Family EMPIDIDAE

 

Empididae, commonly called balloon flies or dance flies, are found worldwide, but are found most frequently in the northern temperate and montane regions. The family consists of over 3,000 species exhibiting a wide array of size and form. Both adults and larvae are predaceous on other insects. Adults of extant forms can be found on trees, in vegetation, or on the surface of water. Larval habitats range from aquatic to terrestrial and include such substrates as soil, leaf litter, dung, and rotten wood.


Members of this family are frequently found in amber inclusions and are one of the more generically diverse families represented in the fossil Diptera record. However, much undetermined material exists in collections awaiting description. In addition to the described species listed below, Langenheim et al. (1960) recorded a specimen from the Cretaceous amber deposits of coastal Alaska, and Kovalev (1986: 152­54) described and illustrated two undetermined forms from the Cretaceous deposits of western Mongolia. Jarzembowski (1979: 251) recorded the family from the Eocene/Oligocene Bembridge Marls Formations of the Isle of Wight, England, Fujiyama (1985: 43) described and illustrated a specimen from the Miocene deposits of Sado Island, Japan, Schumann & Wendt (1989: 42) recorded material from the Miocene Bitterfeld amber of Germany, and Hennig in Bachmayer & Schulz (1978: 114­19) described an unnamed hemerodromiine from the Miocene amber of Austria. An undetermined hybotine was recorded from the Eocene/Oligocene Hampstead Beds on the Isle of Wight by Jarzembowski (in litt.).


Legg (1942) mentioned undetermined Empididae from the Upper Cretaceous amber of Canada. Later, Hennig (1970: 10) determined some of this material as belonging to the Microphorinae and Zherichin (1978: 98) determined other of this material as belonging to the Ocydromiinae.


Carpenter (1992) and Grimaldi (1990) most recently listed empidid fossils. However, Grimaldi's list should be taken with caution as it lists the rhagionid genus Zarzia and the mythicomyiid genera Procyrtosia and Proplatypygus as empidoid fossils.

Ref.: Melander (1928, review of world genera).

 

Genus APALOCNEMIS Philippi

*APALOCNEMIS Philippi, 1865: 752. Type species: Apalocnemis obscura Philippi, 1865, by monotypy.
*TIMALPHES Melander, 1928: 97. Type species: Clinocera fumosa Hutton, 1901, by original designation.

acuticornis Loew, 1850b: 41 (Gloma). Nomen nudum.
gracilis Meunier, 1908i: 113 (Brachystoma). PA: Baltic Region (Eocene/Oligocene) [A].
palpata Loew, 1850b: 41 (Gloma). Nomen nudum.

 

Genus ARCHICHRYSOTUS Negrobov

ARCHICHRYSOTUS Negrobov, 1978: 84 (1979: 223). Type species: Archichrysotus hennigi Negrobov, 1978, by original designation.

hennigi Negrobov, 1978: 85 (1979: 224). PA: Russia (Siberia) (Upper Cretaceous) [A].
minor Negrobov, 1978: 86 (1979: 225). PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus ARCHIPLATYPALPUS Kovalev

ARCHIPLATYPALPUS Kovalev, 1974: 84 (195). Type species: Archiplatypalpus cretaceus Kovalev, 1974, by original designation.

cretaceus Kovalev, 1974: 86 (197). PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus BRACHYSTOMA Meigen

*BRACHYSTOMA Meigen, 1822: 12. Type species: Syrphus vesiculosus Fabricius, 1794, by subsequent designation of Blanchard (1840: 582).
[In the interests of stability of nomenclature and common usage, an earlier type designation by Westwood (1840: 132) of Brachystoma longicornis Meigen, which would make Brachystoma a senior synonym of Trichopeza Rondani, is here set aside. Application to the I.C.Z.N. to suppress Westwood's designation in favor of Blanchard's is necessary.]

spinulosa Loew, 1850b: 41. Nomen nudum.
spirulosa Meunier, 1899e: 178. Nomen nudum.
vesiculosa Fabricius, 1794: 299 (Syrphus). PA: Italy; Europe; PA: Baltic Region (Eocene/ Oligocene) [A].

