Bibionids, commonly called march flies, love bugs, or harlequin
flies, are found in all regions of the globe and comprise some
1,000 living and fossil species. They are the largest family of
fossil Diptera with 344 species described in 12 genera. Adults
of extant forms are commonly found hovering in mating swarms.
Some are nectar and pollen feeders. Immature stages are herbivorous,
with some feeding on roots and tubers. Most larvae breed in rotting
leaves and other decaying vegetation.
The oldest recorded compression fossils of Bibionidae are from
the Upper Triassic deposits of the Dan River formation of North
America (Olsen et al., 1978). Peterson (1975) described a species
of Bibio from Upper Cretaceous Canadian amber, Rayner (1987) discussed
an assemblage of Mid- to Upper Cretaceous bibionids from Botswana,
and Zherichin & Sukacheva (1973) mentioned an undescribed
species of the family from Upper Cretaceous Siberian amber. Other
undescribed fossils include the records of bibionids from the
Pliocene of Turkmenia (Rohdendorf, 1964), Miocene of Austria (Bachmayer,
1974), the Oligocene/Miocene amber of the Dominican Republic (Poinar,
1992), the Upper Cretaceous deposits of the Isle of Mull, Scotland
(Jarzembowski, 1978: 251), and the Lower Cretaceous Santana Formation
of Brazil (Grimaldi, 1990: 178; Maisey, 1991: 422).
Gentilini (1991a) asserted that the majority of Rice's (1959)
species of Plecia may have to be transferred to Penthetria. These
taxa are here left in Plecia until examination of the type material
can be conducted.
Recently, Zhang (1993a) transferred Clothonopsis miocenica
Hong & Wang from the Embioptera to the Bibionidae after restudying
the description and illustrations provided by Hong & Wang
(1987).
Ref.: Rice (1959, review of Canadian taxa); Dürrenfeldt (1968, new German Pliocene species).
*BIBIO Geoffroy, 1762: 568. Type species: Tipula
hortulana Linnaeus, 1758, by subsequent designation of Latreille
(1810: 442) (as "Hirtea hortulana Fabr."). [Validated
by I.C.Z.N. (1957a: 86).]
*BIBIONUS Rafinesque, 1815: 130 (unnecessary new replacement
name for Bibio Linnaeus). Type species: Tipula hortulana
Linnaeus, 1758, automatic.
DICHANEURUM Aymard, 1856: 42. Nomen nudum.
acaptus Dürrenfeldt, 1968: 59. PA: Germany (Pliocene)
[C].
aerosus Statz, 1943: 50. PA: Germany (Oligocene) [C].
alacris Oustalet, 1870: 127. PA: France (Oligocene) [C].
albangulus Dürrenfeldt, 1968: 69. PA: Germany (Pliocene)
[C].
anasiformis Dürrenfeldt, 1968: 70. PA: Germany (Pliocene)
[C].
angustatus Heer, 1849: 215. PA: Switzerland (Miocene) [C].
atavus Cockerell, 1909f: 173. NE: USA (Oligocene) [C].
auripes Statz, 1943: 51. PA: Germany (Oligocene) [C].
aymardi Maneval, 1936: 34. PA: France (Oligocene) [C].
infossum Aymard, 1856: 42 (Dichaneurum). Nomen
nudum.
primaevum Aymard, 1856: 42 (Dichaneurum). Nomen
nudum.
batheri Pongrácz, 1928: 180. PA: Croatia (Miocene)
[C].
baumbergeri Théobald, 1937a: 223. PA: France (Oligocene)
[C].
brachypodus Zhang, 1989a: 351. PA: China (Miocene) [C].
brachypteroides Meunier, 1915c: 215. PA: Germany (Oligocene)
[C].
*brunettii Hardy, 1973: 435 (new replacement name for abdominalis
Brunetti). OR: India; PA: China (Miocene) [C].
*abdominalis Brunetti, 1911: 276. OR: India; PA: China
(Miocene) [C]. [Preoccupied by Fabricius, 1805.]
brunneifemus Zhang, 1989a: 343. PA: China (Miocene) [C].
brunneus Zhang, 1989a: 351. PA: China (Miocene) [C].
bullatus Hong, 1985: 27 (Fungivora). PA: China (Miocene)
[C].
capnodes Melander, 1949: 22. NE: USA (Oligocene) [C].
carnificus Dürrenfeldt, 1968: 59. PA: Germany (Pliocene)
[C].
celasensis Théobald, 1937a: 139. PA: France (Oligocene)
[C].
cockerelli James, 1937: 242. NE: USA (Oligocene) [C].
colossoceps Zhang, 1989a: 347. PA: China (Miocene) [C].
comosella Statz, 1943: 47. PA: Germany (Oligocene) [C].
