type species: Lalokia tetraspilota Hardy, 1987, orig. des.
tetraspilota Hardy, 1987: 309. PNG (PNG).

Genus MYOLEJA Rondani

MYOLEJA Rondani, 1856: 112. Type species: Tephritis lucida Fallén, 1826, orig. des.
conjuncta de Meijere, 1913a: 61 (Anomoea). Indonesia (Irian Jaya); Indonesia (Maluku), PNG (Bismarck Arch), Solomon Is.
     apicifasciatus Malloch, 1939d: 267 (Pseudospheniscus). Solomon Is.
megaloba Hardy, 1987: 327. PNG (PNG).
nigrescens Shiraki, 1968: 42 (Hendelina). Ryukyu Is; Indonesia (Irian Jaya), PNG (PNG).
propreincerta Hardy, 1987: 331. PNG (Bougainville I); Solomon Is.
superflucta Enderlein, 1911b: 428 (Trypeta). Taiwan; Indonesia (Maluku); Malaysia (Pen, Sarawak), Philippines, Ryukyu Is.


PARACRISTOBALIA Hardy, 1987: 345. Type species: Paracristobalia polita Hardy, 1987, orig. des.
polita Hardy, 1987: 346. PNG (PNG).


PHILOPHYLLA Rondani, 1870a: 9. Type species: Musca caesio Harris, 1780, orig. des.
HENDELINA Hardy, 1951b: 179. Type species: Pseudopheniscus angulatus Hendel, 1913, orig. des.
PSEUDOSPHENISCUS, authors, not Hendel, 1914, misid.
australina Hardy, 1951b: 180 (Hendelina). Australia (Qld).
bifida Bezzi, 1928: 112 (Pseudospheniscus). Fiji.
curvinervis Bezzi, 1928: 114 (Pseudospheniscus). Fiji.
dividua Hardy, 1987: 277 (Anomoia). PNG (PNG).
erebia Hering, 1941d: 62 (Pseudospheniscus). PNG (PNG); Australia (NSW, Qld); Malaysia (Sabah).
fossata Fabricius, 1805: 320 (Tephritis). India; Australia (Qld), Indonesia (Maluku), PNG (Bismarck Arch, PNG), Solomon Is; Japan, widesp. Oriental Reg.
     elimia Walker, 1849: 1033 (Trypeta). Philippines.
     regularis Doleschall, 1858: 119 (Ortalis). Indonesia (Maluku).
     lieftincki Hering, 1952: 286 (Neanomoeba). Indonesia (Java).
humeralis Hendel, 1915: 452 (Pseudospheniscus). PNG (PNG); Australia (Qld), PNG (Bismarck Arch).
nigroscutellata Hering, 1939b: 18 (Neanomoea). Burma; Guam, Northern Marianas; Indonesia (Java), Malaysia (Sarawak).
     bisecta Hardy & Adachi, 1956: 17 (Hendelina). Guam.
quadrata Malloch, 1939d: 275 (Anomoia). Solomon Is; Australia (NT, Qld), PNG (Bismarck Arch, PNG).
     shirakii Hardy, 1987: 335. Solomon Is.
taylori Malloch, 1939f: 450 (Pseudospheniscus). PNG (PNG); Indonesia (Irian Jaya), Micronesia, PNG (Bismarck Arch), Solomon Is.
     parva Hardy & Adachi, 1956: 18 (Hendelina). Micronesia.


RHAGOLETIS Loew, 1862d: 44. Type species: Musca cerasi Linnaeus, 1758, mon.
conversa Brèthes, 1919: 43 (Spilographa). Chile; Easter I.


TANAODEMA Hardy, 1987: 350. Type species: Tanaodema porrecta Hardy, 1987, orig. des.
porrectum Hardy, 1987: 351. PNG (PNG).


TARPHOBREGMA Hardy, 1987: 353. Type species: Tarphobregma pandani Hardy, 1987, orig. des.
carinatifrons Hardy, 1987: 354. PNG (PNG).
pandani Hardy, 1987: 358. PNG (PNG).

Genus VIDALIA Robineau-Desvoidy

VIDALIA Robineau-Desvoidy, 1830: 719. Type species: Vidalia impressifrons Robineau-Desvoidy, 1830, mon.
STEMONOCERA Rondani, 1870a: 30 [1870c: 26]. Type species: Musca cornuta Scopoli, 1772, mon.
PSEUDINA Malloch, 1939f: 446. Type species: Pseudina buloloae Malloch, 1939, orig. des.
bicolor Hardy, 1987: 364. PNG (PNG).
buloloae Malloch, 1939f: 446 (Pseudina). PNG (PNG).
dualis Permkam & Hancock, 1995b: 1202. Australia (Qld).



