Tipulidae -- a/o cat.

Australasian/Oceanian Diptera Catalog -- Web Version

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[Originally] By Pjotr Oosterbroek

Editor's Note: The original chapter included groups now split into separate families and treated as such in this online version: Limoniidae, Cylindrotomidae, and Pediciidae. The introduction below serves for all these families.

Although phylogenetically one of the most primitive families of Diptera, the Tipulidae is a very successful group comprising ca. 15,000 described species. The majority of the species were described by Alexander in over 1,000 publications. These publications have been indexed to keys, geographical area, and (sub)genera by Oosterbroek & Theowald (1980a,b). More than 2,100 species are known from the regions covered by this catalog.

As might be expected in a group of insects as large and diverse as the Tipulidae, a wide variety of habitats are occupied by the larvae and adults. But considering their popular familiarity as "daddy-longlegs", surprisingly little is known about the species of the Australasian/Oceanian Regions. Larval habitats include freshwater, especially rapidly flowing streams; intertidal zones or brackish waters; all kinds of semiaquatic environments; moss- and algae-covered steep cliffs, which are kept constantly wet; moist to wet cushions of mosses or liverworts; decaying wood or sodden logs in streams; leaf litter or mud such as that found along swamps, marshes, streams, and lakes; sandy or gravelly riversides; dry soils of pasturelands, lawns, steppe, and even deserts; fungi and mycelia; decaying plant materials or fruits in various stages of putrefaction; and leaves of terrestrial higher plants and mosses (mostly externally, rarely internally).

The life cycle of crane flies usually consists of a brief egg stage (1-2 weeks), 4 larval stages, and a short pupal stage (1-2 weeks) before emergence of the short-lived adult. Summer as well as winter diapause of certain stages is often noted and, depending on the species and the environmental conditions, the entire cycle may be as short as 6 weeks or as long as 6 years.

Tipulids are found from the northernmost Arctic lands to equatorial forests and from sea level to about 5,200 meters in elevation. They are usually associated with moist and temperate environments of forested or more open areas such as swamps, marshes, and meadows. Some species can be abundant in localized areas and are therefore of great ecological importance as food for a variety of animals. Larvae of a few species are destructive feeders on the subterranean parts of pasture grasses, seedlings, and crops and may sometimes cause severe commercial losses.

The earliest fossils of Tipulidae are found in Eocene and Oligocene sediments and amber (40-45 mya), but the family is supposedly much older: about mid-Mesozoic. In particular, the Baltic amber fauna can be considered rather diversified, comprising 2 extant genera of Tipulinae and 29 genera of Limoniinae, 25 of which are extant. Mid-Tertiary fossil finds have also yielded tipulid genera, some of which are extant.

An earlier version of this chapter is by Oosterbroek & Jonas (1986), which includes a species index for all the major islands and island groups, secondary references for the taxa listed, and more detailed information on type localities and distribution of species. Works omitted from Oosterbroek & Jonas (1986) are Alexander (1952), and Byers (1982, 1985).
Refs.: Skuse (1890a, 1890b, rev. Australian spp.), Edwards (1923a, rev. NZ spp.; 1928a, rev. Samoan spp.), Alexander (1929d, checklist NZ spp.; 1948f, rev. New Caledonian spp.; 1961c, rev. New Guinean spp.; 1964b, rev. Campbell I spp.; 1972b, rev. Micronesian spp.; 1973a, rev. Kermadec Is spp.), Miller (1950, cat. NZ spp.), Hardy (1960, rev. Hawaiian Is spp.), Dobrotworsky (1968a-1974e, rev. Australian Tipulinae), Oosterbroek & Jonas (1986, cat. Australasian/Oceanian spp.), de Jong (1987, Austrotipula); Fijian Holorusia (Evenhuis, 2006).


ACRACANTHA Skuse, 1890b: 109. Type species: Acracantha sydneyensis Skuse, 1890, des. Alexander, 1920e: 53.
inornata Skuse, 1890b: 113. Australia (WA).
monticola Skuse, 1890b: 112. Australia (NSW).
sydneyensis Skuse, 1890b: 111. Australia (NSW); Australia (ACT, Tas, Vic).
     tasmaniensis Alexander, 1922c: 264. Australia (Tas).
     octava Alexander, 1929a: 50. Australia (Tas).

Genus AUSTROTIPULA Alexander

AUSTROTIPULA Alexander, 1920g: 129. Type species: Pachyrhina hudsoni Hutton, 1900, mon.
hudsoni Hutton, 1900: 24 (Pachyrhina; n. n. for dux Hudson). NZ (NZ).
     dux Hudson, 1895: 293 (Tipula). NZ (NZ). [Preocc. Kirby, 1884.]
     legatus Edwards, 1923a: 332 (Longurio). NZ (NZ).

Genus BRACHYPREMNA Osten Sacken

BRACHYPREMNA Osten Sacken, 1886: 161. Type species: Tipula dispellens Walker, 1861, des. Coquillett, 1910: 516.
tigriventris Alexander, 1922c: 251. Australia (Qld).
waigeuensis Alexander, 1948b: 389. Indonesia (Irian Jaya).


CLYTOCOSMUS Skuse, 1890b: 74. Type species: Clytocosmus helmsi Skuse, 1890, mon.
alexanderi Dobrotworsky, 1968b: 509. Australia (Qld); Australia (NSW).
edwardsi Alexander, 1922b: 158. Australia (Vic).
helmsi Skuse, 1890b: 76. Australia (NSW); Australia (ACT, Vic).
lichwardti Riedel, 1920d: 85. Australia (Qld).
nichollsi Paramonov, 1953f: 51. Australia (Vic).
tillyardi Alexander, 1920b: 184. Australia (NSW).
     skusei Alexander, 1922b: 159 (key) [1922c: 252]. Australia (NSW).


FLABELLIFERA Meigen, 1800: 13. Suppressed by I.C.Z.N., 1963b: 339.
CTENOPHORA Meigen, 1803: 263. Type species: Tipula pectinicornis Linnaeus, 1758, des. Latreille, 1804: 188.

Subgenus PSELLIOPHORA Osten Sacken

PSELLIOPHORA Osten Sacken, 1887: 165 (as gen.). Type species: Tipula laeta Fabricius, 1794, des. Brunetti, 1912b: 290.
ardens Wiedemann, 1820b: 20 [1821a: 20; 1821b: 20]. Indonesia (Java); Indonesia (Maluku); Oriental Reg.
compedita Wiedemann, 1820b: 21 [1821a: 21; 1821b: 21]. Indonesia (Java); Indonesia (Maluku); Oriental Reg.
     competida, error for compedita.