conformans Théobald, 1937a: 335. PA: France (Oligocene)
[C].
consolidus Théobald, 1937a: 225. PA: France (Oligocene)
[C].
constringtus Dürrenfeldt, 1968: 60. PA: Germany (Pliocene)
[C].
contererus Dürrenfeldt, 1968: 54. PA: Germany (Pliocene)
[C].
cratopodus Zhang, 1989a: 354. PA: China (Miocene) [C].
cruciformis Dürrenfeldt, 1968: 68. PA: Germany (Pliocene)
[C].
curtisii Heer, 1856: 34. PA: France (Oligocene) [C].
cylindratus Oustalet, 1870: 133. PA: France (Oligocene)
[C].
deflectus Dürrenfeldt, 1968: 62. PA: Germany (Pliocene)
[C].
deformatus Zhang, 1989a: 345. PA: China (Miocene) [C].
deletus Heyden, 1859: 14. PA: Germany (Oligocene) [C].
detractus Théobald, 1937a: 334. PA: France (Oligocene)
[C].
discerptus Dürrenfeldt, 1968: 65. PA: Germany (Pliocene)
[C].
disjectus Théobald, 1937a: 334. PA: France (Oligocene)
[C].
dislocatus Théobald, 1937a: 224. PA: France (Oligocene)
[C].
dormitus Dürrenfeldt, 1968: 55. PA: Germany (Pliocene)
[C].
dubius Germar, 1837: pl. 24 (Phthiria). PA: Germany
(Oligocene) [C].
elegantulus Novák, 1877: 85. PA: Czech Republic
(Oligocene) [C].
elongatipennis Théobald, 1937a: 141. PA: France
(Oligocene) [C].
elongatus Heer, 1849: 214. PA: Switzerland (Miocene) [C].
enormus Dürrenfeldt, 1968: 55. PA: Germany (Pliocene)
[C].
enterodelus Unger, 1841: 428. PA: Croatia (Miocene) [C].
excurvatus Melander, 1949: 23. NE: USA (Oligocene) [C].
expansus Zhang, 1989a: 344. PA: China (Miocene) [C].
explanatus Melander, 1949: 23. NE: USA (Oligocene) [C].
extremus Cockerell, 1921c: 468. PA: UK (England) (Eocene/Oligocene)
[C].
firmus Heer, 1849: 226. PA: Croatia (Miocene) [C].
*flavissimus Brunetti, 1925: 448. OR: India; Borneo, Philippines,
AU: New Guinea; PA: China (Miocene) [C].
formosus Novák, 1877: 85. PA: Czech Republic (Oligocene)
[C].
fusiformis Heer, 1849: 219. PA: Switzerland (Miocene) [C].
germari Meunier, 1920a: 652. PA: Germany (Oligocene) [C].
giganteus Unger, 1841: 427. PA: Croatia (Miocene) [C].
gigas Oustalet, 1870: 122. PA: France (Oligocene) [C].
gracilis Unger, 1841: 426. PA: Croatia (Miocene) [C].
gurnetensis Cockerell, 1917b: 375. PA: UK (England) (Eocene/Oligocene)
[C].
heroica Heyden & Heyden, 1865a: 29 (Plecia). PA: Germany
(Oligocene) [C].
inflexus Théobald, 1937a: 404. PA: France (Oligocene)
[C].
infumatus Meunier, 1915c: 214. PA: Germany (Oligocene)
[C].
jamesi Melander, 1949: 24. NE: USA (Oligocene) [C].
janus Heyden, 1870: 257. PA: Germany (Oligocene) [C].
jourascencis Théobald in Piton & Théobald,
1935: 69. PA: France (Oligocene) [C].
kochii Staub, 1883: 280. PA: Germany (Miocene) [C].
larteti Oustalet, 1870: 137. PA: France (Oligocene) [C].
latahensis Lewis, 1969: 110. NE: USA (Miocene) [C].
latiantennatus Dürrenfeldt, 1968: 51. PA: Germany
(Pliocene) [C].
lignarius Germar, 1837: pl. 23. PA: Germany (Miocene) [C];
Croatia (Miocene) [C].
ungeri Heer, 1849: 218. PA: Croatia (Miocene) [C].
linearis Heer, 1849: 214. PA: Croatia (Miocene) [C].
lividus Heer, 1849: 223. PA: Croatia (Miocene) [C].
longineurus Zhang, 1989a: 357. PA: China (Miocene) [C].
luteipennis Théobald, 1937a: 333. PA: France (Oligocene)
[C].
macer Oustalet, 1870: 126. PA: France (Oligocene) [C].
maceratus Théobald, 1937a: 142. PA: France (Oligocene)
[C].
maculatus Heer, 1849: 219. PA: Croatia (Miocene) [C].
marginatus Oustalet, 1870: 125 (as ungeri subspecies).
