Family CHIRONOMIDAE

 

Members of the family Chironomidae, commonly called non-biting midges, are found worldwide in a wide range of habitats where the immature stages are an important component of aquatic ecosystems. Approximately, 5,000 species in over 300 genera are known from fossil and living forms. Adults of extant forms are short-lived but can be found in large mating swarms. Adults have been observed to feed on honeydew; other plant exudates may also serve as food sources. Larvae can be found in nearly every moist or aquatic habitat including the periphery of oceans. Some are free-living, but others produce tube-like cases composed of fine substrate. Chironomid larvae feed primarily on decaying organic matter and small plants and animals; others are totally carnivorous.


Certain groups or species are characteristic of certain aquatic environments, which makes the family an excellent indicator group of these habitats. Many chironomids are used in reconstructing paleoclimatic and paleoecological conditions in lentic and lotic habitats. Due to the abundance of extant genera and species that have been identified from numerous paleoenvironmental studies, it was not feasible to include all of them in this catalog. Some of these records have been included in the chironomid chapter of the Palearctic catalog (Ashe & Cranston, 1990) and the reader is referred to this work for more information on this family. Where species have not been included in previous catalogs, every attempt has been made to include named species here.


Chironomidae are abundant in the fossil record. Many are known from amber deposits where they have been recorded from the Cretaceous of Siberia (Zherichin & Sukacheva, 1973; Kalugina, 1976, 1980b), Canada (Boesel, 1937), and New Jersey (Grimaldi, 1990); the Miocene of Saxony (Zherichin, 1978), Myanmar (Schlee & Glockner, 1978), and Mexico (Hurd et al., 1962); the Eocene of China (Hong, 1981) and the Caucasus (Zherichin, 1978); the Oligocene of the Dominican Republic (Poinar, 1992); and the Eocene/Oligocene of the Baltic region (Meunier, 1904a,c,d). The oldest amber chironomid fossil described thus far is Libanochlites neocomicus from the Lower Cretaceous of Lebanon (Brundin, 1976). Jurassic compression fossils have been described from eastern Asia by Kalugina in Kalugina & Kovalev (1985) and Zhang (1989b), and pupae of unnamed tanypodines were described and illustrated by Jell & Duncan (1986: 173­77) from the Lower Cretaceous deposits of Koonwarra, South Gippsland, Australia. Fujiyama & Nomura (1986: 11) recorded an unidentified specimen of this family from the Miocene deposits of Hachiya, Japan.


According to Borkent (1993), the undetermined "mesotendipedid" [= Chironomapterinae] recorded from the Cretaceous Santana Formation of Brazil by Grimaldi (1990) and Maisey (1991: 434) is actually a chironomid and not a chaoborid. Szadziewski (1988: 12) pointed out that one of two Oligocene Sicilian amber specimens identified as Dasypogon by Guérin-Méneville (1838a: 170) is most likely a chironomid.

Ref.: Meunier (1904c, review of Baltic amber taxa); Ashe (1983, catalog of world genera).

 

Genus AMBERASPINUS Evenhuis

ASPINUS Hong, 1981: 52. Type species: Aspinus orientalus Hong, 1981, by original designation. [Preoccupied by Brandorff, 1973.] New synonymy.
AMBERASPINUS Evenhuis, 1994: in this work (new replacement name for Aspinus Hong). Type species: Aspinus orientalus Hong, 1981, automatic.

amblopteres Hong, 1981: 57 (Aspinus). PA: China (Eocene) [A].
ambropteres. Incorrect original spelling of amblopteres (Hong, 1981: 6).
orientalus Hong, 1981: 6, 53 (Aspinus). PA: China (Eocene) [A].
orientales. Incorrect original spelling of orientalus (Hong, 1981: 53).
stenopteres Hong, 1981: 54 (Aspinus). PA: China (Eocene) [A].

 

Genus BYTHOMYIA Zhang

BYTHOMYIA Zhang, 1989c: 155. Type species: Bythomyia oryctes Zhang, 1989, by original designation.

oryctes Zhang, 1989c: 155. PA: China (Miocene) [C].

 

Genus CAMPTOCLADIUS Wulp

*CAMPTOCLADIUS Wulp, 1874: 133. Type species: Tipula byssina Schrank, 1803 [= Tipula stercorarius De Geer, 1776], by subsequent designation of Coquillett (1910: 518).

flexuosus Meunier, 1904c: 221 [1904d: 232]. PA: Baltic Region (Eocene/Oligocene) [A].
sinuosus Meunier, 1904c: 221 [1904d: 232]. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus CHIRONOMOPSIS Handlirsch

CHIRONOMOPSIS Handlirsch, 1906a: 632. Type species: Chironomus arrogans Giebel, 1856, by monotypy.
[In the original description of Chironomopsis, Chironomus extinctus Westwood was questionably included, making the type species Chironomus arrogans by monotypy.]

arrogans Giebel, 1856: 250 (Chironomus). PA: UK (England) (Lower Cretaceous) [C].
extincta Westwood in Brodie, 1845: 121, pl. 4, fig. 5 (Chironomus). PA: UK (England) (Lower Cretaceous) [C].

