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Family BLEPHARICERIDAE
Blepharicerids, commonly called net-winged
midges, are slender, small to medium-length nematocerous flies whose extant
forms are most often found in association with swift moving streams where they
rest on overhanging vegetation, bridges, and the like. Immature stages of the
extant forms are aquatic, and live in the waters of the swift moving streams
where they attach themselves to hard, smooth surfaces via ventral suckers.
Larvae are grazers; most adult males do not feed; some females feed on other
insects that they catch with specialized hind tarsi.
The described fossils
listed here may not belong to the Blephariceridae. Alexander (1958) alleged
that those fossils that were previously described in this family may belong in
other nematocerous families: Paltostomopsis Cockerell
possibly being a tipulid and Philorites Cockerell
possibly being a bibionid. This view was subsequently
followed by Hennig (1969c). These two genera are listed here provisionally
pending further study to ascertain their true position within the Diptera.
Despite these two
questionable genera, since publication of the 1994 catalog there have been a
number of true blepharicerids described from Mesozoic
strata and a few different localities. In addition to those described taxa
listed below, undetermined material identified as Blephariceridae has been
recorded from the Upper Cretaceous amber deposits of Burma (Grimaldi et al., 2002) and the Eocene/Oligocene
deposits of the Isle of Wight (Jarzembowski, 1978: 251).
Ref:: Carpenter
(1992, world fossils).
Genus BLEPHADEJURA Lukashevich, Huang & Lin
BLEPHADEJURA Lukashevich, Huang & Lin, 2006: 132.
type species Blephadejura propria
Lukashevich, Huang & Lin, 2006, by original designation.
propria Lukashevich,
Huang & Lin, 2006: 133. PA: China (Middle Jurassic) [C].
Genus BRIANINA Zhang & Lukashevich
BRIANINA Zhang & Lukashevich, 2007: 303. Type
species: Brianina longitibialis
Zhang & Lukashevich
longitibialis
Zhang & Lukashevich, 2007: 303. PA: China (Middle Jurassic) [C].
Genus MEGATHON Lukashevich & Shcherbakov
MEGATHON Lukashevich & Shcherbakov, 1997: 640. Type
species: Megathon zwicki
Lukashevich & Shcherbakov, 1997, by original designation.
brodskyi
Zhang & Lukashevich, 2007: 103. PA: China (Middle Jurassic) [C].
zwicki
Lukashevich & Shcherbakov, 1997: 642. PA: Russia (Siberia) (Upper
Cretaceous) [C].
Questionably
Included in BLEPHARICERIDAE
Genus PALTOSTOMOPSIS
Cockerell
PALTOSTOMOPSIS Cockerell, 1915c: 489. Type species: Paltostomopsis ciliatus Cockerell, 1915, by
original designation.
ciliata Cockerell, 1915c: 490. PA: UK (England) (Eocene/Oligocene) [C].
Genus PHILORITES
Cockerell
PHILORITES Cockerell, 1908f: 264. Type species: Philorites
johannseni Cockerell, 1908, by monotypy.
johannseni Cockerell, 1908f: 264. NE: USA (Eocene)
[C].
pallescens Cockerell, 1920a: 250. NE: USA (Eocene) [C].