PA: France (Oligocene) [C].
marmoratus Pongrácz, 1928: 186. PA: Croatia (Miocene)
[C].
martinsi Heer, 1861: 153. Nomen nudum.
medioalbus Dürrenfeldt, 1968: 53. PA: Germany (Pliocene)
[C].
mimas Heyden, 1870: 258. PA: Germany (Oligocene) [C].
minor Oustalet, 1870: 134 (as gracilis subspecies.).
PA: France (Oligocene) [C].
miocenicus Hong & Wang, 1985: 3. PA: China (Miocene)
[C].
moestus Heer, 1849: 224. PA: Germany & France (Miocene)
[C].
morio Heer, 1849: 222. PA: Croatia & France (Miocene)
[C].
incrassatus Heer, 1849: 221. PA: Croatia (Miocene) [C].
murchisonis Unger, 1841: 426. PA: Croatia (Miocene) [C].
nebulosus Dürrenfeldt, 1968: 66. PA: Germany (Pliocene)
[C].
nigripennis Théobald, 1937a: 286. PA: France (Oligocene)
[C]. [Preoccupied by Brunetti, 1913; no new replacement name proposed
here.]
oblitus Cockerell, 1921c: 468. PA: UK (England) (Eocene/Oligocene)
[C].
oblongus Heer, 1849: 227. PA: Switzerland (Miocene) [C].
obtusus Théobald, 1937a: 140. PA: France (Oligocene)
[C].
oeningensis Heer, 1849: 221. PA: Switzerland (Miocene)
[C].
oligocenus Cockerell, 1917b: 375. PA: UK (England) (Eocene/Oligocene)
[C].
painvini Meunier, 1915b: 16. PA: Germany (Oligocene) [C].
parenti Théobald, 1937a: 224. PA: France (Oligocene)
[C].
partschii Heer, 1849: 216. PA: Croatia (Miocene) [C].
penisularis Zhang, 1993a: 144. PA: China (Miocene) [C].
peraffinis Zhang, 1989a: 346. PA: China (Miocene) [C].
pinguis Heer, 1849: 220. PA: Switzerland (Miocene) [C].
podager Melander, 1949: 24. NE: USA (Oligocene) [C].
praecidus Dürrenfeldt, 1968: 61. PA: Germany (Pliocene)
[C].
pulchellus Heer, 1849: 217. PA: Switzerland (Miocene) [C].
radobojanus Heer, 1949: 220 (as pinguis subspecies).
PA: Croatia (Miocene) [C].
robustus Oustalet, 1870: 128. PA: France (Oligocene) [C].
rohli Statz, 1943: 46. PA: Germany (Oligocene) [C].
romani Théobald, 1937a: 336. PA: France (Oligocene)
[C].
rottensis Statz, 1943: 48. PA: Germany (Oligocene) [C].
ruptineurus Zhang, 1989a: 356. PA: China (Miocene) [C].
sadoensis Fujiyama, 1985: 51. PA: Japan (Miocene) [C].
sereri Massalongo, 1856: 21. PA: Italy (Eocene) [C].
serresi Théobald, 1937a: 336. PA: France (Oligocene)
[C].
shanwangensis Hong, 1984a: 3. PA: China (Miocene) [C].
similis Dürrenfeldt, 1968: 64. PA: Germany (Pliocene)
[C]. [Preoccupied by James, 1936; no new replacement name proposed
here.]
soldatus Dürrenfeldt, 1968: 56. PA: Germany (Pliocene)
[C].
spadicus Statz, 1943: 51. PA: Germany (Oligocene) [C].
spectabilis Théobald, 1937a: 333. PA: France (Oligocene)
[C].
sticheli Handlirsch, 1911: 180. PA: Slovenia (Miocene)
[C]; PA: Germany (Oligocene) [C].
pannosus Heyden & Heyden, 1865a: 20. PA: Germany (Oligocene)
[C].
heydeni Meunier, 1920a: 653. PA: Germany (Oligocene) [C].
tenuiapicalis Théobald, 1937a: 40. PA: France (Oligocene)
[C].
tenuis Dürrenfeldt, 1968: 71. PA: Germany (Pliocene)
[C].
tertiarius Heyden, 1862: 78. PA: Germany (Oligocene) [C].
testus Lewis, 1969: 111. NE: USA (Miocene) [C].
tranopterus Zhang, 1989a: 353. PA: China (Miocene) [C].
tripus Dürrenfeldt, 1968: 58. PA: Germany (Pliocene)
[C].
varineurus Zhang, 1989a: 355. PA: China (Miocene) [C].
ventricosus Zhang, 1989a: 341. PA: China (Miocene) [C].
vetus James, 1937: 243. NE: USA (Oligocene) [C].
vixdus Dürrenfeldt, 1968: 67. PA: Germany (Pliocene)
[C].
vulcanius Melander, 1949: 25. NE: USA (Oligocene) [C].
vulpinus Statz, 1943: 49. PA: Germany (Oligocene) [C].
wickhami Cockerell, 1915a: 647. NE: USA (Oligocene) [C].
xylophilus Germar, 1837: pl. 22. PA: Germany (Oligocene)
[C].
