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39. Family THEREVIDAE

[Originally] by Michael E. Irwin & Leif Lyneborg

Flies of the family Therevidae are nearly worldwide in distribution, occurring in all geographical regions except Antarctica. The Australasian/Oceanian Therevidae are poorly known. A total of 327 species and 25 genera are herein recognized as valid; many additional undescribed genera and species occur in these regions. Some species from these regions have heretofore been assigned to Thereva, Psilocephala Zetterstedt, and Dialineura Rondani. These genera, as defined by Irwin & Lyneborg (1981), are solely Holarctic in distribution and are not found in the Australasian/Oceanian Regions. A single species of Thereva was introduced into Hawai`i from California as a potential biological control agent but did not become established (Evenhuis 1982a); it is included in the catalog. Genera listed in this catalog are highly conservative and many species have temporarily been assigned to the category of "unplaced species." Around 50% (162) of the 327 valid species listed here are placed in the genus Anabarhynchus. The genus Clesthentia is considered not to belong to the family Therevidae. It is listed in this catalog in the family Apsilocephalidae.

Therevid bioecology is poorly understood. Therevids inhabit a wide variety of environments with a dominant proportion occupying arid and semiarid zones. Adults are not considered to be predaceous; most imbibe water and some take nourishment from insect excretions/secretions or plant nectar/exudates. Adults are diurnal and often alight in sunny patches on paths, males usually awaiting passing females. Some adults, especially of the genus Agapophytus, perch on tree trunks while others (e.g., Ectinorhynchus) can be observed hovering in abbreviated swarms above and among vegetation. Some (e.g., Platycarenum and some species of Anabarhynchus) frequent coastal dunes. In arid environments, adults often concentrate around water sources. A few species of therevids have been regarded as mimics of various wasp models (Nicholson 1927).

Ovipositional and egg-hatching sequences have been recorded by Irwin (1976, 1977). Females lay between 25 and 90 eggs (M.E. Irwin, unpubl.). Five instars are known, the last of which can pupate or diapause. Therevids are usually univoltine, but cases where 2 or more years are needed to complete a life cycle are known (K. Spitzer, pers. comm.). The active snakelike larvae usually burrow into and through sandy and sandy-loam soils and litter, moving rapidly underground. They are voracious predators, even cannibals, feeding on a large variety of fossorial arthropods and earthworms, preferring larvae of Coleoptera, Lepidoptera, and Diptera. The pupal stadium lasts for about 10 days, depending on species and environmental conditions.

Members of the family Therevidae play an important, although as yet unascertained, role in the suppression of actual and potential fossorial target-pest larvae in agroecosystems and should be considered as existing or latent biological control agents.

Refs.: Kröber (1912a-d,f,g, 1932, rev.), White (1915, rev.), Mann (1928, 1929, 1933, rev.), Bhatia (1934, biol.), English (1950, larval & pupal keys, biol.), Miller (1950, cat. NZ spp.), Irwin & Lyneborg (1981, keys, rev.).

Genus ACATOPYGIA Kröber

ACATOPYGIA Kröber, 1912a: 117 (key) [1912b: 149]. Type species: Acatopygia pulchella Kröber, 1912, subs. mon. Kröber, 1912b: 149.
SPATULIPALPA Kröber, 1912a: 118 (key) [1912d: 221]. Type species: Spatulipalpa paradoxa Kröber, 1912, des. Irwin & Lyneborg, 1989: 353.
olivacea Winterton, 2007d: 53. Australia (Qld); Australia (NSW).
paradoxa Kröber, 1912d: 221 (Spatulipalpa). Australia (NSW).
pulchella Kröber, 1912b: 149. Australia (NSW); Australia (Qld).

Genus ACTENOMEROS Winterton & Irwin

ACTENOMEROS Winterton & Irwin, in Winterton, et al., 1999b: 280. Type species: Actenomeros corniculaticaudus Winterton & Irwin, 1999, orig. des.
aureilineata Winterton & Irwin in Winterton, et al., 1999a: 295 (Nanexila). Australia (Qld.).
budawang Winterton, 2011: 58. Australia (NSW).
corniculaticaudus Winterton & Irwin, in Winterton, et al., 1999b: 280. Australia (Tas)
intermedia Winterton & Irwin in Winterton, et al., 1999a: 296 (Nanexila). Australia (NSW).
onyx Winterton & Irwin, in Winterton, et al., 1999b: 280. Australia (Tas) .
paradoxa Winterton & Irwin in Winterton, et al., 1999a: 298 (Nanexila). Australia (Qld.).

Genus ACRASPISA Kröber

ACRASPISA Kröber, 1912g: 238, 286. Type species: Acraspisa trifasciata Kröber, 1912, mon.
PSEUDOLOXOCERA Kröber, 1912g: 286. Type species: Pseudoloxocera pallipes Kröber, 1912, mon.
STRATIOBELONALYS Frey, 1934: 309. Type species: Stratiobelonalys handschini Frey, 1934, orig. des.
ACRASPIZA, error for Acraspisa.
handschini Frey, 1934: 309 (Stratiobelonalys). Indonesia (Timor); Indonesia (Java).
nigrinota Mann, 1929: 20. Australia (Qld).
obscuripes Mann, 1929: 21. Australia (Qld).
pallipes Kröber, 1912g: 287 (Pseudoloxocera). Australia (Qld).
parva Mann, 1933: 342 (Parapsilocephala). Australia (Vic); Australia (Qld).
similis Hardy, 1939a: 48. Australia (Qld).
trifasciata Kröber, 1912g: 286. Australia (Qld); Australia (NSW).

Genus ACRASPISOIDES Hill & Winteron

ACRASPISOIDES Hill & Winterton, 2004: 4. Type species: Acraspispoides helviarta Hill & Wiunterton, 2004, orig. des.
helviarta Hill & Winterton, 2004: 4. Australia (NSW).
monticola Winterton, 2007c: 3. Australia (NSW).