Genus UROPHORA Robineau-Desvoidy

EURIBIA Meigen, 1800: 36. Suppressed by I.C.Z.N., 1963b: 339.
UROPHORA Robineau-Desvoidy, 1830: 769. Type species: Musca cardui Linnaeus, 1758, des. Westwood, 1840: 149.
quadrifasciata Meigen, 1826: 331 (Trypeta). Germany; Australia (NT); Palearctic Reg.
     quadrifaciata. Incorr. orig. spell. of quadrifasciata (Meigen, 1826: 331).
stylata Fabricius, 1775: 785 (Musca). "Angliae nemoribus" [= ?England]; Australia (NT); Pakistan, Palearctic Reg.



SCHISTOPTERUM Becker, 1903: 137. Type species: Schistopterum moebiusi Becker, 1903, mon.
ismayi Hardy, 1982a: 88. PNG (PNG).



CURTICELLA Hardy, 1959: 209. Type species: Trypeta approximans Walker, 1860, orig. des.
approximans Walker, 1860a: 160 (Trypeta). Indonesia (Sulawesi); Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG).


SPHAENISCUS Becker, 1908: 138. Type species: Sphaeniscus brevicauda Becker, 1908 [= Aciura filiola Loew, 1869], mon.
SPHENISCOMYIA Bezzi, 1913b: 146 (unjust. n. n. for Sphaeniscus).
SPHENISCUS Bezzi, 1913b: 146 (unjust. emend. of Sphaeniscus). [Preocc. Brisson, 1760.]
atilius Walker, 1849: 1021 (Trypeta). China; widesp. Australasian/Oceanian Regs.; widesp. Oriental Reg.
     melaleuca Walker, 1864: 238 (Trypeta). Indonesia (Maluku).
     sexmaculata, authors, not Macquart, 1843, misid.
binoculatus Bezzi, 1928: 115. Fiji.


Genus ACINIA Robineau-Desvoidy

ACINIA Robineau-Desvoidy, 1830: 775. Type species: Acinia jaceae Robineau-Desvoidy, 1830 [= Tephritis corniculata Zetterstedt, 1819], des. Rondani, 1871a: 6 [1871b: 4].
picturata Snow, 1894: 173 (Tephritis). USA; Hawaiian Is, Johnston Atoll, Wake I [all introduced]; West Indies, Mexico to Guatemala.
     furcata, authors, not Fabricius, 1794, misid.


AUSTRALASINIA Hancock, 2001: 113. Type species: Tephrella sexincisa Malloch, 1939, orig. des.
sexincisa Malloch, 1939d: 272 (Tephrella). Solomon Is; Australia (NSW, Qld, SA), Indonesia (Nusa Tenggara).

Genus AUSTROTEPHRITIS Hancock & Drew

AUSTROTEPHRITIS Hancock & Drew, 2003: 150. Type species: Trypeta poenia Walker, 1849, orig. des.
brunneimacula Hardy, 1988b: 37 (Paroxyna). PNG (PNG).
fuscata Macquart, 1851: 239(266) (Acinia). Australia (Tas); Australia (ACT, NSW, Qld, SA, Vic).
transversa Hardy & Drew, 1996: 223 (Campiglossa). Australia (Vic).
turneri Hardy & Drew, 1996: 226 (Campiglossa). Australia (WA).
bushi Hardy & Drew, 1996: 354 (Tephritis). Australia (ACT); Australia (NSW, Vic).
cassiniae Malloch, 1931h: 395 (Tephritis). NZ (NZ).
distigmata Hardy & Drew, 1996: 356 (Tephritis). Australia (WA).
hesperia Hardy & Drew, 1996: 360 (Tephritis). Australia (WA).
marginata Malloch, 1931h: 394 (Tephritis). NZ (NZ).
pelia Schiner, 1868: 271 (Tephritis). Australia (NSW); widesp. Australia.
phaeostigma Hardy & Drew, 1996: 367 (Tephritis). Australia (Vic); Australia (SA, WA).
plebeia Malloch, 1931h: 393 (Tephritis). NZ (NZ).
poenia Walker, 1849: 1025 (Trypeta). "New Holland"; widesp. Australia, Indonesia (Irian Jaya), PNG (PNG).
     leontodontis, authors, not De Geer, 1776, ?misid.
     pelia, Malloch, 1939, not Schiner, 1868, misid.
protrusa Hardy & Drew, 1996: 373 (Tephritis). Australia (Qld); Australia (NSW).
pumila Hardy & Drew, 1996: 376 (Tephritis). Australia (ACT); Australia (NSW, Qld, SA, Vic, WA).
quasiprolixa Hardy & Drew, 1996: 378 (Tephritis). Australia (SA); Australia (NSW).
tasmaniae Hardy & Drew, 1996: 380 (Tephritis). Australia (Tas).
thoracica Malloch, 1931h: 392 (Tephritis). NZ (NZ).
trupanea Hardy & Drew, 1996: 381 (Tephritis). Australia (SA); Australia (NSW, Qld, SA, WA).
whitei Hardy & Drew, 1996: 229 (Campiglossa). Australia (Tas).