Unidentified spp.-PA: Italy (Miocene) [C] (Statz, 1943: 56; Gentilini,
1991b: 44244).
Unidentified spp.-PA: Japan (Pliocene/Pleistocene) [C] (Fujiyama
& Iwao, 1974: 95; Fujiyama, 1980: 28).
*BIBIODES Coquillett, 1904: 171. Type species: Bibiodes halteralis Coquillett, 1904, by original designation.
intermedia James, 1937: 243. NE: USA (Oligocene) [C].
BIBIODITES Cockerell, 1915c: 493. Type species: Bibiodites confluens Cockerell, 1915, by original designation.
confluens Cockerell, 1915c: 493. PA: UK (England) (Eocene/Oligocene) [C].
CLOTHONOPSIS Hong & Wang, 1987: 258. Type species: Clothonopsis miocenica Hong & Wang, 1987, by original designation.
miocenica Hong & Wang, 1987: 258. PA: China (Miocene) [C].
PHILIA Meigen, 1800: 20. Suppressed by I.C.Z.N. (1963:
339).
*DILOPHUS Meigen, 1803: 264. Type species: Tipula febrilis
Linnaeus, 1758, by subsequent designation of Latreille (1810:
442).
campbelli Harris, 1983: AU: New Zealand (Eocene) [C].
krantzii Heyden, 1870: 255. PA: Germany (Oligocene) [C].
magnus Dürrenfeldt, 1968: 50. PA: Germany (Pliocene)
[C].
priscus Loew, 1850b: 38. PA: Baltic Region (Eocene/Oligocene)
[A].
Unidentified sp.-PA: France (Oligocene) [C] (Statz, 1943: 56).
*HESPERINUS Walker, 1848: 81. Type species: Hesperinus brevifrons Walker, 1848, by monotypy.
immutabilis Melander, 1949: 20. NE: USA (Oligocene) [C].
LITHOBIBIO Beier, 1952: 133. Type species: Lithobibio styriacus Beier, 1952, by monotypy.
styriacus Beier, 1952: 133. PA: Austria (Miocene) [C].
MESOMYIA Pongrácz, 1928: 174. Type species: Bibio
brevis Heer, 1849, by monotypy. [Preoccupied by Macquart,
1850.]
LITHOSOMYIA Carpenter, 1986: 576 (new replacement name
for Mesomyia Pongrácz). Type species: Bibio brevis
Heer, 1849, automatic.
brevis Heer, 1849: 225 (Bibio). PA: Switzerland (Miocene) [C].
MEGEANA Meunier, 1899e: 175. Type species: Megeana
hardyi Evenhuis, 1994, by present designation.
[Megeana originally proposed without included species.
The type designated is from first inclusion of a named species
within the genus in accordance with the Code.]
hardyi Evenhuis, 1994: in this work (new species). PA:
Baltic Region (Eocene/Oligocene) [A].
[Species-group name validated by bibliographic reference to characters
given in Meunier (1899e: 175) for the genus Megeana.]
*PENTHETRIA Meigen, 1803: 264. Type species: Penthetria
funebris Meigen, 1804, by subsequent monotypy in Meigen (1804:
104).
PROTOMYIA Heer, 1849: 231. Type species: Protomyia lygaeoides
Heer, 1849, by subsequent designation of Carpenter (1992: 414).
EPIPLECIA Giard, 1879: 13. Type species: Protomyia joannisi
Oustalet, 1870, by monotypy.
abacula Lewis, 1971b: 1466. NE: USA (Oligocene) [C].
adusta Oustalet, 1870: 145 (Protomyia). PA: France
(Oligocene) [C].
aestimata Statz, 1943: 39. PA: Germany (Oligocene) [C].
affinis Heer, 1849: 235 (Protomyia). PA: Switzerland
(Miocene) [C].
alderensis Lewis, 1971b: 1466. NE: USA (Oligocene) [C].
amoena Heer, 1849: 237 (Protomyia). PA: Switzerland
(Miocene) [C].
anthracina Heer, 1849: 236 (Protomyia). PA: Switzerland
(Miocene) [C].
blanchardi Oustalet, 1870: 148 (Protomyia). PA:
France (Oligocene) [C].
bohemica Novák, 1877: 86 (Protomyia). PA:
Czech Republic (Oligocene) [C].
brevipennis Heer, 1856: 35 (Protomyia). PA: France
(Oligocene) [C].
