Platystomatidae are worldwide in distribution and one of the larger families
of acalypterate Diptera (over 1,000 species of fossil and living forms in
some 50 genera have been described). Adults of living taxa contain some
of the more morphologically bizarre forms of all the Diptera and can be
found on tree trunks and foliage. They are attracted to flowers, decaying
fruit, excrement, sweat, and decomposing snails. Larvae are found on both
fresh and decaying vegetation, carrion, human corpses, and root nodules.
Most larvae are either phytophagous or saprophagous; however, some have
been recorded as predaceous on other insects and others have been observed
feeding on human lesions.
Only two fossil Diptera species have been described in Platystomatidae.
One (Scholastes foordi) is from amber found in Norfolk, UK, which
may be of Pliocene age if the medium is found to be true amber, or Pleistocene
to Holocene in age, if the specimen is found to be copal from Africa (as
the inclusion suggests). The other species (Ceroxys ethiopia), originally
described as a member of an otitid genus from copal of Zanzibar, has been
recently transferred by Crosskey (1980) to the Platystomatidae. This familial
placement is followed here and the species is listed under generically unplaced
Platystomatidae.
*SCHOLASTES Loew, 1873c: 38. Type species: Platystoma cincta
Guérin-Méneville, 1831, by original designation.
foordi Cockerell, 1921a: 30. PA: UK (England) (Pliocene) [A].
[If found to be African copal (as the inclusion suggests), the age will
have to be revised to either Pleistocene or Holocene.]
ethiopia Meunier, 1908g: 254 (Ceroxys). AF: Tanzania (Pleistocene/Holocene)
[K].