 

Genus CHIRONOMUS Meigen

TENDIPES Meigen, 1800: 17. Suppressed by I.C.Z.N. (1963: 339).
*CHIRONOMUS Meigen, 1803: 260. Type species: Tipula plumosa Linnaeus, 1758, by subsequent designation of Latreille (1810: 442).
PALAEOLYCUS Etheridge & Olliff, 1890: 11. Type species: Palaeolycus problematicus Etheridge & Olliff, 1890, by monotypy. [Preoccupied by Marck, 1863.]

abietarius Meunier, 1904c: 203 [1904d: 214]. PA: Baltic Region (Eocene/Oligocene) [A].
aequalis Keilbach, 1982: 351. Nomen nudum.
amelanderi Arnaud, 1966: 162 (new replacment name for sepultus Melander).

sepultus Melander, 1949: 16. NE: USA (Oligocene) [C]. [Preoccupied by Heer, 1849.]

anchora Keilbach, 1982: 352. Nomen nudum.
aquisextanus Théobald, 1937a: 347. PA: France (Oligocene) [C].
berendtii Keilbach, 1982: 351. Nomen nudum.
bicalcaratus Keilbach, 1982: 352. Nomen nudum.
bitramus Keilbach, 1982: 352. Nomen nudum.
bituminosus Heyden, 1870: 247. PA: Germany (Oligocene) [C].
brevirostris Giebel, 1856: 251. PA: Baltic Region (Eocene/Oligocene) [A].
caliginosus Meunier, 1904c: 264 [1904d: 275]. PA: Baltic Region (Eocene/Oligocene) [A].
uliginosus. Incorrect original spelling of caliginosus Meunier, 1904c: 202 [1904d: 213].
coarctatus Keilbach, 1982: 351. Nomen nudum.
cognatus Keilbach, 1982: 351. Nomen nudum.
conformis Keilbach, 1982: 351. Nomen nudum.
conjunctus Keilbach, 1982: 351. Nomen nudum.
depletus Scudder, 1877a: 744. NE: USA (Eocene) [C].
elegantulus Meunier, 1904c: 199 [1904d: 210]. PA: Baltic Region (Eocene/Oligocene) [A].
erosus Keilbach, 1982: 352. Nomen nudum.
escheri Giebel, 1856: 252. PA: Baltic Region (Eocene/Oligocene) [A].
excellens Keilbach, 1982: 351. Nomen nudum.
gaudini Heer, 1864: 394. PA: Germany (Oligocene) [C].
gracilicornis Keilbach, 1982: 352. Nomen nudum.
haustus Meunier, 1912c: 190 [1912d: 362]. AF: Tanzania (Pleistocene/Holocene) [K].
inclusus Meunier, 1912c: 190 [1912d: 361]. AF: Tanzania (Pleistocene/Holocene) [K].
incrassatus Keilbach, 1982: 351. Nomen nudum.
inglorius Meunier, 1904c: 201 [1904d: 212]. PA: Baltic Region (Eocene/Oligocene) [A].
kirklandi Sublette, 1969: 119. NE: USA (Pliocene/Pleistocene) [C].
lacunus Meunier, 1904c: 205 [1904d: 216]. PA: Baltic Region (Eocene/Oligocene) [A].
lacus Meunier, 1904c: 205 [1904d: 216]. PA: Baltic Region (Eocene/Oligocene) [A].
lanceatus Keilbach, 1982: 352. Nomen nudum.
leptocerus Keilbach, 1982: 352. Nomen nudum.
leucomelas Gistl, 1831: 247. NT: Brazil (Holocene) [K].
longicornis Keilbach, 1982: 352. Nomen nudum.
longipalpus Keilbach, 1982: 352. Nomen nudum.
meticulosus Meunier, 1904c: 200 [1904d: 211]. PA: Baltic Region (Eocene/Oligocene) [A].
meunieri Arnaud, 1966: 162 (new replacment name for sepultus Meunier, 1912). AF: Tanzania (Pleistocene/Holocene) [K].
sepultus Meunier, 1912c: 190 [1912d: 362]. AF: Tanzania (Pleistocene/Holocene) [K]. [Preoccupied by Heer, 1849.]