bucklandi Heer, 1849: 238 (Protomyia). PA: Croatia
(Miocene) [C].
dubia Geinitz, 1884: 582 (Protomyia). PA: Germany
(Lower Jurassic) [C].
edwardsi Oustalet, 1870: 130 (Bibio). PA: France
(Oligocene) [C].
elegans Heer, 1856: 36 (Protomyia). PA: France (Oligocene)
[C].
formicoides Oustalet, 1870: 150 (Protomyia). PA:
France (Oligocene) [C].
fryi Rice, 1959: 7. NE: Canada (Eocene) [C].
fuchsi Handlirsch, 1907c: 961. PA: Greece (Miocene) [C].
marinosi Bachmayer et al., 1971: 170. PA: Greece
(Miocene) [C].
fusca Oustalet, 1870: 145 (Protomyia). PA: France
(Oligocene) [C].
globularis Oustalet, 1870: 147 (Protomyia). PA:
France (Oligocene) [C].
gracilis Heer, 1856: 36 (Protomyia). PA: France
(Oligocene) [C].
incerta Oustalet, 1870: 151 (Protomyia). PA: France
(Oligocene) [C].
inflata Oustalet, 1870: 142 (Protomyia). PA: France
(Oligocene) [C].
joannisi Oustalet, 1870: 143 (Protomyia). PA: France
(Oligocene) [C].
jucunda Heer, 1849: 234 (Protomyia). PA: Croatia
(Miocene) [C].
latipennis Heer, 1849: 237 (Protomyia). PA: Croatia
(Miocene) [C].
livida Heer, 1856: 35 (Protomyia). PA: France (Oligocene)
[C].
longa Heer, 1849: 233 (Protomyia). PA: Croatia (Miocene)
[C].
longifurca Melander, 1949: 20. NE: USA (Oligocene) [C].
longipennis Oustalet, 1870: 141 (Protomyia). PA:
France (Oligocene) [C].
lugens Oustalet, 1870: 142 (Protomyia). PA: France
(Oligocene) [C].
lygaeoides Heer, 1849: 232 (Protomyia). PA: Croatia
& France (Miocene) [C].
major Oustalet, 1870: 114 (Plecia). PA: France (Oligocene)
[C].
matheroni Heer, 1861: 153. Nomen nudum.
nigra Statz, 1943: 39. PA: Germany (Oligocene) [C].
nigrescens Oustalet, 1870: 115 (Plecia). PA: France
(Oligocene) [C].
oustaleti Brongniart, 1876a: 1 [1876b: 459] (Protomyia).
PA: France (Oligocene) [C].
pallida Oustalet, 1870: 118 (Plecia). PA: France (Oligocene)
[C].
parschlugiana Heer, 1849: 235 (Protomyia; as jucunda
subspecies). PA: Switzerland (Miocene) [C]; PA: Austria (Miocene)
[C].
rottensis Statz, 1943: 40. PA: Germany (Oligocene) [C].
rubescens Oustalet, 1870: 149 (Protomyia). PA: France (Oligocene)
[C].
rubiensis Lewis, 1971b: 1465. NE: USA (Oligocene) [C].
sauvagii Oustalet, 1870: 146 (Protomyia). PA: France
(Oligocene) [C].
scita Statz, 1943: 40. PA: Germany (Oligocene) [C].
speciosa Heer, 1864: 394 (Protomyia). PA: Switzerland
(Miocene) [C].
togoensis Fujiyama & Iwao, 1974: 93. PA: Japan (Pliocene/Pleistocene)
[C].
tropica Statz, 1943: 39. PA: Germany (Oligocene) [C].
vaillanti Oustalet, 1870: 112. PA: France (Oligocene) [C].
whipsawensis Rice, 1959: 7. NE: Canada (Eocene) [C].
Unidentified sp.-PA: Italy (Miocene) [C] (Gentilini, 1991a: 65).
*PLECIA Wiedemann, 1828: 72. Type species: Hirtea
fulvicollis Fabricius, 1805, by subsequent designation of
Blanchard (1840: 576).
BIBIOPSIS Heer, 1849: 28. Type species: Bibiopsis cimicoides
Heer, 1849, by subsequent designation of Carpenter (1992: 414).
MYCETOPHAETUS Scudder, 1892: 20. Type species: Mycetophaetus
intermedius Scudder, 1892, by monotypy.
abava Heyden & Heyden, 1865a: 20 (Protomyia).
PA: Germany (Oligocene) [C].
acourti Cockerell, 1921c: 467. PA: UK (England) (Eocene/Oligocene)
[C].
aculeolata Zhang, 1989a: 332. PA: China (Miocene) [C].
acutiventris Théobald, 1937a: 410. PA: France (Oligocene)
[C].
amagua Cockerell, 1925c: 13. PA: Russia (Amur) (Oligocene/Miocene)
[C].
angustipennis Handlirsch, 1910: 104 (Penthetria).