Genus BEZZINA Munro

BEZZIELLA Munro, 1937: 9. Type species: Oxyna margaritifera Bezzi, 1908, orig. des. [Preocc. Enderlein, 1937.]
Munro, 1957: 893 (n.n. for Bezziella Munro).
trimaculata Hardy & Drew, 1996: 320 (Platensina). Australia (Qld).


CAMPIGLOSSA Rondani, 1870a: 123 [1870c: 49] (Oxyna subg.). Type species: Tephritis irrorata Fallén, 1814, orig. des.
PAROXYNA Hendel, 1927b: 146. Type species: Trypeta tessellata Loew, 1844, orig. des.
angustipennis Malloch, 1938a: 115 (Paroxyna). French Polynesia (Austral Is).
longirostris Thomson, 1869: 586 (Trypeta). Tonga. [Preocc. Loew, 1846.]
media Malloch, 1938a: 116 (Paroxyna). French Polynesia (Austral Is).
paula Hering, 1941d: 64 (Paroxyna). PNG (PNG); Indonesia (Maluku).
putrida Hering, 1941b: 39. (Paroxyna) Indonesia (Nusa Tenggara); PNG (PNG).
stigmosa de Meijere, 1916b: 83 (Tephritis). Indonesia (Sumatra); PNG (PNG); Indonesia (Nusa Tenggara, Timor).
     timorensis Hering, 1940: 8 (Paroxyna). Indonesia (Nusa Tenggara).
vaga Hardy & Drew, 1996: 228. Australia (Qld).

Genus COLLESSOMYIA Hardy & Drew

COLLESSOMYIA Hardy & Drew, 1996: 231. Type species: Collessomyia setiger Hardy & Drew, 1996, orig. des.
setiger Hardy & Drew, 1996: 231. Australia (WA); Australia (NT, Qld).
heringi Hardy, 1970: 129 (Tephrella). PNG (Bismarck Arch).

Genus COORONGA Hardy & Drew

COORONGA Hardy & Drew, 1996: 233. Type species Cooronga mcalpinei Hardy & Drew, 1996, orig. des.
mcalpinei Hardy & Drew, 1996: 234. Australia (SA).

Genus DIOXYNA Frey

STYLIA Robineau-Desvoidy, 1830: 754. Type species: Stylia bidentis Robineau-Desvoidy, 1830, des. Hering, 1954: 167. [First valid type- species designation of Stylia maculata Robineau-Desvoidy, 1830, by Desmarest in d'Orbigny, 1848: 78 is of a species considered a nomen dubium (Foote & Blanc, 1979: 170; White, 1986: 148).
DIOXYNA Frey, 1945: 62. Type species: Trypeta sororcula Wiedemann, 1830, orig. des.
brachybasis Hardy, 1988b: 21. PNG (PNG); Australia (Qld), Fiji, Indonesia (Irian Jaya).
conflicta Curran, 1929a: 11 (Ensina). New Caledonia (New Caledonia); Indonesia (Irian Jaya), Niue, PNG (Bismarck Arch), Tonga, Vanuatu, Western Samoa.
fouica Hering, 1951a: 14 (Paroxyna; n. n. for longirostris Thomson). Tonga.
Hering, 1944a: 8 (Paroxyna). "New Guinea."
hyalina Hardy & Drew, 1996: 239. Australia (NSW); Australia (Qld, SA, Vic, WA).
sororcula Wiedemann, 1830a: 509 (Trypeta). Canary Is; widesp. Australia (excl. Tas), Bonin Is, Cook Is, Easter I, Fiji, French Polynesia (Austral Is, Marquesas, Society Is), Hawaiian Is, Indonesia (Irian Jaya), Marshall Is, New Caledonia (New Caledonia), Niue, Northern Marianas, PNG (Bismarck Arch, PNG), Western Samoa; pantrop.
     picciola Bigot, 1857c: 347 [1857d: 827] (Acinia). Cuba.
     plicicollis, error for picciola.