meyeri Heer, 1849: 188. PA: Baltic Region (Eocene/Oligocene) [A].
microcephalus Giebel, 1856: 251. PA: Baltic Region (Eocene/Oligocene) [A].
minimus Hong in Hong et al., 1974: 124. PA: China (Eocene) [A]. [Preoccupied by Meigen, 1818; no new replacement name proposed here.]
monilis Keilbach, 1982: 352. Nomen nudum.
obsoletus Heer, 1849: 190. PA: Switzerland (Miocene) [C].
oeningensis Heer, 1849: 189. PA: Switzerland (Miocene) [C].
paludosus Meunier, 1904c: 204 [1904d: 215]. PA: Baltic Region (Eocene/Oligocene) [A].
patens Scudder, 1877a: 744. NE: USA (Eocene) [C]. [Preoccupied by Walker, 1856; no new replacement name proposed here.]
pausatus Melander, 1949: 14. NE: USA (Oligocene) [C].
pliocenicus Piton, 1936: 23. Nomen nudum.
primaevus Melander, 1949: 13. NE: USA (Oligocene) [C].
primitivus Mani, 1945: 62. OR: India (Miocene) [A]. [Preoccupied by Johannsen, 1912; no new replacement name proposed here.]
pristinus Melander, 1949: 13. NE: USA (Oligocene) [C].
proterus Melander, 1949: 14 NE: USA (Oligocene) [C].
requiescens Melander, 1949: 15. NE: USA (Oligocene) [C].
scopa Keilbach, 1982: 351. Nomen nudum.
scopula Keilbach, 1982: 351. Nomen nudum.
scudderiellus Cockerell, 1916c: 98. NE: USA (Oligocene) [C].
separatus Keilbach, 1982: 351. Nomen nudum. [Preoccupied by Walker, 1856.]
sepultus Heer, 1849: 190. PA: Croatia (Miocene) [C].
serresi Théobald, 1937a: 347. PA: France (Oligocene) [C].
spectabilis Keilbach, 1982: 351. Nomen nudum.
stylatus Keilbach, 1982: 352. Nomen nudum.
styliger Keilbach, 1982: 352. Nomen nudum.
subobscurus Meunier, 1904c: 202 [1904d: 213]. PA: Baltic Region (Eocene/ Oligocene) [A].
tenebricosus Meunier, 1904c: 203 [1904d: 214]. PA: Baltic Region (Eocene/ Oligocene) [A].
tenebrosus Meunier, 1904c: 199 [1904d: 210]. PA: Baltic Region (Eocene/Oligocene) [A].
tenuicornis Keilbach, 1982: 352. Nomen nudum.
uliginosus Meunier, 1904c: 204 [1904d: 215]. PA: Baltic Region (Eocene/Oligocene) [A].
umbraticus Meunier, 1904c: 200 [1904d: 211]. PA: Baltic Region (Eocene/ Oligocene) [A].
umbrosus Meunier, 1904c: 201 [1904d: 212]. PA: Baltic Region (Eocene/Oligocene) [A].
ungulatus Keilbach, 1982: 351. Nomen nudum.
ungulinus Keilbach, 1982: 351. Nomen nudum.
vagabundus Meunier, 1904c: 198 [1904d: 209]. PA: Baltic Region (Eocene/ Oligocene) [A].
venerabilis Etheridge & Olliff, 1890: 7. AU: Australia (Pliocene) [C].
problematicus Etheridge & Olliff, 1890: 11 (Palaeolycus). AU: Australia (Pliocene) [C].
verticillatus Keilbach, 1982: 351. Nomen nudum.
?Unidentified sp.-PA: Japan (Pliocene/Pleistocene) [C] (Fujiyama & Iwao, 1975: 45).

 

Genus CLINOTANYPUS Kieffer

*CLINOTANYPUS Kieffer, 1913a: 157. Type species: Clinotanypus fuscosignatus Kieffer, 1913, by subsequent designation of Sublette & Sublette (1965: 145).

vagans Zhang, 1989a: 327. PA: China (Miocene) [C].

 

Genus COELOCHIRONOMA Zhang, Zhang, Liu & Shangguan

COELOCHIRONOMA Zhang et al., 1989: 24. Type species: Coelochironoma xantha Zhang et al., 1989, by original designation.

xantha Zhang et al., 1989: 24. PA: China (Upper Jurassic) [C].

 

Genus CORYNOCERA Zetterstedt

*CORYNOCERA Zetterstedt, 1837: 65 [1838: 856]. Type species: Corynocera ambigua Zetterstedt, 1838, by monotypy.
DRYADOTANYTARSUS Anderson, 1938: 320. Nomen nudum.
DRYADOTANYTARSUS Anderson, 1943: 174. Type species: Dryadotanytarsus edentula Anderson, 1943 [= Corynocera ambigua Zetterstedt, 1838], by monotypy.

*ambigua Zetterstedt, 1838: 856. PA: Sweden; Holarctic; PA: Denmark (Pleistocene) [C].
edentula Anderson, 1943: 174 (Dryadotanytarsus). PA: Denmark (Pleistocene) [C].
duffi Deevey, 1955: 313 (Dryadotanytarsus). AU: New Zealand (Pleistocene) [C].