 

Genus CHELIFERA Macquart

*CHELIFERA Macquart, 1823: 150. Type species: Chelifera raptor Macquart, 1823 [= Hemerodromia monostigma Meigen, 1822], by monotypy.

detestata Meunier, 1908i: 103 (Hemerodromia). PA: Baltic Region (Eocene/ Oligocene) [A].


Genus CHELIPODA Macquart

*CHELIPODA Macquart, 1823: 148. Type species: Tachydromia mantispa Panzer, 1806 [misidentification, = Tachydromia vocatoria Fallén, 1816], by original designation.
*PHYLLODROMIA Zetterstedt, 1837: 31 (as Hemerodromia subgenus). Type species: Empis melanocephala Fabricius, 1794, by subsequent designation of Rondani (1856: 148).
*LEPIDOMYA Bigot, 1857: 563. Type species: Tachydromia mantispa Panzer, 1806 [= Empis melanocephala Fabricius, 1794], by monotypy.
*LEPIDOMYIA Bezzi, 1903: 271 (unjustified emendation of Lepidomya Bigot). Type species: Tachydromia mantispa Panzer, 1806 [= Empis melanocephala Fabricius, 1794], automatic. [Preoccupied by Loew, 1864.]

delicata Meunier, 1908i: 105 (Phyllodromia). PA: Baltic Region (Eocene/Oligocene) [A].
dolosa Meunier, 1908i: 104 (Phyllodromia). PA: Baltic Region (Eocene/Oligocene) [A].
vaga Meunier, 1908i: 106 (Lepidomyia). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus CRETOMICROPHORUS Negrobov

CRETOMICROPHORUS Negrobov, 1978: 82 (1979: 222). Type species: Cretomicrophorus rohdendorfi Negrobov, 1978, by original designation.
JANTARDACHIA Zherichin, 1978: 86. Nomen nudum.

rohdendorfi Negrobov, 1978: 83 (1979: 222). PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus CRETOPLATYPALPUS Kovalev

CRETOPLATYPALPUS Kovalev, 1978: 72 (1979: 353). Type species: Cretoplatypalpus archeus Kovalev, 1978, by original designation.

archeus Kovalev, 1978: 72 (1979: 353). PA: Russia (Siberia) (Upper Cretaceous) [A].

 

Genus DRAPETIELLA Meunier

DRAPETIELLA Meunier, 1908i: 97. Type species: Drapetiella definitum Meunier, 1908, by monotypy.

definitum Meunier, 1908i: 97. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus DRAPETIS Meigen

*DRAPETIS Meigen, 1822: 91. Type species: Drapetis exilis Meigen, 1822, by monotypy.

brevis Meunier, 1908i: 95. PA: Baltic Region (Eocene/Oligocene) [A].
decolorata Meunier, 1908i: 95. PA: Baltic Region (Eocene/Oligocene) [A].
decoratum Meunier, 1908i: 96. PA: Baltic Region (Eocene/Oligocene) [A].
mortuum Meunier, 1908i: 97. PA: Baltic Region (Eocene/Oligocene) [A].
vitiosum Meunier, 1908i: 96. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus DYSALETRIA Loew

*DYSALETRIA Loew, 1864b: 30. Type species: Dysaletria melanocephala Loew, 1864 [= Tachypeza atriceps Boheman, 1852], by monotypy.

diabolica Meunier, 1908i: 100 (Elaphropeza). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus ECOMMOCYDROMIA Schlüter

ECOMMOCYDROMIA Schlüter, 1978b: 103. Type species: Ecommocydromia difficilis Schlüter, 1978, by monotypy.

difficilis Schlüter, 1978b: 103. PA: France (Upper Cretaceous) [A].

 

Genus ELECTROCYRTOMA Cockerell

ELECTROCYRTOMA Cockerell, 1917c: 22. Type species: Electrocyrtoma burmanica Cockerell, 1917, by monotypy.

burmanicum Cockerell, 1917c: 22. OR: Myanmar (?Cretaceous) [A].
electrica, error for burmanicum (Chvála, 1981: 233).

 

Genus EMPIS Linnaeus

*EMPIS Linnaeus, 1758: 603. Type species: Empis pennipes Linnaeus, 1758, by subsequent designation of Latreille (1810: 443).

