Fossil Diptera Catalog - Dolichopodidae

Family DOLICHOPODIDAE

Dolichopodids, commonly called long-legged flies, are found worldwide and consist of over 7,300 species in over 250 genera (Pape et al., 2011). Adults and most larvae are predaceous on other insects and small invertebrates. Adults of extant forms inhabit moist, cool, sylvan habitats near water sources where they can be found resting on tree trunks, vegetation, or skating on the water surface in pursuit of prey items. Larvae are found in rotting vegetation, wet soil, and under bark where they are scavengers or predators.

  The Dolichopodidae are well represented in the fossil record, especially as amber inclusions. Although only Baltic amber material of this family has been described to any extent, the family is present in most other faunas as well (Spahr, 1985). Many adults of this family are tree associates and patrol up and down trunks in search of both mates and food. Hence, they are especially susceptible to being entrapped in tree resins and fossilized in amber. Modern trunk-associated dolichopodids include most Medeterinae (especially Medetera Fischer von Waldheim and Systenus Loew), all Neurigoninae, Achalcus Loew, many Sciapodinae (especially Sciapus Zeller s. str.), and some Diaphorinae. All of these taxa occur in Baltic amber.

  As seen in Baltic amber faunas generally (Larsson, 1978), most recent insect families had radiated by the Early Tertiary. This is the case with the Dolichopodidae, and genera appear well defined and essentially modern at that time. However, the Baltic amber fauna described by Meunier is badly in need of revision, especially in light of large unworked collections of fossils in many institutions. Particular attention must be paid to Meunier’s generic concepts and species placements. Hong (2002) described 16 genera and 14 new species of dolichopodids from the Eocene Fushun amber of China; however, unfortunately, most of the taxa proposed are unavailable because all the new species proposed in the work failed to specify a type depository. See the Introduction to this catalog for more details.

  In addition to the determined material listed below, undetermined material of this family has been recorded from the Lower Cretaceous amber of Burma (Grimaldi et al., 2002), the Upper Cretaceous ambers of Canada (McAlpine & Martin, 1969: 837) and Siberia (Zherichin & Sukacheva, 1973: 18, 37), the Miocene amber of the Dominican Republic and the Miocene amber of Chiapas, Mexico (Poinar, 1992), the Lower Miocene deposits of Sado Island, Japan (Fujiyama, 1985: 43), the Miocene Bitterfeld amber of Germany (Schumann & Wendt, 1989), the Eocene/Oligocene deposits of the Bembridge Marls on the Isle of Wight (Jarzembowski, 1976: 12), and Pliocene deposits of Hungary (Krzeminski et al., 1997).

Ref.: Meunier (1907cj,1908ad, f; review of Baltic amber taxa).

Genus ACHALCUS Loew

*ACHALCUS Loew, 1857: 30. Type species: Porphyrops flavicollis Meigen, 1824, by subsequent designation of Coquillett (1910: 502).

latipennis Meunier, 1907d: 209 [1907g: 245]. PA: Baltic Region (Eocene) [A]. latipennis Meunier, 1907c: 198. Nomen nudum.

Genus ANEPSIOMYIA Bezzi

*ANEPSIUS Loew, 1857: 45. Type species: Porphyrops flaviventris Meigen, 1824, by original designation. [Preoccupied by Lec, 1851.] 

*ANEPSIOMYIA Bezzi, 1902: 192 (new replacement name for Anepsius Loew, 1857). Type species: Porphyrops flaviventris Meigen, 1824, automatic.

planipedia Meunier, 1907e: 221 [1908f: 58] (Anepsius). PA: Baltic Region (Eocene) [A]. 

planipidius Meunier, 1907c: 198. Nomen nudum.

Genus ARCHICHRYSOTUS Negrobov

ARCHICHRYSOTUS Negrobov, 1978: 84 (1979: 223). Type species: Archichrysotus hennigi Negrobov, 1978, by original designation.

hennigi Negrobov, 1978: 85 (1979: 224). PA: Russia (Siberia) (Upper Cretaceous) [A]. 

incompletus Grimaldi & Cumming, 1999: 65. NE: USA (New Jersey) (Upper Cretaceous) [A].

manitobus Grimaldi & Cumming, 1999: 65. NE: Canada (Upper Cretaceous) [A].

minor Negrobov, 1978: 86 (1979: 225). PA: Russia (Siberia) (Upper Cretaceous) [A].