NE: Canada (Eocene) [C].
angustiventris Théobald, 1937a: 135. PA: France
(Oligocene) [C].
antenata Heyden & Heyden, 1865a: 26 (Protomyia).
PA: Germany (Oligocene) [C].
assonensis Gentilini, 1993: 161. PA: France (Oligocene)
[C].
avunculus Handlirsch, 1910: 104 (Penthetria). NE:
Canada (Eocene) [C].
avus Handlirsch, 1910: 115 (Penthetria). NE: Canada
(Eocene) [C].
axeliana Cockerell, 1915a: 646. NE: USA (Oligocene) [C].
baglii Gentilini, 1991a: 433. PA: Italy (Miocene) [C].
bivalvula Zhang, 1993a: 142. PA: China (Miocene) [C].
borussica Meunier, 1907b: 395. PA: Baltic Region (Eocene/Oligocene)
[A].
brevicollis Heer, 1849: 231 (Bibiopsis). PA: Croatia
(Miocene) [C].
brevipes Handlirsch, 1910: 106 (Penthetria). NE:
Canada (Eocene) [C].
brevithoracis Théobald, 1937a: 409. PA: France (Oligocene)
[C].
brunneipennis Piton, 1939: 107. PA: France (Oligocene)
[C].
cairnesi Rice, 1959: 11. NE: Canada (Eocene) [C].
canadensis Handlirsch, 1910: 118 (Penthetria). NE:
Canada (Eocene) [C].
carbonum Heyden & Heyden, 1865b: 34 (Bibiopsis).
PA: Germany (Miocene) [C].
catellaroi Gentilini, 1991a: 427. PA: Italy (Miocene) [C].
cerestensis Théobald, 1937a: 409. PA: France (Oligocene)
[C].
cimicoides Heer, 1849: 229 (Bibiopsis). PA: Croatia
(Miocene) [C].
claripennis Théobald, 1937a: 228. PA: France (Oligocene)
[C].
collossea Heyden & Heyden, 1865a: 21 (Protomyia).
PA: Germany (Oligocene) [C].
conica Théobald, 1937a: 232. PA: France (Oligocene)
[C].
constricta Théobald, 1937a: 342. PA: France (Oligocene)
[C].
crassicornis Keilbach, 1982: 347. Nomen nudum.
crassiventris Théobald, 1937a: 227. PA: France (Oligocene)
[C].
creedensis James, 1938a: 114. NE: USA (Eocene) [C].
curtula Handlirsch, 1910: 119 (Penthetria). NE:
Canada (Eocene) [C].
decapitata Cockerell, 1917e: 391. NE: USA (Oligocene) [C].
dejecta Scudder, 1890: 586. NE: USA (Eocene) [C].
diatoma Zhang, 1989a: 331. PA: China (Miocene) [C].
difficilis Théobald, 1937a: 234. PA: France (Oligocene)
[C].
dilatata Handlirsch, 1910: 120 (Penthetria). NE:
Canada (Eocene) [C].
dilatativentris Théobald, 1937a: 338. PA: France
(Oligocene) [C].
dumasi Théobald, 1937a: 138. PA: France (Oligocene)
[C].
egerana Novák, 1877: 83 (Bibiopsis). PA:
Czech Republic (Oligocene) [C].
elegantula Meunier, 1915d: 226 (Protomyia). PA:
Germany (Oligocene) [C].
eliator Handlirsch, 1910: 108 (Penthetria). NE:
Canada (Eocene) [C].
elongata Heyden & Heyden, 1865a: 27 (Protomyia).
PA: Germany (Oligocene) [C].
exigua Statz, 1943: 26. PA: Germany (Oligocene) [C].
explanata Cockerell, 1917b: 376. NE: USA (Oligocene) [C].
exposita Heyden & Heyden, 1865a: 24 (Protomyia).
PA: Germany (Oligocene) [C].
falcatula Handlirsch, 1910: 110 (Penthetria). NE:
Canada (Eocene) [C].
flichei Théobald, 1937a: 406. PA: France (Oligocene)
[C].
foersteri Théobald, 1937a: 230. PA: France (Oligocene)
[C].
fragmentum Handlirsch, 1910: 111 (Penthetria). NE:
Canada (Eocene) [C].
fusciventris Théobald, 1937a: 407. PA: France (Oligocene)
[C].
fushunensis Hong, 1980: 46 [1981: 76]. PA: China (Eocene)
[A].
gigantea Théobald, 1937a: 226. PA: France (Oligocene)
[C].
goreti Théobald, 1937a: 409. PA: France (Oligocene)
[C].
gracilenta Heyden & Heyden, 1865a: 28 (Protomyia).