Subgenus ACALLOMYIA Melander

*ACALLOMYIA Melander, 1928: 140 (as genus). Type species: Iteaphila peregrina Melander, 1902, by original designation.

probolaea Melander, 1949: 38. NE: USA (Oligocene) [C].

Unplaced to Subgenus of EMPIS Linnaeus

arcuata Keilbach, 1982: 368. Nomen nudum.
bulbirostris Meunier, 1899e: 178. Nomen nudum.
carbonum Germar, 1837: pl. 21. PA: Germany (Miocene) [C].
externa Keilbach, 1982: 366. Nomen nudum.
florissantana Cockerell, 1915a: 645. NE: USA (Oligocene) [C].
infossa Melander, 1949: 39. NE: USA (Oligocene) [C].
inscita Meunier, 1908i: 121. PA: Baltic Region (Eocene/Oligocene) [A].
longipes Keilbach, 1982: 366. Nomen nudum. [Preoccupied by Meigen, 1804.]
macilenta Meunier, 1908i: 121 (Hilara). PA: Baltic Region (Eocene/Oligocene) [A].
macrophthalma Förster, 1891: 482. PA: Germany (Oligocene) [C].
mala Meunier, 1908i: 123. PA: Baltic Region (Eocene/Oligocene) [A].
malefica Meunier, 1908i: 122. PA: Baltic Region (Eocene/Oligocene) [A].
melia Heyden, 1870: 259. PA: Germany (Oligocene) [C].
miocenica Cockerell, 1915a: 645. NE: USA (Oligocene) [C].
mordax Meunier, 1908i: 123. PA: Baltic Region (Eocene/Oligocene) [A].
morosella Melander, 1928: 164 (new replacement name for morosa Meunier). PA: Baltic Region (Eocene/Oligocene) [A].
morosa Meunier, 1908i: 125. PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Meigen, 1822.]
orapaensis Waters, 1989a: 235. AF: Botswana (Upper Cretaceous) [C].
perdita Cockerell, 1916c: 92. NE: USA (Oligocene) [C].
personata Meunier, 1908i: 122. PA: Baltic Region (Eocene/Oligocene) [A].
poeppigi Giebel, 1856: 207. PA: Baltic Region (Eocene/Oligocene) [A].
pulvillata Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].
simplex Keilbach, 1982: 366. Nomen nudum.
spinifera Statz, 1940: 132. PA: Germany (Oligocene) [A]. [Preoccupied by Bezzi, 1909; no new replacement name proposed here.]
spinitarsis Keilbach, 1982: 366. Nomen nudum.
stilicornis Meunier, 1899e: 178. Nomen nudum.
tibialis Meunier, 1899e: 178. Nomen nudum.
tritava Melander, 1928: 176 (new replacement name for tristis Meunier). PA: Baltic Region (Eocene/Oligocene) [A].
tristis Meunier, 1908i: 124. PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Loew, 1867.]
tumidirostris Keilbach, 1982: 366. Nomen nudum.
umbonata Statz, 1940: 133. PA: Germany (Oligocene) [C].

 

Genus EUHYBUS Coquillett

*EUHYBUS Coquillett, 1895: 437. Type species: Hybos purpureus Walker, 1849, by subsequent designation of Coquillett (1903: 250).

Unidentified sp.-PA: Baltic Region (Eocene/Oligocene) [A] (Hennig, 1973: 46).

 

Genus EUTHYNEURIELLA Meunier

EUTHYNEURIELLA Meunier, 1908i: 112. Type species: Euthyneuriella longirostris Meunier, 1908, by monotypy.

longirostris Meunier, 1908i: 113. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus GLOMA Meigen

*GLOMA Meigen, 1822: 14. Type species: Gloma fuscipennis Meigen, 1822, by monotypy.

fuscipennis Loew, 1850b: 41. Nomen nudum.
hirta Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus HEMERODROMIA Meigen

*HEMERODROMIA Meigen, 1822: 61. Type species: Tachydromia oratoria Fallén, 1815, by subsequent designation of Rondani (1856: 148).
[Suspension of I.C.Z.N. rules required to validate Rondani's type designation, and to set aside earlier type designations by Westwood (1840: 132) and Duponchel in d'Orbigny (1845: 528) in the interests of stability and common usage. See Melander (1928: 252) and Collin (1961: 715).]

praedatrix Keilbach, 1982: 372. Nomen nudum.
raptatrix Keilbach, 1982: 372. Nomen nudum.
Unidentified sp.-PA: Baltic Region (Eocene/Oligocene) [A] (Loew, 1850b: 41).