ELAPHROMYIA Bigot, 1859b: 314. Type species: Elaphromyia melas Bigot, 1859 [= Trypeta adatha Walker, 1849], mon.
PARALLOPTERA Bezzi, 1913b: 154. Type species: Paralloptera pterocallaeformis Bezzi, 1913, orig. des.
transversa Hardy, 1988b: 29. PNG (PNG).

Genus ENSINA Robineau-Desvoidy

ENSINA Robineau-Desvoidy, 1830: 751. Type species: Ensina scorzonerae Robineau-Desvoidy, 1830 [= Musca sonchi Linnaeus, 1767], des. Foote & Freidberg, 1981: 30.
ENSINIA, error for Ensina.
sonchi Linnaeus, 1767: 998 (Musca). Sweden; Hawaiian Is [introduced]; widesp. Old World, Neotrop. Reg.


EUARESTA Loew, 1873b: 296. Type species: Trypeta festiva Loew, 1862, des. Coquillett, 1910: 540.
CAMAROMYIA Hendel, 1914c: 95. Type species: Trypeta bullans Wiedemann, 1830, orig. des.
EVARESTA, error for Euaresta.
aequalis Loew, 1862a: 86 (Trypeta). USA; Australia (ACT, Qld) [introduced], Fiji [introduced; ?established]; S Canada.
bullans Wiedemann, 1830a: 506 (Trypeta). Uruguay; Australia (NSW, Qld, Vic) [introduced]; Israel, South Africa, Turkey, USA, USSR, S Europe.

Genus EUTRETA Loew

EUTRETA Loew, 1873b: 276. Type species: Trypeta sparsa Wiedemann, 1830, des. Coquillett, 1910: 543.
xanthochaeta Aldrich, 1923b: 261. Mexico; Hawaiian Is [introduced], Australia [introduced; ?established]; ?Guatemala.
     sparsa, authors, not Wiedemann, 1830, misid.

Genus HYALOPEZA Hardy & Drew

HYALOPEZA Hardy & Drew, 1996: 253. Type species: Hyalopeza schneiderae Hardy & Drew, 1996, orig. des.
aristae Hancock & Drew, 2003: 142. Australia (Qld).
schneiderae Hardy & Drew, 1996: 253. Australia (Vic); Australia (ACT, NS, SAW, Qld).

Genus LIEPANA Hardy & Drew

LIEPANA Hardy & Drew, 1996: 256. Type species: Liepana latifrons Hardy & Drew, 1996, orig. des.
apiciclara Hardy & Drew, 1996: 267 (Oedaspis). Australia (NSW).
helichrysii Hardy & Drew, 1996: 257. Australia (NSW).
latifrons Hardy & Drew, 1996: 253. 259 (Vic).
lugubris Macquart, 1847a: 43 [1847b: 109] (Tephritis). "Nouvelle-Hollande"; Australia (Tas).