DOLICHOPEZA Curtis, 1825: 62. Type species: Dolichopeza sylvicola Curtis, 1825 [= Tipula albipes Ström, 1768], orig. des.

Subgenus DOLICHOPEZA Curtis

DOLICHOPEZA Curtis, 1825: 62 (as gen.). Type species: Dolichopeza sylvicola Curtis, 1825 [= Tipula albipes Ström, 1768], orig. des.
APEILESIS Macquart, 1846: 136(8) (as gen.). Type species: Apeilesis cinerea Macquart, 1846, mon.
aequalis Theischinger, 1993b: 880. Australia (Qld).
albescens Alexander, 1937: 331. Australia (NSW).
annulipes Skuse, 1890b: 61. Australia (NSW); Australia (Qld, Vic).
anthema Alexander, 1971: 271. Indonesia (Irian Jaya).
asymmetrica Theischinger, 1993b: 874. Australia (NSW); Australia (Qld).
atropos Hudson, 1895: 295 (Tipula). NZ (NZ).
austrocaledonica Alexander, 1948e: 138. New Caledonia (New Caledonia).
ballaratiensis Dobrotworsky, 1974e: 4. Australia (Vic); Australia (Tas).
berrimilla Theischinger, 1993b: 860. Australia (Qld); Australia (NSW).
bibasis Alexander, 1971: 273. Indonesia (Irian Jaya).
bickeli Theischinger, 1993b: 846. Australia (NSW).
boogoo Theischinger, 1993b: 858. Australia (Qld).
brevifurca Skuse, 1890b: 66. Australia (NSW); Australia (Qld, Tas, Vic).
     fuscoradialis Alexander, 1934a: 241. Australia (NSW).
cairnensis Alexander, 1934a: 242. Australia (Vic); Australia (NSW).
cinerea Macquart, 1846: 136(8) (Apeilesis). Australia (Tas); Australia (NSW, Vic).
     longidigitalis Alexander, 1924c: 182. Australia (NSW).
collessi Theischinger, 1993b: 881. Australia (Qld).
corinnaiensis Dobrotworsky, 1974e: 8. Australia (Tas).
cyatheti Alexander, 1934a: 244. Australia (Vic); Australia (Tas).
danbulla Theischinger, 1993b: 868. Australia (Qld).
davidsoni Alexander, 1930a: 137. Australia (NSW); Australia (Qld, Vic).
directa Theischinger, 1993b: 877. Australia (Qld).
distivena Alexander, 1971: 274. Indonesia (Irian Jaya).
dorrigensis Alexander, 1930a: 138. Australia (NSW).
fenwicki Alexander, 1923h: 280. NZ (NZ).
ferox Alexander, 1928g: 337. Australia (Tas).
geometrica Theischinger, 1993b: 848. Australia (Qld); Australia (NSW).
howesi Alexander, 1922k: 204. NZ (NZ).
illingworthi Alexander, 1930a: 136. Australia (Qld).
kongoola Theischinger, 1993b: 869. Australia (NSW).
kraussiana Alexander, 1971: 280. Vanuatu.
kurandensis Alexander, 1937: 332. Australia (Qld).
leonardi Theischinger, 1993b: 860. Australia (Qld).
longifurca Skuse, 1890b: 65. Australia (NSW).
minnamurra Theischinger, 1993b: 862. Australia (NSW).
mongas Alexander, 1971: 276. Indonesia (Irian Jaya).
monticola Skuse, 1890b: 63. Australia (NSW); Australia (Qld).
nephalia Alexander, 1948b: 397. PNG (PNG).
nigrina Dobrotworsky, 1974e: 17. Australia (Tas).
niveitarsis Skuse, 1890b: 62. Australia (NSW); ?Australia (Vic).
nokensis Alexander, 1948b: 393. Indonesia (Irian Jaya).
oresitropha Alexander, 1928d: 55. Australia (NSW).
palliditarsis Alexander, 1928g: 341. Australia (Tas).
pallidula Alexander, 1928g: 338. Australia (NSW).
parvicauda Edwards, 1923a: 330. NZ (NZ).
percuneata Alexander, 1936b: 173. PNG (PNG).
planidigitalis Skuse, 1890b: 65. Australia (NSW).
pygmaea Alexander, 1928g: 344. Australia (Qld).
queenslandica Alexander, 1920e: 55 (brevifurca ssp.). Australia (Qld).
segnis Alexander, 1937: 334. Australia (NSW).
setistyla Alexander, 1971: 277. Indonesia (Irian Jaya).
spetai Theischinger, 1993b: 851. Australia (Qld).
subannulipes Alexander, 1934a: 245. Australia (NSW).
subposticata Alexander, 1928g: 342. Australia (Tas); Australia (Vic).
tayloriana Alexander, 1936b: 171. PNG (PNG).
thiasa Alexander, 1971: 279. Indonesia (Irian Jaya).
thowla Theischinger, 1993b: 875. Australia (NSW).
thysbe Alexander, 1948b: 395. Indonesia (Irian Jaya).
     disseminata Alexander, 1953a: 326 (unnec. n. n. for thysbe Alexander).
tropica Theischinger, 1993b: 849. Australia (Qld).
tyilye Theischinger, 1993b: 865. Australia (NSW).
varipes Skuse, 1890b: 67. Australia (NSW).
victoriae Alexander, 1928g: 340. Australia (Vic); Australia (SA).
yourula Theischinger, 1993b: 863. Australia (Qld).
zborowskii Theischinger, 1999: 503. Australia (Qld).
zenta Theischinger, 1993b: 869. Australia (Qld); Australia (NSW).

Subgenus NESOPEZA Alexander

NESOPEZA Alexander, 1914a: 157 (as gen.). Type species: Dolichopeza gracilis de Meijere, 1911, orig. des.
costalis Brunetti, 1918: 277. India; Indonesia (Maluku).
gracilis de Meijere, 1911a: 60. Indonesia (Java); Indonesia (Maluku); Oriental Reg.
jobiensis Alexander, 1948b: 392. Indonesia (Irian Jaya).
praesultator Alexander, 1948b: 390. Indonesia (Irian Jaya).