PA: Germany (Oligocene) [C].
gracilentula Evenhuis, 1994: in this work (new replacement
name for gracilenta Zhang). PA: China (Miocene) [C].
gracilenta Zhang, 1993a: 143. PA: China (Miocene) [C].
[Preoccupied by Heyden & Heyden, 1865.] New synonymy.
graciliventris Théobald, 1937a: 138. PA: France
(Oligocene) [C].
gracillima Förster, 1891: 480. PA: Germany (Miocene)
[C].
graciosa Théobald, 1937a: 235. PA: France (Oligocene)
[C].
gradata Melander, 1949: 21. NE: USA (Oligocene) [C].
grandaeva Heyden & Heyden, 1865a: 25 (Protomyia).
PA: Germany (Oligocene) [C].
grossa Heyden & Heyden, 1865a: 22 (Protomyia).
PA: Germany (Oligocene) [C].
guinieri Théobald, 1937a: 408. PA: France (Oligocene)
[C].
haennii Gentilini, 1993: 164. France (Oligocene) [C].
heeri Heyden & Heyden, 1865a: 28 (Protomyia).
PA: Germany (Oligocene) [C].
hilaris Heer, 1849: 211. PA: Switzerland (Miocene) [C].
hypogaea Heyden & Heyden, 1865a: 23 (Protomyia).
PA: Germany (Oligocene) [C].
imperialis Novák, 1877: 84 (Bibiopsis). PA:
Czech Republic (Oligocene) [C]. [Preoccupied by Schiner, 1868;
no new replacement name proposed here.]
imperialis Statz, 1943: 13. PA: Germany (Oligocene) [C].
[Preoccupied by Schiner, 1868; no new replacement name proposed
here.]
inflata Hull, 1960: 274. NE: USA (Oligocene) [C].
intermedia Scudder, 1892: 20 (Mycetophaetus). NE:
USA (Oligocene) [C].
intima Fujiyama & Iwao, 1974: 94. PA: Japan (Pliocene/Pleistocene)
[C].
kanetakii Fujiyama, 1970: 69. PA: Japan (Miocene) [C].
kelownaensis Rice, 1959: 8. NE: Canada (Eocene) [C].
kudiella Cockerell, 1925c: 14. PA: Russia (Amur) (Oligocene/Miocene)
[C].
kuznetzovi Cockerell, 1925c: 15. PA: Russia (Amur) (Oligocene/Miocene)
[C].
lambei Handlirsch, 1910: 116 (Penthetria). NE: Canada
(Eocene) [C].
lapidaria Heyden & Heyden, 1865a: 25 (Protomyia).
PA: Germany (Oligocene) [C].
sluiteri Meunier, 1917b: 14 (Protomyia). PA: Germany
(Oligocene) [C].
larguensis Gentilini, 1993: 162. PA: France (Oligocene)
[C].
longiventris Théobald, 1937a: 133. PA: France (Oligocene)
[C].
luctuosa Heyden & Heyden, 1865a: 22 (Protomyia).
PA: Germany (Oligocene) [C].
ludongensis Hong & Wang, 1985: 5. PA: China (Miocene)
[C].
luteipennis Théobald, 1937a: 233. PA: France (Oligocene)
[C].
luteola Heyden & Heyden, 1865a: 26 (Protomyia).
PA: Germany (Oligocene) [C].
macrocephala Heyden & Heyden, 1865a: 23 (Protomyia).
PA: Germany (Oligocene) [C].
macroventris Théobald, 1937a: 229. PA: France (Oligocene)
[C].
maculiventris Théobald, 1937a: 411. PA: France (Oligocene)
[C].
maimensis Gentilini, 1993: 158. PA: France (Oligocene)
[C].
melanderi Cockerell, 1911: 77. NE: USA (Oligocene) [C].
miegi Théobald, 1937a: 232. PA: France (Oligocene)
[C].
minutula Rice, 1959: 8. NE: Canada (Eocene) [C].
myersi Peterson, 1975: 712. NE: Canada (Upper Cretaceous)
[A].
nana Handlirsch, 1910: 111 (Penthetria). NE: Canada
(Eocene) [C].
nervisinuata Théobald, 1937a: 339. PA: France (Oligocene)
[C].
nigra Statz, 1943: 25 (as lapidaria subspecies).
PA: Germany (Oligocene) [C]. [Preoccupied by Philippi, 1865; no
new replacement name proposed here.]
obsitula Cockerell, 1925c: 15. PA: Russia (Amur) (Oligocene/Miocene)
[C].
obsoleta Heer, 1849: 227 (Bibio). PA: Switzerland
(Miocene) [C].
orycta Melander, 1949: 21. NE: USA (Oligocene) [C].
ovalis Handlirsch, 1910: 117 (Penthetria). NE: Canada
(Eocene) [C].
oviformis Théobald, 1937a: 229. PA: France (Oligocene)
[C].
pealei Scudder, 1890: 585. NE: USA (Eocene) [C].
pennata Statz, 1943: 19. PA: Germany (Oligocene) [C].
philippi Statz, 1943: 10. PA: Germany (Oligocene) [C].
pictipennis Handlirsch, 1910: 106 (Penthetria).