 

Genus CRETODIAMESA Kalugina

CRETODIAMESA Kalugina, 1976: 88 (79). Type species: Cretodiamesa taimyrica Kalugina, 1976, by original designation.

taimyrica Kalugina, 1976: 89 (90). PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus CRICOTOPIELLA Meunier

CRICOTOPIELLA Meunier, 1916a: 283. Type species: Cricotopiella rostrata Meunier, 1916, by monotypy.

rostrata Meunier, 1916a: 284. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus CRICOTOPUS Wulp

*CRICOTOPUS Wulp, 1874: 132. Type species: Chironomus tibialis Meigen, 1804, by subsequent designation of Coquillett (1910: 528).

abiegnus Meunier, 1904c: 212 [1904d: 223]. PA: Baltic Region (Eocene/Oligocene) [A].
alluvionis Meunier, 1904c: 207 [1904d: 218]. PA: Baltic Region (Eocene/Oligocene) [A].
ambiguus Meunier, 1904c: 211 [1904d: 222]. PA: Baltic Region (Eocene/Oligocene) [A].
amniculus Meunier, 1904c: 207 [1904d: 218]. PA: Baltic Region (Eocene/Oligocene) [A].
antiquus Meunier, 1904c: 206 [1904d: 217]. PA: Baltic Region (Eocene/Oligocene) [A].
coniferus Meunier, 1904c: 214 [1904d: 225]. PA: Baltic Region (Eocene/Oligocene) [A].
crassicornis Meunier, 1904c: 205 [1904d: 216]. PA: Baltic Region (Eocene/Oligocene) [A].
delicatus Meunier, 1904c: 208 [1904d: 219]. PA: Baltic Region (Eocene/Oligocene) [A].
dilapsus Meunier, 1904c: 212 [1904d: 223]. PA: Baltic Region (Eocene/Oligocene) [A].
exstinctus Meunier, 1904c: 215 [1904d: 226]. PA: Baltic Region (Eocene/Oligocene) [A].
extinctus, error for exstinctus Meunier.
insolitus Meunier, 1904c: 213 [1904d: 224]. PA: Baltic Region (Eocene/Oligocene) [A].
minutissimus Meunier, 1904c: 216 [1904d: 227]. PA: Baltic Region (Eocene/ Oligocene) [A].
minutulus Meunier, 1916a: 281. PA: Baltic Region (Eocene/Oligocene) [A].
minutus Meunier, 1904c: 216 [1904d: 227]. PA: Baltic Region (Eocene/Oligocene) [A].
*motatrix Linnaeus, 1758: 587 (Tipula). PA: "Europa"; Holarctic; NE: USA (Undetermined Tertiary) [T].
motilator, error for motatrix.
nemorivagus Meunier, 1904c: 215 [1904d: 226]. PA: Baltic Region (Eocene/ Oligocene) [A].
paganus Meunier, 1904c: 213 [1904d: 224]. PA: Baltic Region (Eocene/Oligocene) [A].
parvulus Meunier, 1916a: 282. PA: Baltic Region (Eocene/Oligocene) [A].
permutabilis Meunier, 1904c: 209 [1904d: 220]. PA: Baltic Region (Eocene/ Oligocene) [A].
pulchellus Meunier, 1904c: 210 [1904d: 221]. PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Meigen, 1830; no new replacement name proposed here.]
pygmaeus Meunier, 1904c: 208 [1904d: 219]. PA: Baltic Region (Eocene/Oligocene) [A].
robustus Meunier, 1904c: 209 [1904d: 220]. PA: Baltic Region (Eocene/Oligocene) [A].
saltuosus Meunier, 1904c: 214 [1904d: 225]. PA: Baltic Region (Eocene/Oligocene) [A].
variabilis Meunier, 1904c: 210 [1904d: 221]. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus DIAMESA Meigen

*DIAMESA Meigen in Gistl, 1835: 66. Type species: Diamesa cinerella Meigen, 1835, by monotypy.
[The genus-group name Diamesa was originally proposed in synonymy with Lestremia (in Cecidomyiidae) by Meigen (1830: 308), but was not validated thereafter; thus, that non-use of Diamesa allows it to be considered a nomen nudum. See Ashe (1983: 20) for details on the nomenclatural history of this name.]

extincta Melander, 1949: 12. NE: USA (Oligocene) [C].

 

Genus ELECTROTENIA Kalugina

ELECTROTAENIA Kalugina, 1974: 49. Nomen nudum.
ELECTROTENIA Kalugina, 1980b: 91. Type species: Electrotenia brundini Kalugina, 1980, by original designation.

brundini Kalugina, 1980b: 90. PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus EOCENITENDIPES Evenhuis, new name

MICROTENDIPES Hong, 1981: 47. Type species: Microtendipes melainus Hong, 1981, by original designation. [Preoccupied by Kieffer, 1915.] New synonymy.
EOCENITENDIPES Evenhuis, 1994: in this work (new replacement name for Microtendipes Hong). Type species: Microtendipes melainus Hong, 1981, automatic.

labrosus Hong, 1981: 48 (Microtendipes). PA: China (Eocene) [A].
longifemerales Hong, 1981: 50 (Microtendipes). PA: China (Eocene) [A].
melainus Hong, 1981: 47 (Microtendipes). PA: China (Eocene) [A].