HOLORUSIA Loew, 1863b: 276 [1864c: 160]. Type species: Holorusia rubiginosa Loew, 1863 [preocc., = Tipula grandis Bergroth, 1888], mon.
CTENACROSCELIS Enderlein, 1912a: 1. Type species: Ctenacroscelis dohrnianus Enderlein, 1912, orig. des.
aberrans Alexander, 1920d: 108 (Ctenacroscelis). Australia (Qld).
borneensis Brunetti, 1918: 263 (Tipula; n. n. for pallida Walker). "New Guinea"; Indonesia (Irian Jaya), PNG (PNG).
     pallida Walker, 1865a: 105 (Tipula). "New Guinea" [=Indonesia (Irian Jaya)]. [Preocc. Gmelin, 1788.]
conspicabilis Skuse, 1890b: 120. Australia (Qld); PNG (Bismarck Arch).
     ssp. angustilineata Alexander, 1935a: 51 (Ctenacroscelis). PNG (Bismarck Arch).
     ssp. conspicabilis s. str. Australia (Qld).
damuda Evenhuis, 2006: 7. Fiji.
degeneri Alexander, 1978a: 103. Fiji.
fijiensis Alexander, 1921e: 562 (Ctenacroscelis). Fiji.
     vitiana Alexander, 1978a: 104. Fiji.
Alexander, 1935a: 51 (Ctenacroscelis). PNG (Bismarck Arch).
illex Alexander, 1947c: 588 (Ctenacroscelis). PNG (PNG).
lepida Alexander, 1924b: 45 (Ctenacroscelis). Fiji.
mamare Evenhuis, 2006: 14. Fiji.
novaeguineae de Meijere, 1913b: 312 (Tipula). Indonesia (Irian Jaya); Indonesia (Maluku); "Borneo."
palauensis Alexander, 1940: 199 (Ctenacroscelis). Belau.
pallescens Edwards, 1926a: 141 [1926b: 171] (Ctenacroscelis). Indonesia (Maluku).
percontracta Alexander, 1947c: 587 (Ctenacroscelis). PNG (PNG).
picturata Evenhuis, 2006: 16. Fiji.
praepotens Wiedemann--not Australasian/Oceanian.
schlingeri Evenhuis, 2006: 18. Fiji.
tabogo Podenas & Evenhuis, 2009: 63. Fiji.
umbrina Wiedemann, 1828: 49 (Tipula). Indonesia (Irian Jaya); PNG (PNG); Oriental Reg.
     infindens Walker, 1860a: 92 (Tipula). Indonesia (Sulawesi).
     congruens Walker, 1861b: 231 (Tipula). Indonesia (Irian Jaya).
     sumatrana Enderlein, 1912a: 5 (Ctenacroscelis). Indonesia (Sumatra).
     incongruens, error for congruens.
vanua Podenas & Evenhuis, 2009: 63. Fiji.
walkeriana Alexander, 1924b: 47 (Ctenacroscelis). Fiji.
     walkeri, error for walkeriana.


ISCHNOTOMA Skuse, 1890b: 115. Type species: Tipula serricornis Macquart, 1846 [= Tipula eburnea Walker, 1848], des. Alexander, 1920e: 53.
eburnea Walker, 1848: 69 (Tipula). Australia (Tas); Australia (NSW, Qld, SA, Vic).
     serricornis Macquart, 1846: 141(13) (Tipula). Australia (Tas). [Preocc. Zetterstedt, 1838.]
     albovariegata Macquart, 1855: 36(16) (Tipula). Australia (SA).
episema Alexander, 1924c: 184. Australia (NSW); Australia (Vic).
fastidiosa Skuse, 1890b: 69 (Tanypremna). Australia (NSW).
fuscobasalis Alexander, 1937: 329. Australia (Vic).
goldfinchi Alexander, 1924f: 520. Australia (NSW); Australia (Vic).
immaculipennis Alexander, 1924c: 186. Australia (NSW).
par Walker, 1856d: 446 (Ptilogyna). Australia (NSW); Australia (ACT, SA, Tas, Vic).
prionoceroides Alexander, 1922c: 266. Australia (Tas).
rubriventris Macquart, 1846: 142(14) (Tipula). Australia (Tas); Australia (ACT, NSW, Vic).
rubroabdominalis Alexander, 1922c: 267. Australia (Tas).
rufiventris Macquart, 1846: 141(13) (Tipula). Australia (Tas); Australia (NSW, SA, Vic).
     nigricaudata Macquart, 1855: 36(16) (Tipula). Australia (SA).
     abnormalis Alexander, 1922c: 265 (Acracantha). Australia (Tas).
     subabnormalis Alexander, 1928d: 53. Australia (NSW).
scutellumnigra Alexander, 1924c: 185. Australia (NSW); Australia (Tas).
skuseana Alexander, 1928g: 353. Australia (Tas).
tarwinensis Alexander, 1928g: 356. Australia (Vic); Australia (NSW).
terminata Alexander, 1928g: 355. Australia (Tas).

Genus LEPTOTARSUS Guérin-Méneville

LEPTOTARSUS Guérin-Méneville, 1831: pl. 20, fig. 1 [1838: 286]. Type species: Leptotarsus macquartii Guérin-Méneville, 1831, mon.

Subgenus AUROTIPULA Alexander

AUROTIPULA Alexander, 1924j: 305. Type species: Macromastix flavoscapus Alexander, 1922, orig. des.
apertus Edwards, 1923a: 338 (Macromastix). NZ (NZ).
atroflavus Alexander, 1922g: 235 (Macromastix). NZ (NZ).
auroatrus Edwards, 1923a: 336 (Macromastix). NZ (NZ).
bivittatus Edwards, 1923b: 629 (Macromastix). NZ (NZ).
brevitarsis Edwards, 1923a: 337 (Macromastix). NZ (NZ).
clarus Kirby, 1884b: 271 (Tipula). Not given [NZ (NZ)].
     clara Walker, 1848: 71 (Tipula). Nomen nudum.
     clara Hutton, 1874: 166 (Tipula). Nomen nudum.
dux Kirby, 1884b: 270 (Tipula). Not given [NZ (NZ)].
     dux Walker, 1848: 71 (Tipula). Nomen nudum.
     dux Hutton, 1874: 166 (Tipula). Nomen nudum.
ferruginosus Edwards, 1923a: 338 (Macromastix). NZ (NZ).
     ssp. ferruginosus s. str. NZ (NZ).
     ferruginea, error for ferruginosus.
     ssp. ruapehuensis Alexander, 1923b: 208 (Macromastix). NZ (NZ).
flavoscapus Alexander, 1922g: 234 (Macromastix). Not given [?NZ (NZ)].
neali Oosterbroek, 1989: 57 (n. n. for tenera Hutton). NZ (NZ).
     tenera Hutton, 1900: 29 (Tipula). NZ (NZ). [Preocc. Müller, 1764.]
occlusus Edwards, 1924b: 162 (Macromastix). NZ (NZ).
orion Hudson, 1895: 294 (Tipula). NZ (NZ).
subtener Alexander, 1922e: 524 (Macromastix). NZ (NZ).
     pseudotenera, error for subtener.