Genus ARGYRA Macquart

*ARGYRA Macquart, 1834: 456. Type species: Musca diaphana Fabricius, 1775, by subsequent designation of Westwood (1840: 134).

[Porphyrops Meigen, 1824 has priority over Argyra Macquart, 1834. However, acceptance of priority would upset the stability of nomenclature and taxonomy by replacing the commonly used Argyra with the lesser-known Porphyrops. Application to the ICZN to suppress Porphyrops has been submitted (Evenhuis et al., 2012).]

arguta Meunier, 1907d: 209 [1907f: 234] (Porphyrops). PA: Baltic Region (Eocene) [A]. 

argutus Meunier, 1907c: 198 (Porphyrops). Nomen nudum. 

debellata Meunier, 1907d: 210 [1908c: 29]. PA: Baltic Region (Eocene) [A]. 

debellata Meunier, 1907c: 198. Nomen nudum. 

debilis Meunier, 1907d: 210 [1908c: 30]. PA: Baltic Region (Eocene) [A]. 

debilis Meunier, 1907c: 198. Nomen nudum. 

deceptoria Meunier, 1907d: 210 [1908c: 30]. PA: Baltic Region (Eocene) [A]. 

deceptoria Meunier, 1907c: 198. Nomen nudum. 

inclyta Meunier, 1907d: 209 [1907f: 234] (Porphyrops). PA: Baltic Region (Eocene) [A]. 

inclytus Meunier, 1907c: 198. Nomen nudum. 

mutabilis Meunier, 1907d: 210 [1908c: 30]. PA: Baltic Region (Eocene) [A]. 

mutabilis Meunier, 1907c: 198. Nomen nudum. 

succinorum Meunier, 1907d: 209 [1907f: 233] (Porphyrops). PA: Baltic Region (Eocene) [A]. 

succinorum Meunier, 1907c: 198 (Porphyrops). Nomen nudum.

Genus ARPACTODOLICHOPODITES Hong

ARPACTODOLICHOPODITES Hong, 2002: 355. Unavailable name; genus-group name based on an unavailable species-group name.

eocenicus Hong, 2002: 355. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus AVENAPHORA Grimaldi & Cumming

AVENAPHORA Grimaldi & Cumming, 1999: 61. Type species: Avenophora hispida Grimaldi & Cumming, 1999, by original designation.

hispida Grimaldi & Cumming, 1999: 61. PA: Lebanon (Lower Cretaceous) [A].

Genus BICERITES Hong

BICERITES Hong, 2002: 392. Unavailable name; genus-group name based on an unavailable species-group name.

fushunensis Hong, 2002: 392. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus CAMPSICNEMUS Haliday

CAMPTOSCELES Haliday, 1832: 357 (as Medeterus subgenus). Suppressed by I.C.Z.N. (1958b: 349). 

*CAMPSICNEMUS Haliday in Walker, 1851: 187. Type species: Dolichopus scambus Fallén, 1823, by validation of I.C.Z.N. (1958b: 351). 

*CAMPTOSCELUS Kertész, 1909c: 306 (unjustified emendation of Camptosceles Haliday). Type species: Dolichopus scambus Fallén, 1823, automatic.

gracilis Meunier, 1907e: 221 [1908f: 57]. PA: Baltic Region (Eocene) [A]. 

gracilis Meunier, 1907c: 198. Nomen nudum.

Genus CHAETOGONOPTERON de Meijere

*CHAETOGONOPTERON de Meijere, 1914: 96. Type species: Chaetogonopteron appendiculatum de Meijere, 1914, by monotypy.

bethnorrisae Bickel, 2009: 36. AU: Australia (Queensland) (Miocene) [A].

sobrius Meunier, 1910c: 142 (Teuchophorus). AF: Tanzania (Pleistocene/Holocene) [K].