Genus ACUPALPA Kröber

ACUPALPA Kröber, 1912a: 118 (key) [1912b: 152]. Type species: Acupalpa rostrata Kröber, 1912, subs. mon. Kröber, 1912b: 152.
ACUTIPALPA, error for Acupalpa.
albimanus Kröber, 1914: 31 (Ecintorhynchus). "Australien"; Australia (NSW, Qld).
    pollinosa Mann, 1929: 25. Australia (Qld).
    polinosa, error for pollinosa .
albitarsa Mann, 1929: 24. Australia (Qld); Australia (NSW).
boharti Winterton, 2011: 43. Australia (WA).
divisa Walker, 1850b: 3 (Dimassus). Not given [?Australia]; Australia (Qld).
    semiflava Mann, 1929: 28. Australia (Qld).
dolichorhyncha Winterton, 2011: 46. Australia (WA).
glossa Winterton, 2011: 43. Australia (Vic).
imitans White, 1915: 28 (Phycus). Australia (Tas, Qld).
irwini Winterton, 2000: 232. Australia (WA).
melanophaeos Winterton, 2011: 52. Australia (WA).
miaboolya Winterton, 2011: 53. Australia (WA).
minuta Winterton, 2011: 55. Australia (WA).
minutoides Winterton, 2011: 56. Australia (WA).
notomelas Winterton, 2011: 57. Australia (WA).
novayamarna Winterton, 2011: 59. Australia (WA).
rostrata Kröber, 1912b: 152. Australia (NSW).
semirufa Mann, 1929: 27. Australia (NSW); Australia (Qld).
westralica Winterton, 2011: 63. Australia (WA).
yalgoo Winterton, 2011: 64. Australia (WA).
yanchep Winterton, 2011: 65. Australia (WA).

Genus AGAPOPHYTUS Guérin

AGAPOPHYTUS Guérin, 1831: pl. 20. Type species: Agapophytus australasiae Guérin, 1831, mon.
adonis Winterton & Irwin, 2001: 474. Australia (Qld); Australia (NSW, Qld).
albobasalis Mann, 1929: 37. Australia (Qld); Australia (NSW, Qld, SA).
    albibasis, error for albobasalis.
albopunctatus Röder, 1885: 139. "Nova-Hollandia"; Australia (NSW, WA).
annamariae Winterton & Irwin, 2001: 477. Australia (WA); Australia (NSW, NT, SA, WA).
antheliogynaion Winterton & Irwin, 2001: 479. Australia (ACT); Australia (ACT, NSW, Qld), PNG (PNG).
asprolepidotos Winterton & Irwin, 2001: 480. Australia (WA).
aterrimus Mann, 1929: 36. Australia (Qld); Australia (ACT, NSW, Qld, SA, Vic).
atrilaticlavius Winterton & Irwin, 2001: 482. Australia (NSW); Australia (NSW, Qld).
australasiae Guérin, 1831: pl. 20. Australia (Qld); Australia (ACT, NSW, Qld).
bicolor Kröber, 1928: 37 (Parapsilocephala). Australia (Qld); Australia (NSW, SA, WA).
biluteus Winterton & Irwin, 2001: 486. Australia (NSW); Australia (ACT, NSW).
borealis Winterton & Irwin, 2001: 487. Australia (WA); Austrlaia (NT, WA).
caliginosus Winterton & Irwin, 2001: 488. Australia (WA).
cerrusus Winterton & Irwin, 2001: 489. Australia (NT); Australia (NT, WA).
chaetohypopion Winterton & Irwin, 2001: 490. PNG (PNG).
chrysosisyrus Winterton & Irwin, 2001: 491. Australia (ACT).
collessi Winterton, 2013: 191. Australia (NT).
decorus Winterton & Irwin, 2001: 492. Australia (Qld).
dieides Winterton & Irwin, 2001: 493. Australia (NSW); Australia (NSW, Qld).
dioctriaeformis Schiner, 1868: 151 (Phycus). Australia (NSW); Australia (ACT, NSW, Qld, Vic).
discolor Winterton & Irwin, 2001: 495. Australia (Qld); Australia (Qld, NT, WA).
eli Winterton & Irwin, 2001: 496. Australia (Qld); Australia (NSW, Qld, Vic).
fenestratum Winterton & Irwin, 2001: 497. Australia (NSW); Australia (NSW, Qld).
galbicaudus Winterton & Irwin, 2001: 498. Australia (ACT); Australia (ACT, NSW).
labifenestellus Winterton & Irwin, 2001: 499. Australia (Qld); Australia (NT, Qld, WA).
laparoceles Winterton & Irwin, 2001: 501. Australia (WA); Australia (NT, WA).
lissohoplon Winterton & Irwin, 2001: 501. Australia (NSW).
lyneborgi Winterton & Irwin, 2001: 502. Australia (NSW); Australia (NSW, Qld).
notozophos Winterton & Irwin, 2001: 503. Australia (SA); Australia (SA, WA).
novaeguineae Winterton & Irwin, 2001: 505. PNG (PNG).
pallidicornis Kröber, 1912a: 124 (Phycus). Australia (NSW); Australia (NSW, Qld).
    flavicornis Mann, 1929: 32. Australia (Qld).
pallidicrus Winterton & Irwin, 2001: 507. Australia (NSW); Australia (ACT, NSW).
palmulus Winterton & Irwin, 2001: 508. Australia (Qld).
paramonovi Winterton & Irwin, 2001: 509. Australia (NSW).
quatiens White, 1915: 27 (Phycus). Australia (Tas).
queenslandi Kröber, 1929a: 417. Australia (Qld); Australia (ACT, NSW, Qld).
    varipennis Mann, 1929: 39. Australia (NSW).
ruficaudus Mann, 1929: 36. Australia (Qld); Australia (NSW, Qld).
    ruficaudatus, error for ruficaudus.
septentrionalis Winterton & Irwin, 2001: 514. Australia (WA); Australia (NT, Qld, WA).
squamosus Mann, 1929: 39. Australia (Qld); Australia (NSW, Qld).
yeatesi Winterton & Irwin, 2001: 516. Australia (WA).
zebra Winterton & Irwin, 2001: 518. Australia (Qld).