Subgenus BREVICERA Miller

HUDSONIA Edwards, 1923a: 351 (as gen.). Type species: Tipula heterogama Hudson, 1913, mon. [Preocc. Hood, 1893.]
BREVICERA Miller, 1945: 72 (as gen.; n. n. for Hudsonia).
aenigmaticus Alexander, 1926b: 547 (Hudsonia). NZ (NZ).
heterogamus Hudson, 1913: 68 (Tipula). NZ (NZ).
waitakerensis Alexander, 1952: 435 (Longurio). NZ (NZ).

Subgenus CHLOROTIPULA Alexander

CHLOROTIPULA Alexander, 1924j: 305. Type species: Tipula viridis Walker, 1856, orig. des.
albistigma Edwards, 1923a: 340 (Macromastix). NZ (NZ).
elongatus Edwards, 1923b: 630 (Macromastix). NZ (NZ).
holochlorus Nowicki, 1875: 9 (Tipula). NZ (NZ).
     ssp. angustior Alexander, 1923g: 212 (Macromastix). NZ (NZ).
     ssp. holochlorus s. str. NZ (NZ).
     holocrea, error for holochlorus.
virescens Edwards, 1923a: 339 (Macromastix). NZ (NZ).
viridis Walker, 1856d: 445 (Tipula). NZ (NZ).

Subgenus HABROMASTIX Skuse

HABROMASTIX Skuse, 1890b: 93 (as gen.). Type species: Habromastix cinerascens Skuse, 1890, des. Alexander, 1920e: 53.
bulburinensis Dobrotworsky, 1974b: 28. Australia (Qld).
cinerascens Skuse, 1890b: 94 (Habromastix). Australia (NSW); Australia (ACT).
cunninghamensis Dobrotworsky, 1974b: 23. Australia (Qld).
heroni Alexander, 1922c: 263 (Habromastix). Australia (NSW); Australia (Qld).
hilli Alexander, 1922j: 588 (Habromastix). Australia (Vic).
     sublateralis Alexander, 1922j: 588 (Habromastix hilli ssp.). Australia (Vic).
luciae Alexander, 1948g: 392 (Habromastix). PNG (PNG).
novellus Alexander, 1928g: 349 (Habromastix). Australia (Qld); Australia (NSW).
ornatipes Skuse, 1890b: 95 (Habromastix). Australia (NSW).
parallelus Alexander, 1920e: 58 (Habromastix). Australia (Qld); Australia (NSW).
pergrandis Alexander, 1922j: 587 (Habromastix). Australia (Qld).
remotus Walker, 1848: 73 (Tipula). "New Holland"; Australia (Qld).
similior Alexander, 1923j: 258 (Habromastix heroni ssp.). Australia (NSW).
terraereginae Alexander, 1920e: 58 (Habromastix). Australia (Qld); Australia (NSW).
     robinsoni Alexander, 1924d: 375 (Habromastix). Australia (Qld).
     platyxantha Alexander, 1924d: 376 (Habromastix). Australia (Qld).

Subgenus LEPTOTARSUS Guérin-Méneville

LEPTOTARSUS Guérin-Méneville, 1831: pl. 20, fig. 1 [1838: 286] (as gen.). Type species: Leptotarsus macquartii Guérin-Méneville, 1831, mon.
SEMNOTES Westwood, 1876a: 501. Type species: Semnotes imperatoria Westwood, 1876, des. Pierre, 1926: 46.
ardrossanensis Dobrotworsky, 1972b: 22. Australia (SA).
clavatus Macquart, 1850: 318(14) (Tipula). Australia (Tas); Australia (ACT, NSW, Qld, Vic).
     nigrithorax Macquart, 1850: 319(15) (Tipula). Australia (Tas).
     cinereipleurus Alexander, 1929e: 243. Australia (Qld).
     rufosternatus Alexander, 1944: 4 (clavatus ssp.). Australia (Vic).
coolgardiensis Dobrotworsky, 1972b: 23. Australia (WA).
ducalis Westwood, 1876a: 502 (Semnotes). "Australi boreali"; Australia (NSW, Qld).
     ssp. bancrofti Alexander, 1929e: 248 (Semnotes). Australia (Qld); Australia (NSW).
     ssp. ducalis s. str. "Australi boreali"; Australia (NSW).
     ssp. gloria Alexander, 1929e: 247 (Semnotes). Australia (Qld).
esperanceiensis Dobrotworsky, 1972b: 23. Australia (WA).
fletcherensis Dobrotworsky, 1972b: 35. Australia (SA); Australia (NSW, Qld).
fraucai Dobrotworsky, 1972b: 37. Australia (Qld).
imperatorius Westwood, 1876a: 502 (Semnotes). Australia (Vic); Australia (ACT, NSW, Qld).
kalamundaensis Dobrotworsky, 1972b: 26. Australia (WA).
macquartii Guérin-Méneville, 1831: pl. 20, fig. 1 [1838: 286]. Australia (NSW); Australia (ACT, Qld, Vic).
     ssp. flavolateralis Alexander, 1944: 4. Australia (ACT).
     ssp. macquartii s. str. Australia (NSW); Australia (ACT, Qld, Vic).
regificus Alexander, 1922c: 262 (Semnotes). Australia (Qld).
scutellaris Skuse, 1890b: 107. Australia (NSW).
tricinctus Walker, 1848: 73 (Tipula). Australia (Qld).
     caesareus Alexander, 1929e: 245. Australia (Qld).
trivittatus Skuse, 1890b: 108. Australia (WA).