 

Genus EURYCNEMUS Wulp

*EURYCNEMUS Wulp, 1874: 135. Type species: Chironomus elegans Meigen, 1818 [= Chironomus crassipes Panzer, 1813], by original designation.

appendiculatus Meunier, 1904c: 220 [1904d: 231]. PA: Baltic Region (Eocene/ Oligocene) [A].
hyalinus Meunier, 1904c: 220 [1904d: 231]. PA: Baltic Region (Eocene/Oligocene) [A].
pilosellus Meunier, 1904c: 220 [1904d: 231]. PA: Baltic Region (Eocene/Oligocene) [A].
stagnum Meunier, 1904c: 219 [1904d: 230]. PA: Baltic Region (Eocene/Oligocene) [A].
tenellus Meunier, 1904c: 219 [1904d: 230]. PA: Baltic Region (Eocene/Oligocene) [A].
vulgaris Meunier, 1904c: 218 [1904d: 229]. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus FUSHUNITENDIPES Hong

FUSHUNITENDIPES Hong, 1981: 31. Type species: Fushunitendipes eocenicus Hong, 1981, by original designation.
FUSHUNTENDIPES. Incorrect original spelling of Fushunitendipes (Hong, 1981: 6).
FUSHUNITENSIPES. Incorrect original spelling of Fushunitendipes (Hong, 1981: 44).

eocenicus Hong, 1981: 33. PA: China (Eocene) [A].
limnetes Hong, 1981: 44. PA: China (Eocene) [A].
limnetus. Incorrect original spelling of limnetes (Hong, 1981: 6, 36).
lobotes Hong, 1981: 37. PA: China (Eocene) [A].
labotes. Incorrect original spelling of lobotes (Hong, 1981: 6).
longipalpulatus Hong, 1981: 42. PA: China (Eocene) [A].
longipapulatus. Incorrect original spelling of longipalpulatus (Hong, 1981: 160).
platamodes Hong, 1981: 45. PA: China (Eocene) [A].
trichodes Hong, 1981: 41. PA: China (Eocene) [A].
uracanthodes Hong, 1981: 39. PA: China (Eocene) [A].
uracanthoides. Incorrect original spelling of uracanthodes (Hong, 1981: 6, 23, 40, 162).

 

Genus GURVANOMYIA Kalugina

GURVANOMYIA Kalugina, 1986: 121. Type species: Gurvanomyia magna Kalugina, 1986, by original designation.

magna Kalugina, 1986: 122. PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C].
moderata Kalugina, 1986: 122. PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C].
modesta. Incorrect original spelling of moderata (Sinitsa, 1986: 14).

 

Genus JENTZSCHIELLA Meunier

JENTZSCHIELLA Meunier, 1899b: 162. Type species: Jentzschiella jentzschi Evenhuis, 1993, by present designation.
[Jentzschiella originally proposed without included named species. The type designated is from the first inclusion of a named species within the genus in accordance with the Code.]
ZENTSCHIELLA. Incorrect subsequent spelling of Jentzschiella (Meunier, 1899e: pl. 1).

jentzschi Evenhuis, 1994: in this work (new species). PA: Baltic Region (Eocene/ Oligocene) [A].
[Species-group name valid by bibliographic reference to characters given in Meunier (1899b: 162) and Meunier (1899e: pl. 1) for the genus Jentzschiella.]

 

Genus JUROCHLUS Kalugina

JUROCHLUS Kalugina in Kalugina & Kovalev, 1985: 95. Type species: Jurochlus sibiricus Kalugina, 1985, by original designation.

rigor Kalugina in Kalugina & Kovalev, 1985: 96. PA: Russia (Siberia) (Middle Jurassic) [C].
sibiricus Kalugina in Kalugina & Kovalev, 1985: 95. PA: Russia (Siberia) (Middle Jurassic) [C].

 

 

Genus JUROPELOPIA Evenhuis

MESOPELOPIA Kalugina in Kalugina & Kovalev, 1985: 110. Type species: Mesopelopia fittkaui Kalugina, 1985, by original designation. [Preoccupied by Roback, 1971.].
JUROPELOPIA Evenhuis, 1994: in this work (new replacement name for Mesopelopia Kalugina). Type species: Mesopelopia fittkaui Kalugina, 1985, automatic.

fittkaui Kalugina in Kalugina & Kovalev, 1985: 110 (Mesopelopia). PA: Russia (Siberia) (Lower/Middle Jurassic) [C]. New combination.