NE: Canada (Eocene) [C].
pinguis Heyden & Heyden, 1865a: 24 (Protomyia).
PA: Germany (Oligocene) [C].
pisaurensis Gentilini, 1991a: 430. PA: Italy (Miocene)
[C].
platoptera Zhang, 1993a: 141. PA: China (Miocene) [C].
platyptera Handlirsch, 1910: 121 (Penthetria). NE:
Canada (Eocene) [C].
prisca Meunier, 1899e: 176. PA: Baltic Region (Eocene/Oligocene)
[A].
pristina Hardy, 1971: 65. NT: Mexico (Oligocene/Miocene)
[A].
proserpina Heyden & Heyden, 1865a: 22 (Protomyia).
PA: Germany (Oligocene) [C].
pulchella Meunier, 1915c: 216. PA: Germany (Oligocene)
[C].
pulchra Handlirsch, 1910: 113 (Penthetria). NE:
Canada (Eocene) [C].
pulla Handlirsch, 1910: 105 (Penthetria). NE: Canada
(Eocene) [C].
quaesita Novák, 1877: 86. PA: Czech Republic (Oligocene)
[C].
rectivena Zhang, 1989a: 335. PA: China (Miocene) [C].
redempta Cockerell, 1925c: 14. PA: Russia (Amur) (Oligocene/Miocene)
[C].
reducta Handlirsch, 1910: 109 (Penthetria). NE:
Canada (Eocene) [C].
refracta Cockerell, 1925c: 14. PA: Russia (Amur) (Oligocene/Miocene)
[C].
retracta Théobald, 1937a: 341. PA: France (Oligocene)
[C].
rhenana Heyden & Heyden, 1865a: 28. PA: Germany (Oligocene)
[C].
rhodopterina Cockerell, 1924a: 6. NE: USA (Eocene) [C].
satyrus Statz, 1943: 12. PA: Germany (Oligocene) [C].
schineri Heyden & Heyden, 1865a: 27 (Protomyia).
PA: Germany (Oligocene) [C].
separanda Handlirsch, 1910: 112 (Penthetria). NE:
Canada (Eocene) [C].
similkameena Scudder, 1879: 177 (Penthetria). NE:
Canada (Miocene) [C].
solaris Zhang, 1989a: 334. PA: China (Miocene) [C].
spinula Zhang, 1989a: 329. PA: China (Miocene) [C].
splendida Théobald, 1937a: 137. PA: France (Oligocene)
[C].
sturmi Statz, 1943: 20. PA: Germany (Oligocene) [C].
stygia Heyden & Heyden, 1865a: 24 (Protomyia).
PA: Germany (Oligocene) [C].
subterranea Théobald, 1937a: 233. PA: France (Oligocene)
[C].
superba Meunier, 1915c: 207. PA: Germany (Oligocene) [C].
superba Théobald, 1937a: 134. PA: France (Oligocene)
[C]. [Preoccupied by Meunier, 1915; no new replacement name proposed
here.]
tessella Melander, 1949: 21. NE: USA (Oligocene) [C].
transitoria Handlirsch, 1910: 117 (Penthetria).
NE: Canada (Eocene) [C].
tulameenensis Rice, 1959: 11. NE: Canada (Eocene) [C].
undans Zeuner, 1941: 94. PA: UK (Scotland) (Upper Cretaceous)
[C].
varia Théobald, 1937a: 234. PA: France (Oligocene)
[C].
variegata Théobald, 1937a: 234. PA: France (Oligocene)
[C].
vergnei Piton in Piton & Théobald, 1935:
86. PA: France (Oligocene) [C].
veterana Heyden & Heyden, 1865a: 25 (Protomyia). PA:
Germany (Oligocene) [C].
villosa Zhang, 1989a: 333. PA: China (Miocene) [C].
volgeri Heyden, 1859: 15 (Bibiopsis). PA: Germany
(Oligocene) [C].
vulcania Zhang, 1989a: 331. PA: China (Miocene) [C].
vulpina Statz, 1943: 27. PA: Germany (Oligocene) [C].
winchesteri Cockerell, 1917b: 375. NE: USA (Eocene) [C].
winnertzi Heyden & Heyden, 1865a: 21 (Protomyia). PA:
Germany (Oligocene) [C].
woodruffi Cockerell, 1916c: 89. NE: USA (Eocene) [C].