Genus CHRYSOTUS Meigen

*CHRYSOTUS Meigen, 1824: 40. Type species: Musca nigripes Fabricius, 1794, by subsequent designation of Westwood (1840: 134).

antipellus Keilbach, 1982: 375. Nomen nudum. 

apicalis Keilbach, 1982: 375. Nomen nudum. 

ciliatus Keilbach, 1982: 375. Nomen nudum. [Preoccupied by Malloch, 1914.] 

dasycerus Keilbach, 1982: 375. Nomen nudum. 

decorus Meunier, 1907d: 209 [1907i: 269]. PA: Baltic Region (Eocene) [A]. 

decorus Meunier, 1907c: 198. Nomen nudum. 

delecocerus Keilbach, 1982: 375. Nomen nudum. 

furcatus Keilbach, 1982: 375. Nomen nudum. 

lepidus Meunier, 1907d: 209 [1907i: 270]. PA: Baltic Region (Eocene) [A]. 

lepidus Meunier, 1907c: 198. Nomen nudum. 

praeconcinnus Evenhuis, 1994b: 360 (new replacement name for concinnus Meunier). PA: Baltic Region (Eocene) [A]. 

concinnus Meunier, 1907c: 198. Nomen nudum. [Preoccupied by Zetterstedt, 1843.] 

concinnus Meunier, 1907d: 209 [1907i: 270]. PA: Baltic Region (Eocene) [A]. [Preoccupied by Zetterstedt, 1843.] 

semiciliatus Keilbach, 1982: 375. Nomen nudum. 

setosus Giebel, 1856: 205. PA: Baltic Region (Eocene) [A]. 

stanciui Parent, 1936: 211. PA: Romania (Oligocene) [A]. 

stylatus Keilbach, 1982: 375. Nomen nudum. 

terminaeus Keilbach, 1982: 375. Nomen nudum. 

Unidentified sp. - PA: UK (England) (Pliocene) [A] (Reid, 1884: 602). 

Unidentified sp. - PA: Poland (Pleistocene) [C] (Lomnicki, 1894: 100).

Genus COLUMNIOCORNA Hong

COLUMNIOCORNA Hong, 2002: 388. Unavailable name; genus-group name based on an unavailable species-group name.

latilabrosa Hong, 2002: 389. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus CONVEXIVERTEX Hong

CONVEXIVERTEX Hong, 2002: 351. Unavailable name; genus-group name based on an unavailable species-group name.

viridulus Hong, 2002: 351. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus CRETOMICROPHORUS Negrobov

CRETOMICROPHORUS Negrobov, 1978: 82 (1979: 222). Type species: Cretomicrophorus rohdendorfi Negrobov, 1978, by original designation. 

JANTARDACHIA Zherichin, 1978: 86. Nomen nudum.

rohdendorfi Negrobov, 1978: 83 (1979: 222). PA: Russia (Siberia) (Upper Cretaceous) [A].

Genus CURVUS Kaddumi

CURVUS Kaddumi, 2005b: 85 (2007: 121). Type species: Curvus khuludi Kaddumi, 2005, by original designation.

khuludi Kaddumi, 2005b: 85 (2007: 121; as “khuludae”). PA: Jordan (Lower Cretaceous) [A].

Genus DIAPHORUS Meigen

*DIAPHORUS Meigen, 1824: 32. Type species: Diaphorus flavocinctus Meigen, 1824 [= Dolichopus oculatus Fallén, 1823], by subsequent designation of Westwood (1840: 134). 

*LYRONEURUS Loew, 1857: 38. Type species: Lyroneurus coerulescens Loew, 1857, by subsequent designation of Coquillett (1910: 563).

alsiosus Meunier, 1910c: 141 (Nematoproctus). AF: Tanzania (Pleistocene/Holocene) [K]. 

fernandi Meuffels & Grootaert, 1999: 290 (new replacement name for soccatus Meunier). PA: Baltic Region (Eocene) [A].

soccatus Meunier, 1899e: 179 (Dolichopus). PA: Baltic Region (Eocene) [A]. [Preoccupied in Dolichopus by Walker, 1849.] 

soccatus Keilbach, 1982: 377. Nomen nudum. 

modestus Keilbach, 1982: 377. Nomen nudum. 

tertiarius Meunier, 1907d: 210 [1908a: 8]. PA: Baltic Region (Eocene) [A].

tertiarius Meunier, 1907c: 198. Nomen nudum. 

venustus Meunier, 1907i: 269 (Lyroneurus). PA: Baltic Region (Eocene) [A]. 

venustus Meunier, 1907c: 198. Nomen nudum.

Genus DOLICHOPUS Latreille

*DOLICHOPUS Latreille, 1797: 159. Type species: Musca ungulata Linnaeus, 1758, by subsequent designation of Latreille (1810: 443). 