OEDASPIS Loew, 1862b: 46. Type species: Trypeta multifasciata Loew, 1850, des. Rondani (1870: 9).
Hendel, 1927c: 63. Type species: Oedaspoides acuticornis Hendel, 1927, mon.
CHRYSOTRYPANEA Malloch, 1939f: 457. Type species: Chrysotrypanea trifasciata Malloch, 1939, orig. des.
acuticornis Hendel, 1927c: 63 (Oedaspoides). Australia (NSW).
apicalis Hardy & Drew, 1996: 265. Australia (NSW).
australis Malloch, 1939f: 456 (Tephrella). Australia (WA); Australia (NSW, Qld), PNG (PNG).
austrina Hardy & Drew, 1996: 268. Australia (SA); Australia (WA).
continua Hardy & Drew, 1996: 270. Australia (WA).
escheri Bezzi, 1910b: 19 (Oedaspis). Australia (NSW); Australia (NT, WA).
gallicola Hardy & Drew, 1996: 275. Australia (NSW); Australia (Vic).
goodenia Hardy & Drew, 1996: 277. Australia (Vic); Australia (NSW, Qld).
hardyi Norrbom in Norrbom et al., 1999: 175 (n. n. for serrata Hardy & Drew). Australia (NSW).
     serrata Hardy & Drew, 1996: 288. Australia (NSW). [Preocc. Freidberg & Kaplan, 1992.]
Hardy & Drew, 1996: 279. Australia (Qld).
olearia Hardy & Drew, 1996: 282. Australia (Vic); Australia (NSW, Tas).
ouinensis Hancock, 2008: 203. New Caledonia (New Caledonia).
perkinsi Hardy & Drew, 1996: 285. Australia (Vic).
semihyalina Hardy & Drew, 1996: 286. Australia (WA).
trifasciata Malloch, 1939f: 457 (Chrysotrypanea). Australia (Vic).
trimaculata Hardy & Drew, 1996: 292. Australia (WA); Australia (NT).
whitei Hardy & Drew, 1996: 293. Australia (WA); Australia (NT, Vic).


PARACTINOPTERA Hardy & Drew, 1996: 299. Type species: Paractinoptera collessi Hardy & Drew, 1996, orig. des.
collessi Hardy & Drew, 1996: 301. Australia (WA).
danielsi Hancock & Drew, 2003: 153. Australia (Qld).
prolixa Hardy & Drew, 1996: 371 (Tephritis). Australia (WA); Australia (NT, SA).

Genus PARAHYALOPEZA Hardy & Drew

PARAHYALOPEZA Hardy & Drew, 1996: 302. Type species: Parahyalopeza bushi Hardy & Drew, 1996, orig. des.
bushi Hardy & Drew, 1996: 303. Australia (Vic).
multipunctata Hancock & Drew, 2003: 155. Australia (Qld).
pantosticta Hardy & Drew, 1996: 362 (Tephritis). Australia (Qld).


PARARHABDOCHAETA Hardy, 1985: 61. Type species: Rhabdochaeta convergens Hardy, 1974, orig. des.
albolineata Hardy, 1985: 62. Indonesia (Irian Jaya).


PARASPATHULINA Hardy & Drew, 1996: 304. Type species: Paraspathulina apicomacula Hardy & Drew, 1996, orig. des.
apicomacula Hardy & Drew, 1996: 305. Australia (ACT); widesp. Australia.
eremostigma Hardy & Drew, 1996: 309. Australia (NSW); Australia (NT, Qld, SA, Vic, WA).
trimacula Hancock & Drew, 2003: 148. Australia (Qld).

Genus PENEPAROXYNA Hardy & Drew

PARAPENOXYNA Hardy & Drew, 1996: 314. Type species: Peneparoxyna minuta Hardy & Drew, 1996, orig. des.
minuta Hardy & Drew, 1996: 315. Australia (NSW); Australia (NT).

Genus PHAEOGRAMMA Grimshaw

PHAEOGRAMMA Grimshaw, 1901: 47. Type species: Phaeogramma vittipennis Grimshaw, 1901, orig. des.
hispida Hardy in Hardy & Delfinado, 1980a: 54. Hawaiian Is.
vittipennis Grimshaw, 1901: 48. Hawaiian Is.

Genus PLATENSINA Enderlein

PLATENSINA Enderlein, 1911b: 453. Type species: Platensina sumbana Enderlein, 1911, mon.
TEPHROSTOLA Bezzi, 1913b: 193. Type species: Trypeta acrostacta Wiedemann, 1824, orig. des.
ampla de Meijere, 1914b: 217. Indonesia (Java); Indonesia (Maluku), PNG (PNG).
amplipennis Walker, 1860a: 159 (Trypeta). Indonesia (Sulawesi); Australia (Qld), Guam, Micronesia, Northern Marianas, PNG (PNG), Solomon Is; Indonesia, Laos, Malaysia (Pen), Taiwan, Thailand, Vietnam.
     platyptera Hendel, 1915: 461. Taiwan.
     malaita Curran, 1936: 29. Solomon Is.
     dubia Malloch, 1939f: 459. Australia (Qld).
     dilatata Hering, 1941d: 63. PNG (PNG).
parvipuncta Malloch, 1939f: 458. Australia (Qld).
zodiacalis Bezzi, 1913b: 165 (Tephritis). India; Australia (NT, Qld), Indonesia (Maluku); widesp. Oriental Reg.
     zodiakalis, error for zodiacalis.