 

Genus HILARA Meigen

*HILARA Meigen, 1822: 1. Type species: Empis maura Fabricius, 1777, by subsequent designation of Curtis (1826: pl. 130).

exilis Meunier, 1908i: 121 (Empis). PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Coquillett, 1903; no new replacement name proposed here.]
heeri Meunier, 1915b: 12. PA: France (Oligocene) [C].
litigiosa Meunier, 1908i: 120. PA: Baltic Region (Eocene/Oligocene) [A].
royoi Gil Collado, 1926: 98. PA: Spain (Paleocene/Pliocene) [C].
tarda Meunier, 1908i: 120. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus HILARITES Heer

HILARITES Heer, 1856: 38. Type species: Hilarites bellus Heer, 1856, by monotypy.

bellus Heer, 1856: 38. PA: France (Oligocene) [C].

 

Genus HYBOS Meigen

*NEOZA Meigen, 1800: 27. Suppressed by I.C.Z.N. (1963: 339).
*HYBOS Meigen, 1803: 269. Type species: Hybos funebris Meigen, 1804 [= Musca grossipes Linnaeus, 1767], by subsequent designation of Curtis (1837: pl. 661).

exilis Meunier, 1908i: 109. PA: Baltic Region (Eocene/Oligocene) [A].
minutus Keilbach, 1982: 369. Nomen nudum.
tenuis Meunier, 1908i: 109. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus LEPTOPEZA Macquart

*LEMTOPEZA Macquart, 1827: 143. Type species: Ocydromia flavipes Meigen, 1820 [= Ocydromia ruficollis Meigen, 1820], by monotypy.
*LEPTOPEZA. Incorrect subsequent spelling of Lemtopeza (Macquart, 1834: 320).
*LEPTOPEZA Scudder, 1882: 182 (unjustified emendation of Lemtopeza). Type species: Type species: Ocydromia flavipes Meigen, 1820 [= Ocydromia ruficollis Meigen, 1820], automatic.
[Though traditionally recorded as such, Macquart did not provide a true emendation from Lemtopeza to Leptopeza in his 1834 work according to Article 33(b) of the Code. Lemtopeza is not an incorrect original spelling according to Article 32(c) of the Code and should be treated as the correct orthography for this genus. However, following established usage and in order to maintain stability, Leptopeza is used here as the accepted orthography for this genus.]

clavipes Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].
concinna Meunier, 1908i: 110. PA: Baltic Region (Eocene/Oligocene) [A].
spinigera Meunier, 1899e: 178. Nomen nudum.

 

Genus LOCHMOCOLA Hong

LOCHMOCOLA Hong, 1981: 97. Type species: Lochmocola osterosa Hong, 1981, by original designation.
LOMOCOLA. Incorrect original spelling of Lochmocola (Hong, 1981: 20).

osterosa Hong, 1981: 97. PA: China (Eocene) [A].

 

Genus MEGHYPERIELLA Meunier

MEGHYPERIELLA Meunier, 1908i: 112. Type species: Meghyperiella porphyropsoides Meunier, 1908, by monotypy.

porphyropsoides Meunier, 1908i: 112. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus MICREMPIS Melander

*MICREMPIS Melander, 1928: 298. Type species: Micrempis nana Melander, 1928, by original designation.

eocenica Meunier, 1908i: 97 (Phoneutisca). PA: Baltic Region (Eocene/Oligocene) [A].
suspiciosa Meunier, 1908i: 98 (Phoneutisca). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus MICROPHOR Macquart