Subgenus MACROMASTIX Osten Sacken

MACROTHORAX Jaennicke, 1867: 319 (as gen.). Type species: Macrothorax ornatus Jaennicke, 1867 [= Tipula costalis Swederus, 1787], mon. [Preocc. Lioy, 1864.]
MACROMASTIX Osten Sacken, 1887: 185 (as gen.; n. n. for Macrothorax).
albicollis Alexander, 1924c: 188 (Macromastix). Australia (NSW).
albipedis Hynes, 1993: 79. New Caledonia (New Caledonia).
albiplagia Alexander, 1923b: 202 (Macromastix). NZ (NZ).
     ssp. albiplagia s. str. NZ (NZ).
     ssp. obliteratus Alexander, 1923b: 203 (Macromastix). NZ (NZ).
alexanderi Edwards, 1923a: 343 (Macromastix). NZ (NZ).
alfie Theischinger in Kitching & Theischinger, 1996: 143. Australia (Qld).
amissionis Alexander, 1952: 436 (Longurio). NZ (NZ).
angusticosta Alexander, 1923a: 110 (Macromastix). NZ (NZ).
arenarius Alexander, 1950b: 937 (Macromastix). Australia (WA).
atridorsum Alexander, 1922d: 314 (Macromastix). NZ (NZ).
     harrisi Alexander, 1922f: 98 (Macromastix). NZ (NZ).
aurantioceps Alexander, 1924c: 190 (Macromastix). Australia (NSW).
     subnitidus Alexander, 1924c: 192 (Macromastix aurantioceps ssp.). Australia (NSW).
barringtoniensis Dobrotworsky, 1974c: 48. Australia (NSW).
binnaburrae Alexander, 1951b: 576 (Macromastix). Australia (Qld).
binotatus Hutton, 1900: 32 (Macromastix). NZ (NZ).
brachycerus Alexander, 1978a: 100. New Caledonia (New Caledonia).
brisbaneiensis Dobrotworsky, 1974c: 49. Australia (Qld).
caledonianus Alexander, 1934f: 443 (Macromastix). New Caledonia (New Caledonia).
campbelli Alexander, 1923b: 207 (Macromastix). NZ (NZ).
cinereus Edwards, 1923a: 345 (Macromastix). NZ (NZ).
clitellarius Alexander, 1930a: 139 (Macromastix). Australia (Vic).
cockerellae Alexander, 1929f: 85 (Macromastix). New Caledonia (New Caledonia).
collessi Dobrotworsky, 1974c: 50. Australia (Qld).
constrictus Skuse, 1890b: 134 (Macromastix). Australia (NSW); Australia (Qld, Vic).
     clarkianus Alexander, 1930a: 142 (Macromastix). Australia (Vic).
costalis Swederus, 1787: 286 (Tipula). Australia (Tas).
     pacificus Erichson, 1842: 270 (Macromastix). Australia (Tas).
     macquartianus Alexander, 1924c: 187 (Macromastix). Australia (Tas).
     costata, error for costalis.
dispar Walker, 1835a: 468 (Megistocera). "New Holland"; Australia (ACT, NSW, Qld, Vic).
     dimidiatus Westwood, 1835f: 451 (Megistocera). "Australasi ."
     limbipennis Macquart, 1838a: 64(60) (Megistocera). "Patrie inconnue" [Australia].
     ornatus Jaennicke, 1867: 320 (Macromastix). "Australia."
cubitalis Edwards, 1923a: 347 (Macromastix). NZ (NZ).
decoratus Edwards, 1923a: 342 (Macromastix). NZ (NZ).
dichroithorax Alexander, 1920g: 133 (Macromastix). NZ (NZ).
dorrigensis Alexander, 1924c: 189 (Macromastix). Australia (NSW).
fergusoni Alexander, 1924c: 192 (Macromastix). Australia (NSW); Australia (ACT, Vic).
flavidipennis Alexander, 1923b: 201 (Macromastix). NZ (NZ).
fucatus Hutton, 1900: 31 (Macromastix). NZ (NZ).
fumibasis Edwards, 1923a: 344 (Macromastix). NZ (NZ).
gargettensis Dobrotworsky, 1974c: 52. Australia (Qld).
glabristylus Hynes, 1993: 81. New Caledonia (New Caledonia).
glaucocapillus Alexander, 1952: 437 (Longurio). NZ (NZ).
greyanus Alexander, 1922f: 96 (Macromastix). NZ (NZ).
hackeri Alexander, 1920e: 62 (Macromastix). Australia (Qld); Australia (NSW).
halteratus Alexander, 1923a: 109 (Macromastix). NZ (NZ).
helmsi Skuse, 1890b: 132 (Macromastix). Australia (NSW); Australia (SA, Tas).
hudsonianus Alexander, 1922g: 237 (Macromastix). NZ (NZ).
     hudsonia, error for hudsonianus.
humilis Skuse, 1890b: 136 (Macromastix). Australia (Vic); Australia (NSW, SA, Tas).
huttoni Edwards, 1923a: 342 (Macromastix). NZ (NZ).
igniceps Alexander, 1928g: 358 (Macromastix). Australia (Tas); Australia (NSW, Vic).
incertus Edwards, 1923a: 348 (Macromastix). NZ (NZ).
intermedius Alexander, 1922g: 236 (Macromastix). NZ (NZ).
longioricornis Alexander, 1923a: 108 (Macromastix). NZ (NZ).
lunatus Hutton, 1900: 32 (Macromastix). NZ (NZ).
     ssp. fuscolateratus Alexander, 1922g: 236 (Macromastix). NZ (NZ).
     ssp. lunatus s. str. NZ (NZ).
     luna, error for lunatus.
luteicostus Alexander, 1924c: 193 (Macromastix). Australia (NSW).
luteisubcostatus Alexander, 1934a: 240 (Macromastix). Australia (Vic).
mastersi Skuse, 1890b: 133 (Macromastix). Australia (NSW); Australia (ACT, Vic).
mathewsi Alexander, 1930a: 141 (Macromastix). Australia (WA).
mesocerus Alexander, 1922i: 572 (Macromastix). NZ (NZ).
minor Edwards, 1923a: 346 (Macromastix). NZ (NZ).
minutissimus Alexander, 1923b: 205 (Macromastix). NZ (NZ).
mixtus Hynes, 1993: 82. New Caledonia (New Caledonia).
monstratus Alexander, 1924f: 522 (Macromastix). NZ (NZ).
montanus Hutton, 1900: 31 (Macromastix). NZ (NZ).
mutabilis Alexander, 1928g: 360 (Macromastix). Australia (Tas).
nigropolitus Alexander, 1928g: 359 (Macromastix). Australia (NSW).
noelianus Alexander, 1978a: 102. New Caledonia (New Caledonia).
novocaledonicus Alexander, 1929f: 83 (Macromastix). New Caledonia (New Caledonia).
     productifrons Alexander, 1945: 236 (Macromastix). New Caledonia (New Caledonia).
     mouicola Alexander, 1951b: 578 (Macromastix). New Caledonia (New Caledonia).
     fulvithorax Alexander, 1978a: 101. New Caledonia (New Caledonia).
obliquus Edwards, 1923a: 348 (Macromastix). NZ (NZ).
obscuripennis Kirby, 1884b: 271 (Tipula). Not given [NZ (NZ)].
     obscuripennis Walker, 1848: 71 (Tipula). Nomen nudum.
     obscuripennis Hutton, 1874: 166 (Tipula). Nomen nudum.
obscurirostris Skuse, 1890b: 135 (Macromastix). Australia (NSW).
ohakunensis Alexander, 1923b: 204 (Macromastix). NZ (NZ).
opifex Alexander, 1929a: 53 (Macromastix). Australia (NSW).
pallidistigma Alexander, 1922f: 95 (Macromastix). NZ (NZ).
pallidus Hutton, 1900: 31 (Macromastix). NZ (NZ).
pedestris Alexander, 1939: 221 (Macromastix). NZ (NZ).
pseudoaurantioceps Dobrotworsky, 1974c: 55. Australia (NSW).
pseudotortilis Dobrotworsky, 1974c: 53. Australia (NSW).
repletus Alexander, 1948f: 368 (Macromastix). New Caledonia (New Caledonia).
risbeci Alexander, 1934f: 441 (Macromastix). New Caledonia (New Caledonia).
rufibasis Alexander, 1922f: 97 (Macromastix). NZ (NZ).
rufiventris Edwards, 1923a: 347 (Macromastix). NZ (NZ).
sessilis Alexander, 1924d: 379 (Macromastix). NZ (NZ).
setivena Alexander, 1929a: 55 (Macromastix). Australia (Vic); Australia (Tas).
simillimus Alexander, 1924f: 521 (Macromastix). NZ (NZ).
sinclairi Edwards, 1923a: 350 (Macromastix). NZ (NZ).
spinastylus Hynes, 1993: 84. New Caledonia (New Caledonia).
submancus Alexander, 1923f: 112 (Macromastix). NZ (NZ).
submontanus Edwards, 1923a: 346 (Macromastix). NZ (NZ).
subobsoletus Alexander, 1926b: 547 (Macromastix). NZ (NZ).
subvittatus Alexander, 1939: 222 (Macromastix). NZ (NZ).
tamborineiensis Dobrotworsky, 1974c: 54. Australia (Qld).
tapleyi Alexander, 1923b: 206 (Macromastix). NZ (NZ).
tenuifrons Alexander, 1926b: 546 (Macromastix). NZ (NZ).
tortilis Alexander, 1920e: 61 (Macromastix). Australia (Qld); Australia (NSW).
     brevipetiolatus Alexander, 1922c: 268 (Macromastix). Australia (NSW).
variegatus Edwards, 1923a: 341 (Macromastix). NZ (NZ).
verreauxi Alexander, 1929a: 52 (Macromastix). Australia (Tas).
vittatus Edwards, 1923a: 343 (Macromastix). NZ (NZ).
vulpinus Hutton, 1881: 16 (Megistocera). NZ (NZ).
zeylandiae Alexander, 1920g: 131 (Macromastix). NZ (NZ).