PLIOMELAENA Bezzi, 1918a: 220 (key) [1918b: 930] (Euaresta subg.). Type species: Euaresta brevifrons Bezzi, 1918, des. Munro, 1937: 13.
INDARESTA Hering, 1941b: 36. Type species: Indaresta callista Hering, 1941, orig. des.
callista Hering, 1941b: 36 (Indaresta). Indonesia (Nusa Tenggara); Indonesia (Irian Jaya), PNG (PNG).


PROCECIDOCHARES Hendel, 1914c: 91 [1915: 42]. Type species: Trypeta atra Loew, 1862, orig. des.
alani Steyskal, 1974: 639. Mexico; Australia (Qld [introduced], Hawaiian Is [introduced].
utilis Stone, 1947: 97. Mexico; Australia (NSW, Qld, SA) [introduced], Hawaiian Is [introduced], NZ (NZ) [introduced; ?established].

Genus QUASICOORONGA Hardy & Drew

QUASICOORONGA Hardy & Drew, 1996: 328. Type species: Quasicooronga connecta Hardy & Drew, 1996, orig. des.
connecta Hardy & Drew, 1996: 328. Australia (Vic).
disconnecta Hardy & Drew, 1996: 330. Australia (NSW).

Genus RHABDOCHAETA de Meijere

RHABDOCHAETA de Meijere, 1904b: 109. Type species: Rhabdochaeta pulchella de Meijere, 1904, mon.
asteria Hendel, 1915: 462. Taiwan; PNG (PNG); India, Japan, Laos, Philippines, Ryukyu Is.
crockeri Curran, 1936: 28. Solomon Is; Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG), Solomon Is.
     cockeri, error for crockeri.
guamae Malloch, 1942: 204. Guam.
pluscula Hardy, 1970: 128. PNG (Bismarck Arch).
pulchella de Meijere, 1904b: 109. Indonesia (Java); Australia (Qld), Indonesia (Irian Jaya), PNG (PNG); widesp. SE Asia, Philippines, Ryukyu Is, Sri Lanka.
     bakeri Bezzi, 1914: 328. Philippines.
     mucronata Hering, 1942: 289. Sri Lanka.
     assidua Ito, 1984: 284. Ryukyu Is.
queenslandica Hardy & Drew, 1996: 336. Australia (Qld).
wedelia Hardy & Drew, 1996: 338. Australia (Qld).


RHOCHMOPTERUM Speiser, 1910: 185. Type species: Rhochmopterum neuropteripenne Speiser, 1910, mon.
venustum de Meijere, 1914b: 215 (Rhabdochaeta). Indonesia (Java); Australia (NSW, Qld), Indonesia (Irian Jaya), PNG (PNG); Indonesia (Nusa Tenggara), Japan, Laos, Malaysia (Pen), Philippines, Ryukyu Is, Thailand, Vietnam.
     subsolanum Ito, 1984: 287. Japan.
     dorsosetosa Hardy, 1970: 110 (Rhabdochaeta). Philippines.

Genus SCEDELLA Munro

SCEDELLA Munro, 1957: 988. Type species: Trypeta caffra Loew, 1861, orig. des.
infrequens Hardy & Drew, 1996: 310 (Paroxyna). Australia (NSW); Australia (Qld).
formosella Hendel, 1915: 465 (Euribia). Taiwan; Guam, Indonesia (Irian Jaya, Maluku), PNG (Bismarck Arch, PNG), Solomon Is; Indonesia (Nusa Tenggara), Malaysia (Pen), Philippines, Ryukyu Is.
     pentagonella Bezzi, 1928: 117 (Euribia). Fiji.
de Meijere, 1908a: 130 (Tephritis). Indonesia (Java); Australia (Qld); Sri Lanka, SE Asia.
     apiciclara Hardy, 1973: 326 (Stylia). Vietnam.