*HYGROCELEUTHUS Loew, 1857: 10. Type species: Dolichopus latipennis Fallén, 1823, by subsequent designation Coquillett (1910: 554).

georgi Meuffels & Grootaert, 1999: 290 (new replacement name for minutus Statz). PA: Germany (Oligocene) [C].

minutus Statz, 1940: 136. PA: Germany (Oligocene) [C]. [Preoccupied by Fabricius, 1805.]

incertus Theobald, 1937a: 242. PA: France (Oligocene) [C].

miluus Förster, 1891: 483. PA: Germany (Oligocene) [C].

morbosus Meunier, 1907e: 221. PA: Baltic Region (Eocene) [A].

morbosus Meunier, 1907c: 198. Nomen nudum.

negotiosus Meunier, 1907e: 221 [1908d: 45]. PA: Baltic Region (Eocene) [A].

negotiosus Meunier, 1907c: 198. Nomen nudum.

nigrilamellatus Evenhuis, 1994b: 362. Nomen nudum.

paleungulatus Meuffels & Grootaert, 1999: 291 (unavailable name; new replacement name proposed for a non-existent name).

[Evenhuis (1994b: 362) listed two dolichopodid names thought to have been proposed by Loew (1850: 42) as new: Dolichopus nigrilamellatus and D. ungulatus. Further examination of Loew (1850) in the preparation of this revised version of the fossil fly catalog has shown that Loew was not proposing new species but was instead comparing unnamed amber specimens to nigrilamellatus Macquart, 1827 and ungulatus Linnaeus, 1758. Thus, there is no name ungulatus proposed by Loew, 1850 and no name needing to be replaced because of being preoccupied by ungulatus Linnaeus, 1758.]

scitus Statz, 1940: 136. PA: Germany (Oligocene) [C].

smicrus Meuffels & Grootaert, 1999: 291 (new replacement name for minutus Meunier, 1907). PA: Baltic Region (Eocene) [A].

minutus Meunier, 1907e: 221 (Hygroceleuthus). PA: Baltic Region (Eocene) [A]. [Preoccupied by Fabricius, 1805.]

minuta Meunier, 1907c: 198 (Hygroceleuthus). Nomen nudum.

spinosus Statz, 1940: 135. PA: Germany (Oligocene) [C].

titanus Meunier, 1907e: 221 (Hygroceleuthus). PA: Baltic Region (Eocene) [A].

titanus Meunier, 1907c: 198 (Hygroceleuthus). Nomen nudum.

ungulatus Evenhuis, 1994b: 362. Nomen nudum.

Unidentified sp. - NE: Canada (Oligocene) [C] (Handlirsch, 1910: 126).

Unidentified sp. - PA: Germany (Miocene) [C] (Heyden, 1858: 120).

Genus ELECTROPHORELLA Cumming & Brooks

ELECTROPHORELLA Cumming & Brooks, 2002: 42. Type species: Electrophorella baltica Cumming & Brooks, 2002, by original designation.

baltica Cumming & Brooks, 2002: 43. PA: Baltic Region (Eocene) [A].

grimaldii Cumming & Brooks, 2002: 47. PA: Baltic Region (Eocene) [A].

Genus EODOLICHOPODITES Hong

EODOLICHOPODITES Hong, 2002: 360. [Unavailable name; genus-group name based on an unavailable species-group name.]

floricopulatus Hong, 2002: 362. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

fushunensis Hong, 2002: 365. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus EOEURYOPODITES Hong

EOEURYOPODITES Hong, 2002: 348. [Unavailable name; genus-group name based on an unavailable species-group name.]

wanghuacunensis Hong, 2002: 349. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus GHEYNIA Meunier

GHEYNIA Meunier, 1899c: 322. Type species: Gheynia bifurcatus Meunier, 1907, by subsequent monotypy in Meunier (1907c: 199) (as “Gheynius bifurcatus”). 

GHEYNIUS. Incorrect subsequent spelling of Gheynia (Meunier, 1906d: 617 [1907c: 199]).

bifurcata Meunier, 1907c: 199 [1908a: 8]. PA: Baltic Region (Eocene) [A]. 

bifurcatus Meunier, 1907c: 198. Nomen nudum.