Genus ANABARHYNCHUS Macquart

ANABARHYNCHUS Macquart, 1848: 231, 234(71, 74); pl. 3, fig. 7. Type species: Anabarhinchus fasciatus Macquart, 1848, mon.
PLATYCARENUM Kröber, 1912a: 119 (key) [1912f: 243]. Type species: Platycarenum porrectifrons Kröber, 1912 [= Thereva latifrons Macquart, 1848], subs. mon. (Kröber, 1912f: 243).
ANABARYNCHUS. Incorr. orig. spell. of Anabarhynchus (Macquart, 1848: 191 (31).]
ANABARHINCHUS. Incorr. orig. spell. of Anabarhynchus [Macquart, 1848: 192(32)].
ANABARRHYNCHUS, error for Anabarhynchus.
abdominalis Kröber, 1912d: 219. "Australien"; Australia (NSW).
acuminatus Lyneborg, 1992: 23. NZ (NZ).
adornatus Lyneborg, 2001: 79. Australia (NSW).
albellus Lyneborg, 2001: 69. Australia (NSW).
albipennis Lyneborg, 1992: 23. NZ (NZ).
albosetosus Lyneborg, 2001: 176. Australia (ACT); Australia (ACT, Qld).
alpinus Lyneborg, 2001: 172. Australia (Vic).
annulatus Lyneborg, 2001: 177. Australia (WA).
apertus Macquart, 1846: 231(103) (Thereva). "Île de Sydney, dans l'Océanie" [= Australia (NSW)]; Australia (NSW, Qld).
    concolor Walker, 1848: 226 (Thereva). "New Holland."
    rufipes Macquart, 1850: 403(99). Australia, "côte orientale.
    ochropus Thomson, 1869: 477 (Thereva). Australia (NSW).
    aureovillosus Kröber, 1912d: 216. "Australien".
arenarius Lyneborg, 1992: 24. NZ (NZ).
argenteus Mann, 1928: 176. Australia (Qld).
atratus Lyneborg, 1992: 25. NZ (NZ).
atrifemoratus Lyneborg, 2001: 51. Australia (Qld).
atripes Lyneborg, 1992: 25. NZ (NZ).
aureosericeus Kröber, 1932: 136. NZ (NZ).
barrington Lyneborg, 2001: 205. Australia NSW).
bicuspidatus Lyneborg, 2001: 82. Australia (Qld).
bigoti Lyneborg, 2001: 55. Australia (ACT); Australia (ACT, NSW, Qld).
boharti Lyneborg, 2001: 104. Australia (SA).
bohemani Thomson, 1869: 479. Australia (NSW).
boomerang Lyneborg, 2001: 232. Australia (NSW); Australia (NSW, Qld).
brevicornis Lyneborg, 1992: 27. NZ (NZ).
brunninervis Kröber, 1932: 128. NZ (NZ).
busseltonensis Lyneborg, 2001: 127. Australia (WA).
calceatus Schiner, 1868: 149. Australia (NSW); Australia (NSW, Qld).
camiro Lyneborg, 2001: 247. Australia (Qld).
carduus Lyneborg, 2001: 149. Australia (NSW); Australia (NSW, Vic).
cinereus Kröber, 1912d: 215. Australia (NSW).
collessi Lyneborg, 2001: 106. Australia (Qld).
completus Lyneborg, 1992: 29. NZ (NZ).
cretatus Ferguson in Ferguson et al., 2014: 555. Australia (Qld).
curvistylus Lyneborg, 1992: 30. NZ (NZ).
danielsi Lyneborg, 2001: 58. Australia (NSW); Australia (ACT, NSW, Tas).
darembal Ferguson in Ferguson et al., 2014: 558. Australia (Qld).
decipiens Lyneborg, 2001: 95. Australia (NSW); Australia (ACT, NSW, Vic).
dentiphallus Lyneborg, 2001: 197. Australia NSW): Australia (ACT, NSW).
dimidiatus Macquart, 1847a: 50 [1847b: 66] (Thereva). "Nouvelle-Hollande"; Australia (ACT, NSW, Qld, SAS, Tas).
    lateralis Walker, 1852: 161 (Thereva). Not given [?Australia]. [Preocc. Eschscholtz, 1822.]
    validus Mann, 1928: 190. Australia (Tas).
    liepae Irwin & Lyneborg, 1989: 355 (n.n. for lateralis Walker). ?Australia
    calceatus, White, 1916, not Schiner, 1868, misid.
disciphallus Lyneborg, 2001: 53. Australia (Qld).
diversicolor Lyneborg, 1992: 31. NZ (NZ).
doncollessi Ferguson in Ferguson et al., 2012: 57. Australia (NT).
dugdalei Lyneborg, 1992: 32. NZ (NZ).
dysmachiiformis Kröber, 1932: 129. NZ (NZ).
embersoni Lyneborg, 1992: 33. NZ (NZ).
ewamin Ferguson in Ferguson et al., 2012: 61. Australia (Qld).
exiguus Hutton, 1901: 26. NZ (NZ).
    novazealandiae Kröber, 1932: 140 (Neothereva.NZ (NZ).
farinosus Lyneborg, 1992: 34. NZ (NZ).
fasciatus Macquart, 1848: 192(32). "Nouvelle-Hollande"; Australia (NSW, Qld).
    tricolor Walker, 1848: 225 (Thereva). "New Holland."
    inconspicuus Walker, 1852: 160 (Thereva). Australia (NSW).
femoralis Kröber, 1932: 135. "Neuseeland."
    caesius Kröber, 1912d: 219. "Neuseeland."
fenwicki Lyneborg, 1992: 36. NZ (NZ).
ferrugineus Lyneborg, 2001: 151. Australia (WA).