Genus SPATHULINA Rondani

SPATHULINA Rondani, 1856: 113. Type species: Spathulina sicula Rondani, 1856, orig. des.
acroleuca Schiner, 1868: 268 (Tephritis). Australia (NSW); American Samoa, widesp. Australia (excl. Tas), Bonin Is, Fiji, French Polynesia (Society Is), Guam, Indonesia (Irian Jaya), New Caledonia (New Caledonia), PNG (Bismarck Arch, PNG), Solomon Is, Tonga, Vanuatu, Western Samoa; India, Indonesia, Japan, Philippines, Ryukyu Is, Taiwan, Afrotrop., Palearctic Regs.
     undecimguttata Thomson, 1869: 581 (Trypeta; as 11-guttata). Australia (NSW).
     parca Bezzi, 1913b: 159 (Oxyna). India.
     nigrifemorata de Meijere, 1914b: 220. Indonesia (Java).

Genus SPHENELLA Robineau-Desvoidy

SPHENELLA Robineau-Desvoidy, 1830: 773. Type species: Sphenella linariae Robineau-Desvoidy, 1830 [= Tephritis marginata Fallén, 1814], mon.
SINEVRA Lioy, 1864: 1024. Type species: Tephritis marginata Fallén, 1814, des. Hardy (1977: 130).
fascigera Malloch, 1931h: 391 (Tephritis). NZ (NZ).
novaguineensis Hardy, 1988b: 62. PNG (Bismarck Arch); Indonesia (Irian Jaya), PNG (PNG).
ruficeps Macquart, 1851: 261(288) (Urophora). Australia (Tas); widesp. Australia; New Caledonia (New Caledonia).
     heterura Thomson, 1869: 584 (Trypeta). Australia (NSW).
sinensis Schiner, 1868: 267. China; Indonesia (Irian Jaya), PNG (PNG), widesp. Oriental Reg.
     sinensis Thomson, 1869: 585 (Trypeta). China. [Preocc. Schiner, 1868.]

Genus TEPHRITIS Latreille

TEPHRITIS Latreille, 1804: 196. Type species: Musca arnicae Linnaeus, 1758, des. Cresson, 1914a: 278.
brunnea Hardy & Drew, 1996: 352. Australia (Vic); Australia (Qld).
cometa Loew, 1840b: 157 (Trypeta). "aus der Wiener Gegend" [= Austria]; ?Australasian/Oceanian [= prob. misid.]; USSR, Europe, Middle East.
furcata Hardy & Drew, 1996: 358. Australia (Qld).


TETREUARESTA Hendel, 1928b: 368. Type species: Trypeta obscuriventris Loew, 1873, orig. des.
obscuriventris Loew, 1873b: 313 (Trypeta). Brazil; Fiji [introduced], Hawaiian Is [introduced], New Caledonia (New Caledonia) [introduced], Tonga [introduced]; ?Bolivia, ?Colombia, Cuba, Jamaica, Puerto Rico.