Subgenus MAORITIPULA Alexander

MAORITIPULA Alexander, 1924j: 306. Type species: Macromastix maori Alexander, 1920, orig. des.
hudsoni Alexander, 1924j: 306 (Macromastix; n. n. for nebulosa Hudson). NZ (NZ).
     nebulosa Hudson, 1920: 276 (Tipula). NZ (NZ). [Preocc. Bellardi, 1859.]
     hudsonella Alexander, 1964c: 282 (unnec. n. n. for hudsoni Alexander).
maori Alexander, 1920g: 130 (Macromastix). NZ (NZ).

Subgenus PHYMATOPSIS Skuse

PHYMATOPSIS Skuse, 1890b: 97 (as gen.). Type species: Phymatopsis nigrirostris Skuse, 1890, mon.
albidipes Alexander, 1929a: 50 (Phymatopsis). Australia (NSW).
brevipalpis Alexander, 1920d: 105 (Phymatopsis). Australia (Qld).
brevirostratus Alexander, 1922c: 261 (Phymatopsis). Australia (NSW).
flavopygialis Alexander, 1920e: 60 (Macromastix). Australia (Qld).
     flavopygalis, error for flavopygialis.
intricatus Alexander, 1928d: 54 (Phymatopsis). Australia (NSW); Australia (Qld).
nigrirostris Skuse, 1890b: 98 (Phymatopsis). Australia (NSW); Australia (Qld).
nigrolimbatus Alexander, 1929a: 51 (Phymatopsis). Australia (NSW).
nigrosubcostatus Alexander, 1924d: 378 (Macromastix). Australia (NSW).
periplocus Alexander, 1928d: 53 (Phymatopsis). Australia (NSW); Australia (Qld).
tenuirostris Alexander, 1924d: 377 (Macromastix). Australia (NSW).
tonnoiranus Alexander, 1928g: 351 (Phymatopsis). Australia (Tas).
walpoleiensis Dobrotworsky, 1974a: 12. Australia (WA).


PSEUDOLEPTOTARSUS Alexander, 1920e: 57. Type species: Leptotarsus liponeura Alexander, 1920, mon.
liponeura Alexander, 1920e: 57. Australia (Qld); Australia (ACT, NSW).
nigrinus Dobrotworsky, 1972b: 41. Australia (NSW); Australia (Qld).

Genus MEGISTOCERA Wiedemann

MAEKISTOCERA Wiedemann, 1820b: 41 [1821a: 41; 1821b: 41]. Type species: Tipula filipes Fabricius, 1805, des. Macquart, 1838a: 63(59) (for Megistocera).
MEGISTOCERA Wiedemann, 1828: 55. Type species: Tipula filipes Fabricius, 1805, des. Macquart, 1838a: 63(59).
[The name Megistocera is used here in the interest of stability and common usage. Suspension of I.C.Z.N. rules are required to validate this name over Maekistocera, which has priority.]
MOEGISTOCERA Macquart, 1834a: 113 (unjust. emend. of Maekistocera).
filipes Fabricius, 1805: 25 (Tipula). "Guinea"; widesp. Australasian/Oceanian Regs; widesp. Afrotrop., Oriental Regs.
     ssp. fuscana Wiedemann, 1820b: 29 [1821a: 29; 1821b: 29] (Nematocera). Indonesia (Java); Australia (Qld), Indonesia (Maluku), PNG (Bismarck Arch, PNG), Solomon Is; widesp. Oriental Reg.