flaviventris Lyneborg, 1992: 37. NZ (NZ).
flavus Mann, 1928: 181. Australia (Qld); Australia (NSW, Qld).
fluviatilis Lyneborg, 1992: 37. NZ (NZ).
fulvipes Macquart, 1850: 404(100). "Nouvelle-Hollande."
furcatus Lyneborg, 2001: 208. Australia (NSW).
fuscoapicatus Lyneborg, 2001: 112. Australia (Tas).
fuscofemoratus Lyneborg, 1992: 38. NZ (NZ).
gascoyne Lyneborg, 2001: 108. Australia (WA); Australia (NT, WA).
gibbsi Lyneborg, 1992: 38. NZ (NZ).
glorious Lyneborg, 2001: 234. Australia (Qld).
granitensis Lyneborg, 2001: 179. Australia (Tas).
grossus Lyneborg, 1992: 39. NZ (NZ).
harrisi Lyneborg, 1992: 40. NZ (NZ).
hayakawai Lyneborg, 1992: 40. NZ (NZ).
helvenacus White, 1915: 44. Australia (Tas); Austrlaia (ACT, NSW, SA, Tas, Vic).
herpenthus Lyneborg, 2001: 173. Australia (WA).
hudsoni Lyneborg, 1992: 41. NZ (NZ).
huttoni Lyneborg, 1992: 42. NZ (NZ).
hyalipennis Macquart, 1846: 232(104) (Thereva). Australia (Tas); Australia (NSW, Qld).
  ssp. hyalipennis s. str. Australia (NSW, Qld, Tas).
    latifrons Macquart, 1850: 469, 473, pl. 9, fig. 14. [1851: 314, 324]. Australia (Tas).
    hebes Walker, 1852: 160 (Thereva). Australia (Tas).
    latyfrons. Incorr. orig. spell. of latifrons [Macquart 1850: 405 (101)].
  ssp. varicincta Bigot, 1860b: 222 (Thereva). New Caledonia (Loyalty Is); New Caledonia (Loyalty Is, New Caledonia), Vanuatu.
iancommoni Ferguson in Ferguson et al., 2014: 560. Australia (Qld).
indistinctus Lyneborg, 1992: 42. NZ (NZ).
innotatus Walker, 1856d: 455 (Thereva). "New Zealand"; NZ (NZ).
    luridus Schiner, 1868: 148. NZ (NZ).
    tristis Kröber, 1932: 123. NZ (NZ). [Preocc. Bigot, 1890.]
    modestus Kröber, 1932: 132. NZ (NZ).
irwini Lyneborg, 2001: 186. Australia (Tas).
kampmeierae Irwin & Lyneborg, 1989: 355 (n.n. for griseus Mann). Australia (WA).
    griseus Mann, 1933: 337. Australia (WA). [Preocc. Kröber, 1911.]
kangaroo Lyneborg, 2001: 180. Australia (SA).
kosciuskoenis Mann, 1933: 338. Australia (NSW); Australia (ACT, NSW, Qld).
kroeberi Lyneborg, 2001: 109. Australia (Qld).
lacustris Lyneborg, 1992: 44. NZ (NZ).
lanatus Lyneborg, 2001: 187. Australia (Tas).
laterepilosus Lyneborg, 1992: 45. NZ (NZ).
latifrons Lyneborg, 2001: 114 (Thereva). "Exotique" [= Australia]; Australia (NSW, Vic).
    arida Walker, 1857: 133 (Thereva). "Australia".
    porrectifrons Kröber, 1912f: 244 (Platycarenum). Australia (Qld).
latus Lyneborg, 1992: 45. NZ (NZ).
limbatinervis Kröber, 1932: 133. NZ (NZ).
loneae Lyneborg, 2001: 153. Australia (NSW).
longipennis Kröber, 1932: 135. NZ (NZ).
longipilosus Lyneborg, 1992: 47. NZ (NZ).
longiseta Ferguson in Ferguson et al., 2014: 564. Australia (Qld).
luctuosus Lyneborg, 2001: 175. Australia (WA).
lyncurium Ferguson in Ferguson et al., 2014: 567. Australia (Qld).
lyneborgi Ferguson & Irwin in Ferguson et al., 2012: 65. Australia (WA).
macfarlanei Lyneborg, 1992: 48. NZ (NZ).
major Lyneborg, 1992: 49. NZ (NZ).
maori Hutton, 1901: 25. NZ (NZ)
    frontalis Kröber, 1932: 137. NZ (NZ).
maritimus Hardy, 1916: 271. Australia (Tas); Australia (ACT, NSW, Qld, SA, Tas, Vic).
mcalpinei Lyneborg, 2001: 226. Australia (NSW).
megalopyge Lyneborg, 1992: 51. NZ (NZ).
megaphallus Lyneborg, 2001: 235. Australia (NSW, Qld).
microphallus Lyneborg, 1992: 52. NZ (NZ).
milo Lyneborg, 2001: 210. Australia (Qld).
misellus Walker, 1835a: 472 (Thereva). Australia (WA).
moffat Lyneborg, 2001: 140. Australia (Qld).
monstruosus Lyneborg, 2001: 243. Australia (Qld).
montanus White, 1915: 42. Australia (Tas); Australia (ACT, NSW, Qld, SA, Tas, Vic).
monteithi Lyneborg, 2001: 168. Australia (Qld).
monticola Lyneborg, 1992: 52. NZ (NZ).
moretonensis Ferguson in Ferguson et al., 2014: 572. Australia (Qld).
multispinosus Lyneborg, 2001: 199. Australia (NSW).
neboensis Ferguson in Ferguson et al., 2014: 575. Australia (Qld).
nebulosus Hutton, 1901: 26. NZ (NZ).
neglectus Kröber, 1932: 132. NZ (NZ).
nigrofemoratus Kröber, 1932: 134. NZ (NZ).