Genus GYMNOPTERNUS Loew

GYMNOPTERNUS Loew, 1857: 10. Type species: Dolichopus cupreus Fallén, 1823, by subsequent designation of Coquillett (1910: 548).

lacustris Bickel, 1997: 169. NE: USA (Colorado) (Oligocene) [C].

Genus HAODOLICHOPODITES Hong

HAODOLICHOPODITES Hong, 2002: 369. [Unavailable name; genus-group name based on an unavailable species-group name. ]

fasciatus Hong, 2002: 370. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus HERCOSTOMUS Loew

*HERCOSTOMUS Loew, 1857: 9. Type species: Sybistroma longiventris Loew, 1857, by original designation. 

Subgenus POECILOBOTHRUS Mik

*POECILOBOTHRUS Mik, 1878: 1. Type species: Dolichopus regalis Meigen, 1824, by original designation.

ciliatus Meunier, 1907e: 221 (Poecilobothrus). PA: Baltic Region (Eocene) [A].  

ciliatus Meunier, 1907c: 198. Nomen nudum.

Unplaced to Subgenus of HERCOSTOMUS Loew

bauckhorni Meunier, 1915c: 206 (Gymnopternus). PA: Germany (Oligocene) [C]. 

devinctus Meunier, 1907e: 221 (Gymnopternus). PA: Baltic Region (Eocene) [A]. 

devinctus Meunier, 1907c: 198 (Gymnopternus). Nomen nudum. 

devictus. Incorrect original spelling of devinctus (Meunier, 1907e: 221). 

diaphorus Keilbach, 1982: 377 (Gymnopternus). Nomen nudum. 

interceptus Meunier, 1907e: 221 (Gymnopternus). PA: Baltic Region (Eocene) [A]. 

interceptus Meunier, 1907c: 198 (Gymnopternus). Nomen nudum. 

intremulus Meunier, 1907e: 221 (Gymnopternus). PA: Baltic Region (Eocene) [A]. 

intremulus Meunier, 1907c: 198 (Gymnopternus). Nomen nudum. 

inumbratus Meunier, 1907e: 221 (Gymnopternus). PA: Baltic Region (Eocene) [A]. 

inumbratus Meunier, 1907c: 198 (Gymnopternus). Nomen nudum. 

quievreuxi Parent & Quiévreux, 1935: 151. PA: France (Oligocene) [C].

Genus LATICOPULUS Hong

LATICOPULUS Hong, 2002: 380. Unavailable name; genus-group name based on an unavailable species-group name.

arcidorsatus Hong, 2002: 380. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus LEPTODOLICHOPODITES Hong

LEPTODOLICHOPODITES Hong, 2002: 345. Type species Dolichopus longiflagellatus Hong, 1981, by original designation.

longiflagellatus Hong, 1981: 103 (Dolichopus). PA: China (Eocene) [A].

Genus LONGILABIA Hong

LONGILABIA Hong, 2002: 373. [Unavailable name; genus-group name based on an unavailable species-group name.]

arollaris Hong, 2002: 374. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus MEDETERA Fischer von Waldheim

*MEDETERA Fischer von Waldheim, 1819: 7. Type species: Medetera carnivora Fischer von Waldheim, 1819 [= Musca diadema Linnaeus, 1767], by monotypy. 

MEDETERUS Meigen, 1824: 59 (unjustified emendation of Medetera Fischer von Waldheim). Type species: Medetera carnivora Fischer von Waldheim, 1819 [= Musca diadema Linnaeus, 1767], automatic.

elegantula Meunier, 1907d: 209 [1907g: 245]. PA: Baltic Region (Eocene) [A]. 

elegantula Meunier, 1907c: 198. Nomen nudum

flammea Meunier, 1907d: 209 [1907g: 246]. PA: Baltic Region (Eocene) [A]. 

flammea Meunier, 1907c: 198. Nomen nudum. 

frauenfeldi Giebel, 1856: 205. PA: Baltic Region (Eocene) [A]. 

lasciva Meunier, 1907d: 209 [1907g: 246]. PA: Baltic Region (Eocene) [A]. 

lasciva Meunier, 1907c: 198. Nomen nudum. 

mustela Meunier, 1907d: 209 [1907h: 260]. PA: Baltic Region (Eocene) [A]. 

mustela Meunier, 1907c: 198. Nomen nudum. 

vana Meunier, 1907d: 209 [1907h: 260]. PA: Baltic Region (Eocene) [A]. 

vana Meunier, 1907c: 198. Nomen nudum.