 

Genus LIBANOCHLITES Brundin

LIBANOCHLITES Brundin, 1976: 149. Type species: Libanochlites neocomicus Brundin, 1976, by monotypy.

neocomicus Brundin, 1976: 149. PA: Lebanon (Lower Cretaceous) [A].

 

Genus MAILONIA Kalugina

MAILONIA Kalugina in Kalugina & Kovalev, 1985: 106. Type species: Mailonia pallida Kalugina, 1985, by original designation.

pallida Kalugina in Kalugina & Kovalev, 1985: 107. PA: Russia (Siberia) (Middle Jurassic) [C].

 

 

Genus MANLAYAMYIA Kalugina

MANLAYAMYIA Kalugina, 1980a: 63. Type species: Manlayamyia litorhina Kalugina, 1980, by original designation.

dabeigouensis Zhang, 1991: 557. PA: China (Upper Jurassic) [C].
litorhina Kalugina, 1980a: 64. PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C].

 

Genus METRIOCNEMUS Wulp

*METRIOCNEMUS Wulp, 1874: 136. Type species: Chironomus albolineatus Meigen, 1818, by subsequent designation of Coquillett (1910: 569).

cretatus Boesel, 1937: 52. NE: Canada (Upper Cretaceous) [A].

 

Genus MOGSONOMUS Kalugina

MOGSONOMUS Kalugina in Kalugina & Kovalev, 1985: 100. Type species: Mogsonomus tophus Kalugina, 1985, by original designation.

tophus Kalugina in Kalugina & Kovalev, 1985: 101. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus NOMOCHIRUS Gil Collado

NOMOCHIRUS Gil Collado, 1926: 91. Type species: Nomochirus sampelayoi Gil Collado, 1926, by monotypy.

sampelayoi Gil Collado, 1926: 94. PA: Spain (Paleocene) [C].

 

Genus ORTHOCLADIUS Wulp

*ORTHOCLADIUS Wulp, 1874: 132. Type species: Tipula stercoraria De Geer, 1776 [misidentification, = Chironomus oblidens Walker, 1856], by subsequent designation of Coquillett (1910: 581).

Unidentified spp.-PA: Germany (Oligocene) [C] (Statz, 1944c: 131­33).


Genus ORUSA Lin

ORUSA Lin, 1980: 226. Type species: Orusa barba Lin, 1980, by monotypy.

barba Lin, 1980: 226. PA: China (Upper Jurassic) [C].

 

Genus ORYCTOCHLUS Kalugina

ORYCTOCHLUS Kalugina in Kalugina & Kovalev, 1985: 83. Type species: Oryctochlus vulcanus Kalugina, 1985, by original designation.

affinis Kalugina in Kalugina & Kovalev, 1985: 91. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
contiguus Zhang, 1991: 561. PA: China (Upper Jurassic) [C].
longilobus Kalugina in Kalugina & Kovalev, 1985: 89. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
minor Kalugina in Kalugina & Kovalev, 1985: 91. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
minutus Kalugina in Kalugina & Kovalev, 1985: 92. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
vulcanus Kalugina in Kalugina & Kovalev, 1985: 85. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].

 

Genus PALAEOTANYPUS Meunier

PALAEOTANYPUS Meunier, 1904c: 178 [1904d: 189]. Type species: Tanypus filiformis Meunier, 1904, by monotypy.

filiformis Meunier, 1904c: 226 [1904d: 237] (Tanypus). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PODONOMIUS Kalugina

PODONOMIUS Kalugina in Kalugina & Kovalev, 1985: 101. Type species: Podonomius tugnuicus Kalugina, 1985, by original designation.

minimus Kalugina in Kalugina & Kovalev, 1985: 105. PA: Russia (Siberia) (Middle Jurassic) [C].
rotundatus Kalugina in Kalugina & Kovalev, 1985: 104. PA: Russia (Siberia) (Middle Jurassic) [C].
simplex Kalugina in Kalugina & Kovalev, 1985: 104. PA: Russia (Siberia) (Middle Jurassic) [C].
splendidus Kalugina in Kalugina & Kovalev, 1985: 104. PA: Russia (Siberia) (Lower/Middle Jurassic) [C].
tugnuicus Kalugina in Kalugina & Kovalev, 1985: 103. PA: Russia (Siberia) (Lower/Middle Jurassic) [C].
undulatus Kalugina in Kalugina & Kovalev, 1985: 105. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus PROCLADIUS Skuse

*PROCLADIUS Skuse, 1889: 283. Type species: Procladius paludicola Skuse, 1889, by subsequent designation of Coquillett (1910: 594).

perditus Heyden, 1870: 247 (Chironomus). PA: Germany (Oligocene) [C].

 

Genus SHINLUSTIA Kalugina

SHINLUSTIA Kalugina, 1986: 123. Type species: Shinlustia irae Kalugina, 1986, by original designation.
SCHINLUSTIA. Incorrect original spelling of Shinlustia (Sinitsa, 1986: 39, 42).

applanata Sinitsa, 1986: 39, 42 (Schinlustia). Nomen nudum.
irae Kalugina, 1986: 124. PA: Mongolia (Lower Cretaceous) [C].