PALES Meigen, 1800: 14. Suppressed by I.C.Z.N., 1963b: 339.
NEPHROTOMA Meigen, 1803: 262. Type species: Tipula dorsalis Fabricius, 1782, mon.
PACHYRHINA Macquart, 1834a: 88. Type species: Tipula crocata Linnaeus, 1758, des. Westwood, 1840: 128.
PACHYRINA. Incorr. orig. spell. of Pachyrhina (Macquart, 1834a: 88).
PACHYRRHINA, error for Pachyrhina.
atrohirsuta Alexander, 1973b: 262. Indonesia (Irian Jaya).
augustana Alexander, 1973b: 264. PNG (Bougainville I).
australasiae Skuse, 1890b: 126 (Pachyrrhina). Australia (Qld); Australia (NSW, NT), Lord Howe I.
boyesi Alexander, 1969: 259. PNG (PNG).
buruensis Edwards, 1926a: 143 [1926b: 173]. Indonesia (Maluku).
byersina Alexander, 1973b: 267. PNG (Bougainville I).
colorata Walker, 1865c: 131 (Pachyrrhina). Indonesia (Irian Jaya); Indonesia (Maluku).
dimidiata de Meijere, 1904b: 90 (Pachyrrhina). "Neu-Guinea"; Indonesia (Irian Jaya), PNG (Bismarck Arch, PNG); Indonesia (Java).
flavoposticata Alexander, 1936b: 169. PNG (PNG).
fumiscutellata Alexander, 1935a: 54. PNG (Bismarck Arch).
guttipleura Alexander, 1936d: 2. PNG (Bougainville I).
iavana Wiedemann--not Australasian/Oceanian.
javensis Doleschall, 1856: 406 (Tipula). Indonesia (Java); Indonesia (Maluku); widesp. Oriental Reg.
     doleschalli Osten Sacken, 1881a: 399 (Pachyrrhina; unnec. n. n. for javensis Doleschall).
     fallax de Meijere, 1904b: 90 (Pachyrrhina). Indonesia (Java).
kaviengensis Alexander, 1936b: 170. PNG (Bismarck Arch).
kraussiana Alexander, 1978b: 141. Solomon Is.
laffooni Alexander, 1950b: 938. Vanuatu.
lordhowensis Dobrotworsky, 1974d: 70. Lord Howe I.
melanura Osten Sacken, 1881a: 401 (Pachyrrhina). PNG (PNG).
nigroannulata Wulp, 1885a: 8 (Pachyrrhina). Indonesia (Maluku).
opima Alexander, 1924b: 49. Solomon Is.
solomonis Alexander, 1924b: 48. Solomon Is.
     ssp. guadalcanarana Alexander, 1936d: 4. Solomon Is.
     ssp. malaitana Alexander, 1936d: 4. Solomon Is.
     ssp. solomonis s. str. Solomon Is.
speculata de Meijere, 1913b: 314 (Pachyrrhina). Indonesia (Irian Jaya); PNG (Bismarck Arch, PNG).
toxopei Edwards, 1926a: 142 [1926b: 172]. Indonesia (Maluku).
tripartita Walker, 1861b: 231 (Pachyrhina). Indonesia (Irian Jaya).
walkeri Oosterbroek, 1986: 49 (n. n. for tenuis Walker). "New Guinea."
     tenuis Walker, 1865a: 106 (Pachyrhina). "New Guinea" [= Indonesia (Irian Jaya)]. [Preocc. Loew, 1863.]


PLATYPHASIA Skuse, 1890b: 84. Type species: Platyphasia princeps Skuse, 1890, mon.
eximia Alexander, 1928g: 348. Australia (NSW).
pictonensis Dobrotworsky, 1971b: 14. Australia (Tas).
princeps Skuse, 1890b: 85. Australia (NSW).
rawlinsoni Dobrotworsky, 1971b: 9. Australia (NSW).
regina Alexander, 1922j: 586. Australia (NSW); Australia (Qld).
tasmaniensis Dobrotworsky, 1971b: 12. Australia (Tas).
wilsoni Alexander, 1929a: 49. Australia (Vic).

Genus PTILOGYNA Westwood

PTILOGYNA Westwood, 1835c: 280 [1835f: 448]. Type species: Tipula ramicornis Walker, 1835, mon.

Subgenus CTENOGYNA Macquart

CTENOGYNA Macquart, 1838a: 46(42) (as gen.). Type species: Ctenogyna bicolor Macquart, 1838, mon.
EUPLUSIOMYIA Alexander, 1928g: 345 (Plusiomyia subg.). Type species: Plusiomyia nasalis Alexander, 1924, orig. des.
bicolor Macquart, 1838a: 47(43) (Ctenogyna). "Inconnue, mais vraisemblablement de l'Australasie"; Australia (ACT, NSW, Qld, Vic).
     optima Alexander, 1928g: 345 (Plusiomyia). Australia (?Vic).
nasalis Alexander, 1924c: 183 (Plusiomyia). Australia (Qld); Australia (NSW).

Subgenus PLUSIOMYIA Skuse

PLUSIOMYIA Skuse, 1890b: 86 (as gen.). Type species: Plusiomyia olliffi Skuse, 1890, des. Alexander, 1920e: 53.
PHACELODOCERA Enderlein, 1912a: 26 (as gen.). Type species: Ptilogyna flabellifera Loew, 1851, orig. des.
clarki Alexander, 1928g: 346 (Plusiomyia). Australia (WA).
felix Alexander, 1922c: 257 (Plusiomyia). Australia (Tas).
     felixi, error for felix.
gracilis Walker, 1848: 37 (Pedicia). Not given [Australia]; Australia (ACT, NSW, Qld, SA, Vic).
     longipedalis Westwood, 1876a: 503 (Ozodicera). "Australi ."
     ssp. gracilis s. str. Not given [Australia]; Australia (NSW, Qld).
     ssp. spectabilis Skuse, 1890b: 90 (Plusiomyia; as sp.). Australia (Vic); Australia (ACT, NSW, SA).
herroni Alexander, 1948f: 370 (Phacelodocera). New Caledonia (New Caledonia).
inornata Skuse, 1890b: 92 (Plusiomyia). Australia (NSW); Australia (Vic).
kraussiana Alexander, 1966: 135 (Plusiomyia). New Caledonia (New Caledonia).
leucoplagia Alexander, 1957: 103 (Plusiomyia). Australia (Qld); Australia (NSW).
lineata Skuse, 1890b: 91 (Plusiomyia). Australia (WA).
mackerrasi Alexander, 1928d: 52 (Plusiomyia). Australia (NSW).
margaritae Alexander, 1948e: 138 (Phacelodocera). New Caledonia (New Caledonia).
mathesonae Dobrotworsky, 1971a: 20. Australia (Vic).
minor Alexander, 1922c: 258 (Plusiomyia). Australia (Qld).
necopina Alexander, 1922c: 260 (Plusiomyia). Australia (Tas).
neocaledonica Alexander, 1948f: 368 (Plusiomyia). New Caledonia (New Caledonia).
neogama Alexander, 1944: 1 (Plusiomyia). Australia (NSW).
olliffi Skuse, 1890b: 89 (Plusiomyia). Australia (NSW); Australia (ACT, Vic).
pandoxa Alexander, 1922c: 255 (Plusiomyia). Australia (Tas).
spissigrada Alexander, 1922c: 259 (Plusiomyia). Australia (Tas).
tasmaniensis Alexander, 1922c: 254 (Phacelodocera). Australia (Tas).
tripectinata Alexander, 1922c: 257 (Plusiomyia). Australia (Tas).
wellsi Alexander, 1950b: 935 (Plusiomyia). Australia (Qld).
westralis Dobrotworsky, 1971a: 31. Australia (WA).