Genus MICROPHOR Macquart

*MICROPHOR Macquart, 1827: 139. Type species: Microphor velutinus Macquart, 1827, by subsequent designation of Rondani (1856: 151). 

*MICROPHORUS. Incorrect subsequent spelling of Microphor (Macquart, 1834: 345).

*MICROPHORA. Incorrect subsequent spelling of Microphor (Zetterstedt, 1842: 24).

defunctus Handlirsch, 1910: 124. NE: Canada (Oligocene) [C]. 

eocenica Meunier, 1902d: 402 (Rhamphomyia). PA: Baltic Region (Eocene/Oligocene) [A]. 

rusticus Meunier, 1908i: 105 (Phyllodromia). PA: Baltic Region (Eocene/Oligocene) [A].

Genus MICROPHORITES Hennig

MICROPHORITES Hennig, 1971b: 16. Type species: Microphorites extinctus Hennig, 1971, by monotypy.

deploegi Nel, Perrichot, Daugeron & Néraudeau, 2004: 24. PA: France (Lower Cretaceous) [A].

extinctus Hennig, 1971b: 16. PA: Lebanon (Lower Cretaceous) [A].

oculeus Grimaldi & Cumming, 1999: 59. PA: Lebanon (Lower Cretaceous) [A].

similis Grimaldi & Cumming, 1999: 57. PA: Lebanon (Lower Cretaceous) [A].

utrillensis Peñalver in Arillo et al., 2008: 31. PA: Spain (Lower Cretaceous) [A].

Genus NEMATOPROCTUS Loew

*NEMATOPROCTUS Loew, 1857: 40. Type species: Porphyrops annulata Macquart, 1827 [= Chrysotus distendens Meigen, 1824], by subsequent designation of Coquillett (1910: 574).

parcus Meunier, 1907d: 210 [1908b: 21]. PA: Baltic Region (Eocene) [A]. 

parcus Meunier, 1907c: 198. Nomen nudum. 

Genus ONCOPYGIUS Mik

*ONCOPYGIUS Mik, 1866: 307. Type species: Systenus ornatus Mik, 1866 [= Sybistroma distans Loew, 1857, by monotypy.

gracilior Statz, 1940: 137. PA: Germany (Oligocene) [C]. 

oligocaenicus Statz, 1940: 137. PA: Germany (Oligocene) [C]. 

venustus Statz, 1940: 138. PA: Germany (Oligocene) [C].

Genus ORBICAPITIS Hong

ORBICAPITIS Hong, 2002: 345. Unavailable name; genus-group name based on an unavailable species-group name.

borealis Hong, 2002: 346. PA: China (Eocene) [A]. Unavailable name; species-group name proposed after 1999 without specification of a type depository.

Genus ORILABIA Hong

ORILABIA Hong, 2002: 354. Type species Septocellula fera Hong, 1981, by original designation.

fera Hong, 1981: 113 (Septocellula). PA: China (Eocene) [A]. 

ferina. Incorrect original spelling of fera (Hong, 1981: 6). 

Genus PALAEOARGYRA Meunier

PALAEOARGYRA Meunier, 1895b: 5. Type species: Palaeoargyra dytei Evenhuis, 1993, by subsequent designation of Evenhuis (1994b: 365). 

[Palaeoargyra originally proposed without included species. The type designated is the first inclusion of a named species within the genus in accordance with the Code.]

dytei Evenhuis, 1994b: 365. PA: Baltic Region (Eocene) [A].

[Species-group name available by bibliographic reference to characters given in Meunier (1895b: 5) for the genus Palaeoargyra.]

Genus PALAEOMEDETERUS Meunier

PALAEOMEDETERUS Meunier, 1895c: 174. Type species: Tipula culiciforme Meunier, 1899, by subsequent monotypy in Meunier (1899e: 174). 

PALAEOCHRYSOTUS Meunier, 1906d: 617 [1906e: 267]. Nomen nudum. 