 

Genus SINORYCTOCHLUS Zhang

SINORYCTOCHLUS Zhang, 1991: 563. Type species: Sinoryctochlus insolitus Zhang, 1991, by original designation.

insolitus Zhang, 1991: 564. PA: China (Upper Jurassic) [C].


Genus SMITTIA Holmgren

*SMITTIA Holmgren, 1869: 47. Type species: Chironomus brevipennis Boheman, 1865, by monotypy.

veta Boesel, 1937: 53 (Spaniotoma). NE: Canada (Upper Cretaceous) [A].


Genus SPANIOTOMA Philippi

*SPANIOTOMA Philippi, 1865: 629. Type species: Spaniotoma bivittata Philippi, 1865, by subsequent designation of Johannsen (1905: 162).

conservata Boesel, 1937: 52. NE: Canada (Upper Cretaceous) [A].

 

Genus TANYPUS Meigen

PELOPIA Meigen, 1800: 18. Suppressed by I.C.Z.N. (1963: 339).
*TANYPUS Meigen, 1803: 261. Type species: Tipula cincta Fabricius, 1794 [misidentification, = Tipula punctipennis Meigen, 1818], by subsequent designation of Latreille (1810: 442) (as "Chironomus cinctus, Fab.").

compactus Meunier, 1904c: 223 [1904d: 234]. PA: Baltic Region (Eocene/Oligocene) [A].
dorminans Heyden, 1870: 248 (Chironomus). PA: Germany (Oligocene) [C].
eridanus Meunier, 1904c: 224 [1904d: 235]. PA: Baltic Region (Eocene/Oligocene) [A].
fusiformis Meunier, 1904c: 222 [1904d: 233]. PA: Baltic Region (Eocene/Oligocene) [A].
longicornis Meunier, 1904c: 225 [1904d: 236]. PA: Baltic Region (Eocene/Oligocene) [A].
obscurus Statz, 1944c: 129 (Pelopia). PA: Germany (Oligocene) [C]. [Preoccupied by Macquart, 1826; no new replacement name proposed here.]
pagasti Statz, 1944c: 130 (Pelopia). PA: Germany (Oligocene) [C].
palaemon Heyden, 1870: 249 (Chironomus). PA: Germany (Oligocene) [C].
parvus Meunier, 1904c: 225 [1904d: 236]. PA: Baltic Region (Eocene/Oligocene) [A].
porrectus Meunier, 1904c: 223 [1904d: 234]. PA: Baltic Region (Eocene/Oligocene) [A].
subrotundatus Meunier, 1904c: 224 [1904d: 235]. PA: Baltic Region (Eocene/ Oligocene) [A].
thienemanni Statz, 1944c: 128 (Pelopia). PA: Germany (Oligocene) [C].

 

Genus TANYTARSUS Wulp

*TANYTARSUS Wulp, 1874: 134. Type species: Chironomus signatus Wulp, 1858, by subsequent designation of I.C.Z.N. (1961: 361).
*CALOPSECTRA Kieffer, 1909: 50. Type species: Tanytarsus gregarius Kieffer, 1909, by subsequent designation of Kieffer (1921: 26).

insularis Meunier, 1904c: 217 [1904d: 228]. PA: Baltic Region (Eocene/Oligocene) [A].
maritimus Meunier, 1904c: 218 [1904d: 229]. PA: Baltic Region (Eocene/Oligocene) [A].
wulpii Meunier, 1904c: 218 [1904d: 229]. PA: Baltic Region (Eocene/Oligocene) [A].
Unidentified sp.-NE: USA (Miocene) [P] (Palmer, 1957: 274).

 

Genus TENDIPOPSIS Hong & Wang

TENDIPOPSIS Hong & Wang, 1990: 131. Type species: Tendipopsis colorata Hong & Wang, 1990, by original designation.
TENDIPOSIS. Incorrect original spelling of Tendipopsis (Regional Geological Surveying Team, 1990: table 3).

colorata Hong & Wang, 1990: 131. PA: China (Upper Jurassic) [C].

 

Genus TINACTUM Lin

TINACTUM Lin, 1980: 226. Type species: Tinactum solusum Lin, 1980, by original designation.

solusum Lin, 1980: 226. PA: China (Upper Jurassic) [C].