niveus Lyneborg, 2001: 182. Australia (NSW): Australia (ACT, NSW, Qld, SA, Vic).
noosa Lyneborg, 2001: 248. Australia (Qld).
nudifemoratus Macquart, 1846: 231(103) (Thereva). Australia (Tas).
    terrenus White, 1915: 45. Australia (Tas).
oblongicornis Winterton, 2004: 3. Australia (Qld).
occidentalis Lyneborg, 2001: 132. Australia (WA).
oceanus Ferguson & Yeates, 2013: 4. Australia (Vic).
ocypteraeformis Lyneborg, 2001: 193. Australia (NT).
olivaceus Lyneborg, 1992: 56. NZ (NZ).
ornatifrons Kröber, 1914: 59. Australia (Qld); Australia (NSW, Qld).
ostenatus Lyneborg, 1992: 56. NZ (NZ).
pallidus White, 1915: 39. Australia (Tas).
paramonovi Lyneborg, 2001: 229. Australia (NSW).
parilis Ferguson in Ferguson et al., 2012: 69. Australia (Qld).
parkeri Ferguson & Irwinin Ferguson et al., 2012: 73. Australia (WA).
passus White, 1915: 46. Australia (Tas); Australia (ACT, NSW, Tas).
pilbara Ferguson & Irwin in Ferguson et al., 2012: 77. Australia (WA).
planifrons Lyneborg, 2001: 124. Australia (Tas); Australia (NSW, Qld, Tas, Vic).
plumbeoides Lyneborg, 2001: 218. Australia (ACT): Australia (ACT, NSW, Qld, Tas, Vic).
plumbeus Lyneborg, 2001: 220. Australia (NSW); Australia (ACT, NSW, Qld, Tas, Vic).
poona Lyneborg, 2001: 237. Australia (Qld).
postocularis Lyneborg, 1992: 57. NZ (NZ).
projectus Lyneborg, 2001: 158. Australia (NSW); Austraia (ACT, NSW, Qld).
queenslandensis Lyneborg, 2001: 73. Australia (Qld).
quinquevittatum Macquart, 1847a: 49 [1847b: 65] (Thereva). Australia (Tas).
    quinquecellata. Incorr. orig. spell. of quinquevittata (Macquart, 1847a: 104 [1847b: 120]).
ravenshoensis Ferguson in Ferguson et al., 2012: 81. Australia (Qld).
robustus Lyneborg, 1992: 58. NZ (NZ).
rossi Lyneborg, 2001: 160. Australia (NSW); Australia (NSW, Qld).
ruficoxa Lyneborg, 1992: 59. NZ (NZ).
rufobasalis Lyneborg, 1992: 59. NZ (NZ).
rufolateralis Lyneborg, 2001: 203. Australia (WA).
schlingeri Lyneborg, 1992: 60. NZ (NZ).
semioccultus Lyneborg, 2001: 119. Australia (SA).
setulosus Lyneborg, 2001: 133. Australia (WA).
shiptonsflatensis Ferguson in Ferguson et al., 2012: 84. Australia (Qld).
similis Lyneborg, 1992: 61. NZ (NZ).
simplex Lyneborg, 1992: 61. NZ (NZ).
spiniger Lyneborg, 1992: 62. NZ (NZ).
spinosus Lyneborg, 2001: 223. Australia (WA).
spitzeri Lyneborg, 1992: 63. NZ (NZ).
striatifrons Lyneborg, 2001: 195. Australia (Qld).
striatus Lyneborg, 2001: 136. Australia (WA).
stylatus Lyneborg, 2001: 213. Australia (NSW).
tasmanicus Lynebrog, 2001: 75. Australia (Tas).
tauricus Lyneborg, 2001: 85. Australia (Qld).
teddington Lyneborg, 2001: 250. Australia (Qld).
tener Lyneborg, 2001: 76. Australia (NSW); Australia(ACT, NSW).
thoracicus Kröber, 1932: 133. NZ (NZ).
triangularis Lyneborg, 1992: 63. NZ (NZ).
tribulationensis Ferguson in Ferguson et al., 2012: 88. Australia (Qld).
tricoloratus Lyneborg, 1992: 64. NZ (NZ).
tristis Bigot, 1890a: 325. "Australie"; Australia (ACT, NSW, Qld).
turnham Lyneborg, 2001: 230. Australia (Qld).
umbratilis White, 1915: 40. Australia (Tas).
varians Walker, 1852: 161 (Thereva). Australia (Tas).
varipes Macquart, 1847a: 49 [1847b: 65] (Thereva). Australia (Tas); Australia (SA, Tas, Vic, WA).
    conformis Walker, 1848: 225 (Thereva). "New Holland."
    apicalis Walker, 1852: 162 (Thereva). Not given [?Australia]. [Preocc. Wiedemann, 1821.]
waipaensis Ferguson in Ferguson et al., 2012: 91. Australia (Qld).
waitarerensis Lyneborg, 1992: 64. NZ (NZ).
wau Lyneborg, 2001: 246. PNG (PNG).
webbi Lyneborg, 2001: 216. Australia (Tas).
westlandensis Lyneborg, 1992: 65. NZ (NZ).
whitei Lyneborg, 2001: 163. Australia (Tas).
wintertoni Ferguson in Ferguson et al., 2014: 578. Australia (Qld).
wisei Lyneborg, 1992: 66. NZ (NZ).
yarraman Lyneborg, 2001: 226. Australia (Qld).
yeatsei Lyneborg, 2001: 165. Australia (Vic); Australia (ACT, NSW, Vic).
yeppoon Lyneborg, 2001: 87. Australia (Qld).
yuraygirensis Lyneborg, 2001: 119. Australia (NSW).