*MICROPHOR Macquart, 1827: 139. Type species: Microphor velutinus Macquart, 1827, by subsequent designation of Rondani (1856: 151).
*MICROPHORUS. Incorrect subsequent spelling of Microphor (Macquart, 1834: 345).
*MICROPHORA Zetterstedt, 1842: 24 (unjustified emendation of Microphorus Macquart). Type species: Microphor velutinus Macquart, 1827, automatic.

defunctus Handlirsch, 1910: 124. NE: Canada (Oligocene) [C].
eocenica Meunier, 1902d: 402 (Rhamphomyia). PA: Baltic Region (Eocene/Oligocene) [A].
rusticus Meunier, 1908i: 105 (Phyllodromia). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus MICROPHORITES Hennig

MICROPHORITES Hennig, 1971b: 16. Type species: Microphorites extinctus Hennig, 1971, by monotypy.

extinctus Hennig, 1971b: 16. PA: Lebanon (Lower Cretaceous) [A].

 

Genus OEDALEA Meigen

*OEDALEA Meigen, 1820: 355. Type species: Empis hybotina Fallén, 1816, by subsequent designation of Thon in Ersch & Gruber (1832: 38).

putida Meunier, 1908i: 126 (Microphorus). PA: Baltic Region (Eocene/Oligocene) [A].
robusta Meunier, 1908i: 110. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus OUSTALETIMYIA Meunier

OUSTALETIMYIA Meunier, 1893c: cccxxxiii (1894b: cccxxxiii). Type species: Oustaletimyia succinorum Meunier, 1893, by monotypy.

succinorum Meunier, 1893c: cccxxxiii (1894b: cccxxxiii). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PALAEOEDALEA Meunier

PALAEOEDALEA Meunier, 1902a: 101. Type species: Palaeoedalea samlandica Meunier, 1902, by monotypy.

samlandica Meunier, 1902a: 101. PA: Baltic Region (Eocene/Oligocene) [A].
elegans Meunier, 1902a: fig. 5. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PALAEOLEPTOPEZA Meunier

PALAEOLEPTOPEZA Meunier, 1908i: 111. Type species: Palaeoleptopeza gracilis Meunier, 1908, by monotypy.

gracilis Meunier, 1908i: 111. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PALAEOPARAMESIA Meunier

PALAEOPARAMESIA Meunier, 1902a: 98. Type species: Palaeoparamesia proosti Meunier, 1902, by monotypy.

proosti Meunier, 1902a: 99. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PARATHALASSIELLA Meunier

PARATHALASSIELLA Meunier, 1908i: 106. Type species: Parathalassiella problematica Meunier, 1908, by monotypy.

problematica Meunier, 1908i: 106. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PLATYPALPUS Macquart

*PLATYPALPUS Macquart, 1827: 92. Type species: Musca cursitans Fabricius, 1775, by subsequent designation of Westwood (1840: 132).

concitatus Meunier, 1908i: 100. PA: Baltic Region (Eocene/Oligocene) [A].
eversoris Meunier, 1908i: 102. PA: Baltic Region (Eocene/Oligocene) [A].
interfectoris Meunier, 1908i: 102. PA: Baltic Region (Eocene/Oligocene) [A].
predatoris Meunier, 1908i: 101. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PROGLOMA James

PROGLOMA James, 1937: 247. Type species: Progloma rohweri James, 1937, by original designation.

rohweri James, 1937: 247. NE: USA (Oligocene) [C].

 

Genus PROTOEDALEA Cockerell

PROTOEDALEA Cockerell, 1920a: 252. Type species: Protoedalea brachystoma Cockerell, 1920, by original designation.

brachystoma Cockerell, 1920a: 252. NE: USA (Eocene) [C].

 

Genus PSEUDOACARTERUS Waters

PSEUDOACARTERUS Waters, 1989b: 658. Type species: Pseudoacarterus orapaensis Waters, 1989, by original designation.

orapaensis Waters, 1989b: 658. AF: Botswana (Upper Cretaceous) [C].