Subgenus PTILOGYNA Westwood

PTILOGYNA Westwood, 1835c: 280 [1835f: 448] (as gen.). Type species: Tipula ramicornis Walker, 1835, mon.
cheesmanae Alexander, 1947c: 585. PNG (PNG).
     macquartii Loew, 1851a: 392 [unnec. n.n. for specimen misid. as fuliginosa Say by Macquart, 1838a: 50(46)]. "Amerique septentrionale" [error, = Australia; see Osten Sacken, 1878: xvi, 222].
minima Alexander, 1922c: 254. Australia (NT).
ramicornis Walker, 1835a: 469 (Tipula). "New Holland"; Australia (NSW, Qld).
     marginalis Westwood, 1835c: 280 [1835f: 449]. "Nov Hollandi ."
     picta Schiner, 1868: 38. Australia (NSW).

Genus SCAMBONEURA Osten Sacken

SCAMBONEURA Osten Sacken, 1882b: 95. Type species: Scamboneura dotata Osten Sacken, 1882, mon.
curtistyla Alexander, 1971: 270 (Nephrotoma). Indonesia (Irian Jaya).
nigrodorsalis Alexander, 1947c: 593. PNG (PNG).
quadrata de Meijere--not Australasian/Oceanian.
tagensis Alexander, 1961c: 488. Indonesia (Irian Jaya).
vittifrons Walker, 1860b: 144 (Limnobia). Indonesia (Maluku).

Genus TIPULA Linnaeus

TIPULA Linnaeus, 1758: 585. Type species: Tipula oleracea Linnaeus, 1758, des. Latreille, 1810: 442.

Subgenus ACUTIPULA Alexander

ACUTIPULA Alexander, 1924g: 11. Type species: Tipula gaboonensis Alexander, 1920, orig. des.
diclava Alexander, 1922j: 589. "Northern Australia."
leptoneura Alexander, 1922c: 269. Australia (NT).

Subgenus INDOTIPULA Edwards

INDOTIPULA Edwards, 1931a: 81. Type species: Tipula walkeri Brunetti, 1911, orig. des.
gurneyana Alexander, 1978a: 106. PNG (Bougainville I).
laffooniana Alexander, 1971: 282. PNG (Bougainville I).
melanodonta Alexander, 1978b: 142. Solomon Is.
querella Alexander, 1948g: 398. PNG (PNG).
serritergata Alexander, 1947c: 592. PNG (PNG).

Subgenus PAPUATIPULA Alexander

PAPUATIPULA Alexander, 1935a: 52. Type species: Tipula novaebrittaniae Alexander, 1935, orig. des.
artifex Alexander, 1948b: 400. PNG (PNG).
consiliosa Alexander, 1971: 284. Indonesia (Irian Jaya).
cyclopica Alexander, 1948g: 396. Indonesia (Irian Jaya).
divergens de Meijere, 1913b: 312. Indonesia (Irian Jaya).
gressittiana Alexander, 1971: 286. Indonesia (Irian Jaya).
inseparata Young, 1987: 164. PNG (PNG).
koiari Young, 1987: 164. PNG (PNG).
leucosticta Alexander, 1934f: 444. PNG (PNG); PNG (Bismarck Arch).
     venusta, Riedel, 1921, not Walker, 1848, misid.
lieftincki Alexander, 1971: 287. Indonesia (Irian Jaya).
meijereana Alexander, 1935c: 115 (n. n. for dentata de Meijere). Indonesia (Irian Jaya); PNG (PNG).
     dentata de Meijere, 1913b: 313. PNG (PNG). [Preocc. Meigen, 1838.]
melanotis Alexander, 1948b: 398. PNG (PNG).
nigrita Young, 1987: 167. PNG (PNG).
nokicola Alexander, 1953c: 747. Indonesia (Irian Jaya).
novaebrittaniae Alexander, 1935a: 53. PNG (Bismarck Arch).
obediens Alexander, 1947c: 590. PNG (PNG).
omissinervis de Meijere, 1906b: 71 (Tanypremna). Indonesia (Irian Jaya); PNG (PNG).
oneili Young, 1987: 168. PNG (PNG).
pedicioides Alexander, 1948b: 393. PNG (PNG).
pensilis Alexander, 1971: 290. Indonesia (Irian Jaya).
satirica Alexander, 1971: 292. Indonesia (Irian Jaya).
staryi Alexander, 1971: 294. Indonesia (Irian Jaya).
strictistyla Alexander, 1971: 296. Indonesia (Irian Jaya).
surcularia Alexander, 1947c: 591. Indonesia (Irian Jaya).
toxopeina Alexander, 1971: 297. Indonesia (Irian Jaya).
wibleae Young, 1987: 170. PNG (PNG).

Subgenus TIPULODINA Enderlein

TIPULODINA Enderlein, 1912a: 30 (as gen.). Type species: Tipulodina magnicornis Enderlein, 1912, orig. des.
cinctipes de Meijere--not Australasian/Oceanian.
curtissima Alexander, 1948b: 399. Solomon Is.
fumifinis Walker, 1860b: 145. Indonesia (Maluku).
     longicornis Doleschall, 1858: 79. Indonesia (Maluku). [Preocc. Linnaeus, 1758.]
     schummeli Brunetti, 1911b: 247 (n. n. for longicornis Doleschall).

Unplaced species of Tipula Linnaeus

demeijerei Edwards, 1915b: 393. Indonesia (Irian Jaya).
tropica de Meijere, 1913b: 311. PNG (PNG); Indonesia (Irian Jaya, Maluku).
vanewrighti Oosterbroek, 1986: 49 (n. n. for lateralis Walker). "NW coast of Australia" [prob. = Australia (WA)].
     lateralis Walker, 1848: 70. "NW coast of Australia." [Preocc. Meigen, 1818.]


ZELANDOTIPULA Alexander, 1922e: 522 (Holorusia subg.). Type species: Tipula novarae Schiner, 1868, orig. des.
fulva Hutton, 1900: 26 (Tipula). NZ (NZ).
novarae Schiner, 1868: 37 (Tipula). NZ (NZ).
     senex Walker, 1848: 71 (Tipula). Nomen nudum.
     senex White & Butler, 1874: 27 (Tipula). NZ (NZ).
otagana Alexander, 1922e: 523 (Holorusia). NZ (NZ).