Genus BELONALYS Kröber

BELONALYS Kröber, 1912a: 117, 124. Type species: Belonalys obscura Kröber, 1912, mon.
BELONALIS, error for Belonalys.
obscura Kröber, 1912a: 125. Australia (NSW).
    gracilenta Kröber, 1912b: 148. Australia (NSW).
occulta White, 1915: 51 (Psilocephala). Australia (Tas).
    occulata, error for occulta.

Genus BONJEANIA Irwin & Lyneborg

LONCHORHYNCHUS White, 1915: 33. Type species: Anabarhynchus nitidifrons Macquart, 1850, des. Mann, 1929: 45. [Preocc. Wiman, 1910.]
BONJEANIA Irwin & Lyneborg, 1989: 356 (n.n. for Lonchorhynchus White).
actuosa White, 1915: 35 (Lonchorhynchus). Australia (ACT, SA, Tas).
affinis Winterton, 2007c: 6. Australia (Tas).
angelikae Winterton & Skevington in Winterton et al., 2000: 10. Australia (NSW).
apluda Winterton, 2007c: 8. Australia (NSW); Australia (NSW, Qld).
argentea Winterton, 2007c: 9. Australia (Vic); australia (ACT, NSW, Vic).
bapsis Winterton, 2007c: 11. Australia (Qld); Australia (NSW, Qld).
clamosis Winterton & Skevington in Winterton et al., 2000: 12. Australia (Qld., NSW, WA).
dynastis Winterton & Skevington in Winterton et al., 2000: 15. Australia (NSW).
flavofemoralis Winterton & Skevington in Winterton et al., 2000: 15. Australia (Qld.).
irwinae Winterton & Skevington in Winterton et al., 2000: 17. Australia (Qld., NSW).
jefferiesi Winterton, 2007c: 14. Australia (ACT).
lambkinae Winterton, 2007c: 16. Australia (NSW).
nitidifrons Macquart, 1850: 405(101) (Anabarhynchus). Australia (Tas); Australia (NSW, Qld).
orphne Winterton & Skevington in Winterton et al., 2000: 19. Australia (Qld., NSW).
segnis White, 1915: 37 (Lonchorhynchus). Australia (Tas); Australia (NSW).
trilineata Winterton & Skevington in Winterton et al., 2000: 22. Australia (NT).
webbi Winterton, 2007c: 18. Australia (Qld).
zwicki Winterton, 2007c: 20. Australia (NSW).

Genus COLLESSIAMA Lambkin

COLLESSIAMA Lambkin in Lambkin & Turco, 2013: 99. Type species: Collessiama narelleae Lambkin, 2013, orig. des.
narelleae Lambkin in Lambkin & Turco, 2013: 100. Australia (Qld); Australia (NSW).
damani Lambkin in Lambkin & Turco, 2013: 107. Australia (Qld); Australia (NSW).

Genus ECTINORHYNCHUS Macquart

ECTINORHYNCHUS Macquart, 1850: 407(103). Type species: Thereva variabilis Macquart, 1846, orig. des.
DIMASSUS Walker, 1850b: 2. Type species: Xylophagus latistria Walker, 1848, des. Irwin & Lyneborg, 1989: 356.
ECTORRHYNCHUS, error for Ectinorhynchus.
brunneus Kröber, 1912b: 158. Australia (NSW).
castaneus Hutton, 1901: 26 (Anabarhynchus). NZ (NZ).
cupreus Hutton, 1901: 27 (Anabarhynchus). NZ (NZ).
furcatus Lyneborg, 1992: 71. NZ (NZ).
latistria Walker, 1848: 130 (Xylophagus). Not given [Australia]; Australia (NSW).
    basipunctatus Walker, 1857: 121 (Xylophagus). "Australia."
levis Mann, 1933: 334. Australia (WA).
micans Hutton, 1901: 27 (Anabarhynchus). NZ (NZ).
pyrrhotelus Walker, 1854: 321 (Agapophytus). Australia (Qld).
    extinctus Walker, 1854: 321 (Agapophytus). "Patria ignota" [= ?Australasian].
    superbus Schiner, 1868: 150 (Ectinorhynchus). Australia (NSW).
    viduus Schiner, 1868: 150 (Ectinorhynchus). Australia (NSW).
    kinbergi Thomson, 1869: 478 (Anabarhynchus). Australia (NSW).
    kinbergia, error for kinbergi.
rufipes Kröber, 1912b: 157. Australia (NSW).
terminalis Walker, 1848: 129 (Xylophagus). "New Holland."
variabilis Macquart, 1846: 230(102) (Thereva). Australia (Tas); Australia (NSW, Qld, Vic, WA).
    terminalis Walker, 1850b: 3 (Dimassus). Australia (NSW). [Preocc. Walker, 1848.]

Genus EUPSILOCEPHALA Kröber

EUPSILOCEPHALA Kröber, 1912a: 119 (key) [1912g: 255]. Type species:Thereva singula Walker, 1848, subs. mon. Kröber, 1912f: 255.
albodorsalis Winterton, 2006: 20. Australia (Qld).
kroeberi Paramonov, 1950c: 528. Australia (Qld).
singula Walker, 1848: 227 (Thereva). Australia (WA); Australia (Qld).
sulphurea Winterton, 2006: 22. Australia (Qld).

Genus EVANSOMYIA Mann

EVANSOMYIA Mann, 1928: 161. Type species: Ectinorrhynchus phyciformis White, 1915, orig. des.
phyciformis White, 1915: 32 (Ectinorhynchus). Australia (Tas); Australia (NSW, Vic).
scutellus Mann, 1933: 335. Australia (Vic).
venusta Erichson, 1842: 272 (Thereva). Australia (Tas).

Genus IRWINIELLA Lyneborg

IRWINIELLA Lyneborg, 1976: 251. Type species: Thereva nuba Wiedemann, 1828, orig. des.
atripes Kröber, 1912f: 254 (Psilocephala). "Neuguinea"; PNG (PNG).
    novaguineensis Kröber, 1929a: 423 (Psilocephala). PNG (PNG).
    bakeri, authors, not Kröber, 1929, misid.
congrua Walker, 1858b: 90 (Thereva). Indonesia (Maluku); Indonesia (Irian Jaya); Indonesia (Sulawesi).
    lanata Doleschall, 1857: 392 (Dasypogon). Indonesia (Maluku). [Preocc. Walker, 1849.]
    conscita Walker, 1861e: 8 (Thereva). Indonesia (Maluku).