 

Genus RAGAS Walker

*RAGAS Walker, 1837: 229. Type species: Ragas unica Walker, 1837, by monotypy.
*RHAGAS Agassiz, 1846: 320 (unjustified emendation of Ragas Walker). Type species: Ragas unica Walker, 1837, automatic.

generosa Meunier, 1908i: 125. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus RHAMPHOMYIA Meigen

*RHAMPHOMYIA Meigen, 1822: 42. Type species: Empis marginata Fabricius, 1787, by subsequent designation of Guérin-Méneville in Bory de Saint-Vincent (1828: 547).

ablata Meunier, 1908i: 114. PA: Baltic Region (Eocene/Oligocene) [A].
aeterna Melander, 1949: 40. NE: USA (Oligocene) [C].
angusta Meunier, 1908i: 119. PA: Baltic Region (Eocene/Oligocene) [A].
antipedalis Loew, 1850b: 41. Nomen nudum.
brevipila Keilbach, 1982: 367. Nomen nudum.
calvimontis Cockerell, 1916a: 123. NE: USA (Oligocene) [C].
corrupta Meunier, 1908i: 114. PA: Baltic Region (Eocene/Oligocene) [A].
craterae Melander, 1949: 40. NE: USA (Oligocene) [C].
crinitarsis Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].
distans Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].
enena Cockerell, 1921b: 30. NE: USA (Eocene) [C].
errabunda Meunier, 1908i: 117. PA: Baltic Region (Eocene/Oligocene) [A].
formosa Loew, 1850b: 41. Nomen nudum.
fossa Melander, 1949: 41. NE: USA (Oligocene) [C].
fulvescens Statz, 1940: 133. PA: Germany (Oligocene) [C].
hypolitha Cockerell, 1917b: 378. NE: USA (Oligocene) [C].
inanimata Melander, 1949: 41. NE: USA (Oligocene) [C].
infernalis Melander, 1949: 42. NE: USA (Oligocene) [C].
insolita Meunier, 1908i: 118. PA: Baltic Region (Eocene/Oligocene) [A].
interita Melander, 1949: 43. NE: USA (Oligocene) [C].
involuta Meunier, 1908i: 115. PA: Baltic Region (Eocene/Oligocene) [A].
latipennata Statz, 1940: 134. PA: Germany (Oligocene) [C].
media Meunier, 1908i: 119. PA: Baltic Region (Eocene/Oligocene) [A].
morticina Melander, 1949: 43. NE: USA (Oligocene) [C].
obtusa Meunier, 1908i: 118. PA: Baltic Region (Eocene/Oligocene) [A].
oedaloides Meunier, 1908i: 117. PA: Baltic Region (Eocene/Oligocene) [A].
polymorpha Meunier, 1899e: 178. PA: Baltic Region (Eocene/Oligocene) [A].
porrecta Meunier, 1908i: 117. PA: Baltic Region (Eocene/Oligocene) [A].
pteropa Loew, 1850b: 41. Nomen nudum.
ptilopa Loew, 1850b: 41. Nomen nudum.
remitarsis Loew, 1850b: 41. Nomen nudum.
rottensis Statz, 1940: 135. PA: Germany (Oligocene) [C].
senecta Melander, 1949: 44. NE: USA (Oligocene) [C].
sepulta Cockerell, 1916a: 123. NE: USA (Oligocene) [C].
setosa Keilbach, 1982: 366. Nomen nudum. [Preoccupied by Coquillett, 1895.]
spodites Melander, 1949: 44. NE: USA (Oligocene) [C].
tenuicornis Keilbach, 1982: 366. Nomen nudum.
tumulata Melander, 1949: 45. NE: USA (Oligocene) [C].
ungulina Loew, 1850b: 41. PA: Baltic Region (Eocene/Oligocene) [A].
unipile Keilbach, 1982: 367. Nomen nudum.

 

Genus STILPON Loew

*AGATACHYS Meigen, 1830: 343. Type species: Tachydromia celeripes Meigen, 1830, by monotypy.
[Proposed in specific synonymy. Made available by use as a senior synonym of Stilpon by Coquillett (1910: 504). In accordance with stability of nomenclature and common usage, the name Stilpon is used here, though Agatachys has priority. Action by the I.C.Z.N. to suppress Agatachys in favor of Stilpon is desirable.]
*STILPON Loew, 1859: 34 (as Drapetis subgenus). Type species: Tachydromia graminum Fallén, 1815, by subsequent designation of Loew (1864b: 5).

Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Cumming & Cooper, 1992: 956).

 

Genus SYMBALLOPHTHALMUS Becker

*MACROPTERA Becker, 1889a: 80. Type species: Macroptera pictipes Becker, 1889, by monotypy. [Preoccupied by Lioy, 1863.]
*SYMBALLOPHTHALMUS Becker, 1889b: 285 (new replacement name for Macroptera Becker). Type species: Macroptera pictipes Becker, 1889, automatic.

clavilabrosus Hong, 1981: 48. PA: China (Eocene) [A].

 

Genus TACHYDROMIA Meigen

*TACHYDROMIA Meigen, 1803: 269. Type species: Musca cimecoides Fabricius, 1779 [misidentification, = Tachydromia connexa Meigen, 1822], by subsequent designation of Curtis (1833: pl. 477).

Subgenus TACHYPONA Keilbach

TACHYPONA Keilbach, 1982: 371. Type species: Tachydromia postposita Keilbach, 1982, by monotypy.
[Validated by bibliographic reference to characters in Loew (1850b: 42).]

postposita Keilbach, 1982: 371. PA: Baltic Region (Eocene/Oligocene) [A].
[Validated by bibliographic reference to characters in Loew (1850b: 42).]

Unplaced to Subgenus of TACHYDROMIA Meigen

egelata Meunier, 1908i: 98. PA: Baltic Region (Eocene/Oligocene) [A].
theobaldi Timon-David, 1944: 128. PA: France (Oligocene) [C].
voracis Meunier, 1908i: 99. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus TACHPYEZA Meigen

*TACHYPEZA Meigen, 1830: 341. Type species: Tachydromia nervosa Meigen, 1822 [= Tachydromia nubila Meigen, 1804], by subsequent designation of Rondani (1856: 147).

primitiva Melander, 1949: 45. NE: USA (Oligocene) [C].

 

Genus THIRZA Giebel

THIRZA Giebel, 1856: 209. Type species: Thirza naumanni Giebel, 1856, by monotypy.

naumanni Giebel, 1856: 210. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus TRICHINITES Hennig

TRICHINITES Hennig, 1970: 7. Type species: Trichinites cretaceus Hennig, 1970, by monotypy.

cretaceus Hennig, 1970: 7. PA: Lebanon (Lower Cretaceous) [A].

 

Genus TRICHOPEZA Rondani

*TRICHOPEZA Rondani, 1856: 150. Type species: Brachystoma longicornis Meigen, 1822, by original designation.
TRICOPEZA. Incorrect original spelling of Trichopeza (Rondani, 1856: 225).

sucina Melander, 1928: 220 (new replacement name for longicornis Meunier). PA: Baltic Region (Eocene/Oligocene) [A].
longicornis Meunier, 1908i: 108. PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Meigen, 1822.]

 

Unplaced Species in Empididae

stilpon Meunier, 1899e: 178­79 (Tachydromia). PA: Baltic Region (Eocene/ Oligocene) [A].

 

Questionably Placed in EMPIDIDAE

Genus BURMITEMPIS Cockerell

BURMITEMPIS Cockerell, 1917f: 367. Type species: Burmitempis halteralis Cockerell, 1917, by monotypy.

halteralis Cockerell, 1917f: 367. OR: Myanmar (?Cretaceous) [A].

 

Genus EMPIDIA Weyenbergh

EMPIDIA Weyenbergh, 1869: 258. Type species: Empidia wulpi Weyenbergh, 1869, by monotypy.

wulpi Weyenbergh, 1869: 258. PA: Germany (Upper Jurassic) [C].

 

Genus HASMONA Giebel

HASMONA Giebel, 1856: 208. Type species: Hasmona leo Giebel, 1856, by monotypy.

leo Giebel, 1856: 209. PA: UK (England) (Upper Jurassic) [C].

 

Genus RETINITUS Negrobov

RETINITUS Negrobov, 1978: 86 (1979: 225). Type species: Retinitus nervosus Negrobov, 1978, by original designation.

nervosus Negrobov, 1978: 87 (1979: 226). PA: Russia (Siberia) (Upper Cretaceous) [A].