Genus TRUPANEA Schrank

TRUPANEA Guettard, 1762: 170. Unavailable name; group-genus name proposed in a non-binominal work.
TRUPANEA Schrank, 1795: 147. Type species: Trupanea radiata Schrank, 1795 [= Musca stellata Fuessly, 1775], mon.
TRYPANEA Agassiz, 1846b: 380 (unjust. emend. of Trupanea).
TRYPANOIDEA Bryan, 1924: 367 (Tephritis subg.). Type species: Trypeta crassipes Thomson, 1869, orig. des.
alboapicata Malloch, 1931h: 401. NZ (NZ).
amoena Frauenfeld, 1856: 542 (Trypeta). Yugoslavia; Australia (NT, Qld); widesp. Pal., Afrotrop., Orient. Regs.
arboreae Hardy in Hardy & Delfinado, 1980a: 66. Hawaiian Is.
artemisiae Hardy in Hardy & Delfinado, 1980a: 68. Hawaiian Is.
asteria Schiner, 1868: 270 (Tephritis). India & Indonesia (Java); PNG (PNG), Solomon Is.
beardsleyi Hardy in Hardy & Delfinado, 1980a: 69. Hawaiian Is.
bidensicola Hardy in Hardy & Delfinado, 1980a: 72. Hawaiian Is.
bifida Hardy & Drew, 1996: 387. Australia (NSW).
celaenoptera Hardy in Hardy & Delfinado, 1980a: 72. Hawaiian Is.
centralis Malloch, 1931h: 402. NZ (NZ).
completa Malloch, 1931h: 400. NZ (NZ).
crassipes Thomson, 1869: 583 (Trypeta). Hawaiian Is.
cratericola Grimshaw, 1901: 46 (Tephritis). Hawaiian Is.
dempta Hardy in Hardy & Delfinado, 1980a: 78. Hawaiian Is.
denotata Hardy in Hardy & Delfinado, 1980a: 80. Hawaiian Is.
dubautiae Bryan, 1921: 477 (Tephritis). Hawaiian Is.
     dubautiae Terry, 1912: 147. Nomen nudum.
dubia Malloch, 1931h: 401. NZ (NZ).
extensa Malloch, 1931h: 402. NZ (NZ).
fenwicki Malloch, 1931h: 404. NZ (NZ).
glauca Thomson, 1869: 581 (Trypeta). Australia (NSW); widesp. Australia and Pacific; Indonesia (Java).
heronensis Hardy & Drew, 1996: 390. Australia (Qld).
joycei Hardy in Hardy & Delfinado, 1980a: 84. Hawaiian Is.
latinota Hardy, 1988b: 74. PNG (PNG); Indonesia (Irian Jaya).
limpidapex Grimshaw, 1901: 46 (Tephritis). Hawaiian Is.
lipochaetae Hardy in Hardy & Delfinado, 1980a: 88. Hawaiian Is.
longipennis Malloch, 1931h: 398. NZ (NZ).
lyneborgi Hardy, 1970: 131. PNG (Bismarck Arch).
marginalis Hardy in Hardy & Delfinado, 1980a: 90. Hawaiian Is.
megaspila Hardy in Hardy & Delfinado, 1980a: 91. Hawaiian Is.
neoapicalis Hardy, 1989: 530 (n. n. for apicalis Hardy). Hawaiian Islands.
     apicalis Hardy in Hardy & Delfinado, 1980a: 64. Hawaiian Is. [Preocc. Macquart, 1838.]
nigripennis Hardy in Hardy & Delfinado, 1980a: 92. Hawaiian Is.
nigripilosa Hardy in Hardy & Delfinado, 1980a: 48 (Neotephritis). Hawaiian Is.
notata Hardy & Drew, 1996: 393. Australia (Qld); Australia (NSW).
paludosae Hardy in Hardy & Delfinado, 1980a: 51 (Neotephritis). Hawaiian Is.
pantosticta Hardy in Hardy & Delfinado, 1980a: 93. Hawaiian Is.
pekeloi Hardy in Hardy & Delfinado, 1980a: 95. Hawaiian Is.
perkinsi Hardy in Hardy & Delfinado, 1980a: 97. Hawaiian Is.
prolata Hardy & Drew, 1996: 395. Australia (SA); Australia (NSW, Qld, SA, Vic, WA).
pusilla Hardy & Drew, 1996: 397. Australia (WA); Australia (Qld).
queenslandensis Hardy & Drew, 1996: 398. Australia (Qld).
rufa Hardy, 1988b: 79. PNG (PNG).
simplex Malloch, 1932m: 146. French Polynesia (Marquesas).
     simplicissima Malloch, 1929: 413. Nomen nudum.
swezeyi Bryan, 1921: 478 (Tephritis). Hawaiian Is.
unimaculata Malloch, 1931h: 403. NZ (NZ).
vitiosa Foote, 1989: 531 (n. n. for imperfecta Malloch). NZ (NZ).
imperfecta Malloch, 1931h: 403. NZ (NZ). [Preocc. Coquillett, 1902.]
vittigera Malloch, 1931h: 400. NZ (NZ).
watti Malloch, 1931h: 403. NZ (NZ).


XANTHACIURA Hendel, 1914c: 86. Type species: Trypeta chrysura Thomson, 1869, orig. des.
connexionis Benjamin, 1934: 45. USA; Hawaiian Is [introduced; ?established]; Mexico to Nicaragua, Jamaica to Bahamas.

Unplaced species of Tephritidae

albida Walker, 1853: 384 (Trypeta). Australia (SA).
brevivitta Walker, 1865b: 124 (Trypeta). "New Guinea" [= Indonesia (Irian Jaya)].
     brevivittata, error for brevivitta.
diversata Walker, 1865b: 124 (Trypeta). "New Guinea" [= Indonesia (Irian Jaya)].
immaculata Macquart, 1855: 145(125) (Terellia). French Polynesia (Marquesas).
reclusa Hardy, 1987: 334 (Myoleja). Solomon Is.
rufitarsis Macquart, 1847a: 94 [1847b: 110] (Terellia). "Nouvelle-Hollande."

This page last revised 16 December 2011