Genus JOHNMANNIA Irwin & Lyneborg

MANNIA Paramonov, 1950c: 525. Type species: Mannia tasmanica Paramonov, 1950, orig. des. [Preocc. Dewalque, 1868.]
JOHNMANNIA Irwin & Lyneborg, 1989: 357 (n.n. for Mannia Paramonov).
kosciuszkoensis Lambkin & Recsei, 2005: 19. Australia (NSW).
powerae Lambkin & Recsei, 2005: 9. Australia (Qld); Australia (NSW, Qld).
tasmanica Paramonov, 1950c: 526 (Mannia). Australia (Tas).
     singula, authors, not Walker, 1848, misid.

LAXOTELA Winterton & Irwin, 1999: 300. Type species: Laxotela whitei Winterton & Irwin, 1999, orig. des.
elongata Winterton, 2007a: 59. Australia (Qld).
gaimarii Winterton & Irwin, 1999: 302. Australia (Qld., NSW, WA, SA, ACT).
hauseri Winterton & Irwin, 1999: 304. Australia (NSW, ACT, Vic.).
holstoni Winterton & Irwin, 1999: 305. Australia (WA).
metzi Winterton & Irwin, 1999: 307. Australia (Qld., SA).
ornata Kröber, 1912d: 221 (key) [1912f: 242] (Spatulipalpa). Australia (NSW).
plata Winterton, 2007a: 62. Australia (SA); Australia (ACT, NSW, SA)
whitei Winterton & Irwin, 1999: 309. Australia (Tas.).

Genus MANASTELLA Metz

MANASTELLA Metz in Metz et al., 2003: 10. Type species: Psilocephala tristriata Mann, 1933: 331, orig. des.
caesia Winterton & Lambkin, 2012: 6. Australia (WA); Australia (SA, Vic).
campestris Winterton & Lambkin, 2012: 8. Australia (WA).
canities Winterton & Lambkin, 2012: 9. Australia (WA).
cooloola Winterton & Lambkin, 2012: 10. Australia (Qld).
fumosa Winterton & Lambkin, 2012: 12. Australia (WA).
incompleta Winterton & Lambkin, 2012: 13. Australia (WA).
nubis Winterton & Lambkin, 2012: 14. Australia (Qld).
obscura Winterton & Lambkin, 2012: 15. Australia (Qld).
ocellaris Winterton & Lambkin, 2012: 16. Australia (Qld).
persona Winterton & Lambkin, 2012: 17. Australia (WA).
poecilothorax Winterton & Lambkin, 2012: 18. Australia (WA).
tristriata Mann, 1933: 331 (Psilocephala). Australia (Vic).
umbrapennis Winterton & Lambkin, 2012: 21. Australia (WA).
vasta Winterton & Lambkin, 2012: 22. Australia (WA).
vespera Winterton & Lambkin, 2012: 23. Australia (WA).

Genus MEDOMEGA Winterton & Lambkin

MEDOMEGA Winterton & Lambkin, 2012: 24. Type species: Medomega danielsi Winterton & Lambkin, 2012, orig. des.
averyi Winterton & Lambkin, 2012: 26. Australia (WA).
bailmeup Winterton & Lambkin, 2012: 28. Australia (NT).
chlamydos Winterton & Lambkin, 2012: 29. Australia (WA).
danielsi Winterton & Lambkin, 2012: 30. Australia (Qld).
gigasathe Winterton & Lambkin, 2012: 31. Australia (WA).
nebrias Winterton & Lambkin, 2012: 32. Australia (NSW).

Genus MEGATHEREVA Lyneborg

MEGATHEREVA Lyneborg, 1992: 66. Type species: Bibio bilineata Fabricius, 1775, orig. des.
albopilosa Lyneborg, 1992: 67. NZ (NZ).
atritibia Lyneborg, 1992: 68. NZ (NZ).
bilineatus Fabricius, 1775: 757 (Bibio). "Zelandia.".
    viduus Hudson, 1892, not Walker, 1849, misid.

Genus NANEXILA Winterton & Irwin

NANEXILA Winterton & Irwin in Winterton, et al., 1999a: 244. Type species: Anabarhynchus manni Hardy, 1950, orig. des.
argentiquadris Winterton & Irwin in Winterton, et al., 1999a: 263. Australia (WA, NT).
armeniacum Winterton & Irwin in Winterton, et al., 1999a: 248. Australia (SA, WA).
atricauda Winterton, 2007b: 56. Australia (Qld).
atricostalis Winterton & Irwin in Winterton, et al., 1999a: 292. Australia (Qld.).
carminata Winterton & Irwin in Winterton, et al., 1999a: 252. Australia (SA).
cylomelasma Winterton & Irwin in Winterton, et al., 1999a: 268. Australia (WA, Qld., NT).
danielsi Winterton & Irwin in Winterton, et al., 1999a: 272. Australia (NSW, SA, WA).
furcata Winterton & Irwin in Winterton, et al., 1999a: 255. Australia (SA, WA).
gracilis Mann, 1928: 167 (Taenogera). Australia (Qld.).
jimrodmani Winterton, 2007b: 57. Australia (NSW).
lignyos Winterton & Irwin in Winterton, et al., 1999a: 275. Australia (WA).
ligula Winterton & Irwinin Winterton, et al., 1999a: 276. Australia (WA, SA).
livea Winterton & Irwin in Winterton, et al., 1999a: 278. Australia (Qld, NSW).
lutea White, 1915: 49 (Psilocephala). Australia (Tas).
manni Mann, 1955: 178 (Anabarhynchus). Australia (NSW, Qld., ACT, WA)
nana Winterton & Irwinin Winterton, et al., 1999a: 260. Australia (Qld.).
palassa Winterton & Irwin in Winterton, et al., 1999a: 281. Australia (Qld.).
rufa Kröber, 1912f: 253 (Psilocephala). Australia (NSW).
ruficornis Macquart, 1850: 102 (Anabarhynchus). Australia (Qld.).
spilotis Winterton & Irwin in Winterton, et al., 1999a: 284. Australia (WA, SA).
variabilis Winterton & Irwin in Winterton, et al., 1999a: 286. Australia (WA, NT, NSW, SA, Qld.).
vittata Winterton & Irwin in Winterton, et al., 1999a: 289. Australia (Qld.).