 

Genus TOPHOCLADIUS Kalugina

TOPHOCLADIUS Kalugina in Kalugina & Kovalev, 1985: 107. Type species: Tophocladius stygialis Kalugina, 1985, by original designation.

stygialis Kalugina in Kalugina & Kovalev, 1985: 109. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus ULAIA Kalugina

ULAIA Kalugina in Kalugina & Kovalev, 1985: 93. Type species: Ulaia montana Kalugina, 1985, by original designation.

magna Kalugina in Kalugina & Kovalev, 1985: 94. PA: Russia (Siberia) (Middle Jurassic) [C].
montana Kalugina in Kalugina & Kovalev, 1985: 93. PA: Russia (Siberia) (Middle Jurassic) [C].
reducta Kalugina in Kalugina & Kovalev, 1985: 94. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus ULAIMAILIA Kalugina

ULAIMAILIA Kalugina in Kalugina & Kovalev, 1985: 100. Type species: Ulaimailia vetula Kalugina, 1985, by original designation.

vetula Kalugina in Kalugina & Kovalev, 1985: 100. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus ULAIMAILONIA Kalugina

ULAIMAILONIA Kalugina in Kalugina & Kovalev, 1985: 96. Type species: Ulaimailonia mixta Kalugina, 1985, by original designation.

mixta Kalugina in Kalugina & Kovalev, 1985: 97. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus ULAIMAILONIELLA Kalugina

ULAIMAILONIELLA Kalugina in Kalugina & Kovalev, 1985: 99. Type species: Ulaimailoniella fusiformis Kalugina, 1985, by original designation.

fusiformis Kalugina in Kalugina & Kovalev, 1985: 99. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus VIDUATA Lin

VIDUATA Lin, 1980: 226. Type species: Viduata otiosa Lin, 1980, by original designation.

otiosa Lin, 1980: 26. PA: China (Upper Jurassic) [C].

 

Unplaced Species in CHIRONOMIDAE

antiquus Heyden, 1859: 13 (Chironomus). PA: Germany (Oligocene) [C].
decrepitus Heyden, 1870: 248 (Chironomus). PA: Germany (Oligocene) [C].
gorchonensis Kalugina in Kalugina & Kovalev, 1985: 112 ("Tanypodites"). PA: Russia (Siberia) (Middle Jurassic) [C].

 

Questionably Placed in CHIRONOMIDAE

Genus ASUBA Giebel

ASUBA Giebel, 1856: 253. Type species: Asuba brodiei Giebel, 1856, by monotypy.

brodiei Giebel, 1856: 226 (new replacement name for dubius Westwood). PA: UK (England) (Lower Cretaceous) [C].
dubius Westwood in Brodie, 1845: 121, pl. 3, fig. 10 (Tanypus). PA: UK (England) (Lower Cretaceous) [C]. [Preoccupied by Meigen, 1804.]

 

Genus BIBIONITES Handlirsch

BIBIONITES Handlirsch, 1906a: 631. Type species: Macropeza prisca Giebel, 1856, by monotypy.
BIBIONITIS. Incorrect original spelling of Bibionites (Handlirsch, 1906a: 631).

prisca Giebel, 1856: 252 (Macropeza) PA: UK (England) (Lower Cretaceous) [C].

 

Genus CHIRONOMITES Fric

CHIRONOMITES Fric, 1901: 170. Type species: Chironomites adhaerens Fric, by present designation.

adhaerens Fric, 1901: 170. PA: Czech Republic (Upper Cretaceous) [C].
unionis Fric, 1901: 170. PA: Czech Republic (Upper Cretaceous) [C].

 

Genus DARA Giebel

DARA Giebel, 1856: 254. Type species: Culex fossilis Westwood, 1845, by monotypy.

fossilis Westwood in Brodie, 1845: 121, pl. 3, fig. 15 (Culex). PA: UK (England) (Lower Cretaceous) [C].

 

Genus EOPODONOMUS Rohdendorf

EOPODONOMUS Rohdendorf, 1964: 212 (1974: 236). Type species: Eopodonomus nymphalis Rohdendorf, 1964, by original designation.

nymphalis Rohdendorf, 1964: 212 (1974: 237). PA: Kazakhstan (Upper Jurassic) [C].

 

Genus PACHYURONYMPHA Rohdendorf

PACHYURONYMPHA Rohdendorf, 1964: 214 (1974: 238). Type species: Pachyuronympha karatauensis Rohdendorf, 1964, by original designation.

karatauensis Rohdendorf, 1964: 214 (1974: 238). PA: Kazakhstan (Upper Jurassic) [C].

 

Genus SENDELIA Meunier

SENDELIA Duisberg, 1868: 23. Nomen nudum.
SENDELIA Meunier, 1899b: 162. Type species: Sendelia mirabilis Evenhuis, 1993, by present designation.
[Sendelia Meunier originally proposed without included species. The present type species designation is from the first included species within the genus in accordance with the Code.]

mirabilis Evenhuis, 1994: 277. PA: Baltic Region (Eocene/Oligocene) [A].
[Species-group name valid by bibliographic reference to characters in Meunier (1899b: 162) for the genus Sendelia.]
mirabilis Duisberg, 1868: 23. Nomen nudum.

 

This page last revised 21 June 2002