Genus NEODIALINEURA Mann

NEODIALINEURA Mann, 1928: 171. Type species: Neodialineura striatithorax Mann, 1928, orig. des.
ataxia Winterton, 2009: 8. Australia (SA); Australia (Tas, WA).
atmis Winterton, 2009: 11. Australia (WA).
bagdad Winterton, 2009: 11. Australia (Tas).
bifaria Winterton, 2009: 12. Australia (WA).
litura Winterton, 2009: 14. Australia (SA).
nitens White, 1915: 50 (Psilocephala). Australia (Tas); Australia (ACT, NSW, WA).
polygramma Winterton, 2009: 18. Australia (NSW).
saxatilis White, 1915: 52 (Psilocephala). Australia (Tas); Australia (ACT, NSW, Vic).
signum Winterton, 2009: 21. Australia (Vic).
spinosa Winterton, 2009: 23. Australia (WA).
striatithorax Mann, 1928: 172. Australia (Qld).
tessella Winterton, 2009: 26. Australia (ACT).
trichidion Winterton, 2009: 26. Australia (WA).

Genus PARAPSILOCEPHALA Kröber

PARAPSILOCEPHALA Kröber, 1912a: 119 (key) [1912f: 255]. Type species: Parapsilocephala elegans Kröber, 1912, subs. mon. Kröber, 1912f: 255.
OLDENBERGIA Kröber, 1912a: 118 (key) [1912d: 220]. Type species: Oldenbergia frontalis Kröber, 1912, subs. mon. Kröber, 1912d: 220.
ODENBERGIA, error for Oldenbergia.
aggera Mann, 1933: 327 (Psilocephala). Australia (Qld); Australia (NSW).
albofasciata Kröber, 1928: 38. Australia (Qld).
ambocerina Irwin in Irwin & Yeates, 1995: 1311. Australia (NSW); Australia (Qld).
bifasciata White, 1915: 56. Australia (Tas); Australia (ACT, Vic).
duocoloris Mann, 1933: 329 (Psilocephala). Australia (NSW).
elegans Kröber, 1912g: 256. Australia (NSW); Australia (SA, Tas, Vic).
fascifrons Frey, 1934: 311 (Pseudoloxocera). "N. Australien: Kadarri & Barrel's Cr."
frontalis Kröber, 1912d: 220 (Oldenbergia). Australia (NSW); Australia (Qld, Vic).
    curta Kröber, 1913: 30 (Psilocephala). Australia (NSW).

Genus PATANOTHRIX Winterton

PATANOTHRIX Winterton in Winterton et al., 2001: 204. Type species: Patanothrix skevingtoni Winterton, 2001, orig. des.
skevingtoni Winterton in Winterton et al., 2001: 205. Australia (Qld., NSW).
wilsoni Mann, 1933: 342 (Parapsilocephala). Australia (Vic.).

Genus PIPINNIPONS Winterton

PIPINNIPONS Winterton in Winterton et al., 2001: 205. Type species: Pipinnipons kroeberi Winterton, 2001, orig. des.
chauncyvallis Winterton, 2011: 68. Australia (Tas).
fascipennis Kröber, 1928: 36 (Squamopygia). Australia (Qld).
kampmeierae Winterton, 2011: 70. Australia (Qld).
kroeberi Winterton in Winterton, et al., 2001: 205. Australia (NSW); Australia (Qld).
sphecoda Winterton, 2011: 75. Australia (Tas).

Genus SQUAMOPYGIA Kröber

SQUAMOPYGIA Kröber, 1928: 35. Type species: Squamopygia ornata Kröber, 1928, pres. des.
ornata Kröber, 1928: 35. Australia (Qld).

Genus TAENOGERA Kröber

TAENOGERA Kröber, 1912a: 118 (key) [1912b: 150]. Type species: Taenogera longa Kröber, 1912 [= Anabarhynchus nitidus Macquart, 1850], subs. mon. Kröber, 1912b: 151.
brunnea Winterton, 2007b: 62. Australia (Qld).
longa Schiner, 1868: 149 (Anabarhynchus). Australia (NSW).
luteola Winterton, 2007b: 61. Australia (Qld).
nitida Macquart, 1850: 404(100) (Anabarhynchus). Australia, "côte orientale"; Australia (NSW).
    longa Kröber, 1912b: 151. "Australien." [Preocc. Schiner, 1868.]
notatithorax Mann, 1928: 169. Australia (Qld).

Genus TAENOGERELLA Winterton & Irwin

TAENOGERELLA Winterton & Irwin in Winterton, et al., 1999b: 283. Type species: Taenogera nigrapicalis Mann, 1928, orig. des.
elizabethae Winterton & Irwin in Winterton et al., 1999b: 284. Australia (Qld., NSW).
platina Winterton & Irwin in Winterton et al., 1999b: 287. Australia (Qld., NSW, ACT, Vic.).
schlingeri Winterton & Irwin in Winterton et al., 1999b: 288. Australia (WA, SA).
nigrapicalis Mann, 1928: 16 (Taenogera). Australia (NSW, WA, Vic., SA).

Genus THEREVA Latreille

THEREVA Latreille, 1796: 167. Type species: Musca plebeja Linnaeus 1758, subs. mon. Latreille, 1802: 441.
nebulosa Kröber, 1912h: 264. Hawaiian Is [introduced; not established].

Genus VOMERINA Winterton

VOMERINA Winterton, 2007c: 22. Type species: Vomerina humbug Winterton, 2007, orig. des.
comapenis Winterton & Ferguson, 2012: 67. Australia (NSW).
humbug Winterton, 2007c: 22. Australia (NSW).
micoria Winterton & Ferguson, 2012: 71. Australia (NSW).

Unplaced species of Therevidae

funebris Walker, 1865b: 111 (Thereva). "New Guinea" [= Indonesia (Irian Jaya)].
rufiventris Macquart, 1850: 405(101) (Anabarhynchus). "Nouvelle Hollande, cote orientale" [= Australia (Tas)].

This page last revised 20 April 2016