PALAEOCHRYSOTUS Meunier, 1907c: 199 [1907i : 270]. Type species: Palaeochrysotus horridus Meunier, 1907, by subsequent designation of Spahr (1985: 36). [Preoccupied by Linnaeus, 1758.]. 

fessus Meunier, 1907d: 210 [1908a: 7] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

fessus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

hirsutus Meunier, 1907d: 210 [1907i: 270] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

hirsutus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

horridus Meunier, 1907d: 210 [1907i: 270] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

horridus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

ignavus Meunier, 1907d: 210 [1907j: 281] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

ignavus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

languidus Meunier, 1907d: 210 [1907j: 282] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

languidus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

lassatus Meunier, 1907d: 210 [1908a: 8] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

lassatus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum. 

lentus Meunier, 1907d: 210 [1907j: 281] (Palaeochrysotus). PA: Baltic Region (Eocene) [A]. 

lentus Meunier, 1907c: 198 (Palaeochrysotus). Nomen nudum.

meunieri Meuffels & Grootaert, 1999: 291 (new replacement name for culiciformis Meunier). PA: Baltic Region (Eocene) [A].

culiciformis Meunier, 1899e: 174 (Tipula). PA: Baltic Region (Eocene) [A].

Genus PARADOLICHOPODITES Hong

PARADOLICHOPODITES Hong, 2002: 382. [Unavailable name; genus-group name based on an unavailable species-group name.]

xilutianensis Hong, 2002: 383. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus PROCHRYSOTUS Meunier

PROCHRYSOTUS Meunier, 1906d: 617. Nomen nudum. 

PROCHRYSOTUS Meunier, 1906e: 267. Nomen nudum. 

PROCHRYSOTUS Meunier, 1907c: 199. Type species: Prochrysotus magnus Meunier, 1907, by monotypy.

magnus Meunier, 1907c: 198 [1907d: 210, 1907i: 269]. PA: Baltic Region (Eocene) [A].

Genus PROHERCOSTOMUS Grichanov

PROHERCOSTOMUS Grichanov, 1997: 83 (520) (as Hercostomus subgenus). Type species: Hercostomus noxialis Meunier, 1907, by original designation.

bickeli Evenhuis, 1994b: 361 (new replacement name for vulgaris Meunier) (Dolichopus). PA: Baltic Region (Eocene) [A].

vulgaris Meunier, 1907c: 198 (Dolichopus). Nomen nudum. [Preoccupied by Stannius, 1831.].

vulgaris Meunier, 1907e: 221 [1908d: 45] (Dolichopus). PA: Baltic Region (Eocene) [A]. [Preoccupied by Stannius, 1831.].

meunierianus Evenhuis, 1994b: 361 (Dolichopus) (new replacement name for notabilis Meunier). PA: Baltic Region (Eocene) [A].

monotonus Meunier, 1907e: 221 (Dolichopus). PA: Baltic Region (Eocene) [A].

monotonus Meunier, 1907c: 198 (Dolichopus). Nomen nudum.

notabilis Meunier, 1907e: 221 [1908d: 45] (Dolichopus). PA: Baltic Region (Eocene) [A]. [Preoccupied by Zetterstedt, 1843.].

notabilis Meunier, 1907c: 198 (Dolichopus). Nomen nudum.

noxialis Meunier, 1907e: 221 [1908d: 46]. PA: Baltic Region (Eocene) [A].

noxialis Meunier, 1907c: 198. Nomen nudum.

Genus PROSYSTENUS Negrobov

PROSYSTENUS Negrobov, 1976: 122 (1977: 495). Type species: Prosystenus zherichini Negrobov, 1976, by original designation.

zherichini Negrobov, 1976: 122 (1977: 496). PA: Russia (Sakhalin I) (Paleocene/Eocene) [A].

Genus RETINITUS Negrobov

RETINITUS Negrobov, 1978: 86 (1979: 225). Type species: Retinitus nervosus Negrobov, 1978, by original designation.

nervosus Negrobov, 1978: 87 (1979: 226). PA: Russia (Siberia) (Upper Cretaceous) [A].

Genus RHAPHIUM Meigen

*RHAPHIUM Meigen, 1803: 272. Type species: Rhaphium macrocerum Meigen, 1824, by subsequent designation of Curtis (1835: pl. 568). 

*XIPHANDRIUM Loew, 1857: 36. Type species: Rhaphium quadrifilatum Loew, 1857 (as “quadrifilum”) [= Rhaphium ensicorne Meigen, 1824], by subsequent designation of Coquillett (1910: 621).

splendidum Meunier, 1907d: 209 [1907f: 233] (Xiphandrium). PA: Baltic Region (Eocene) [A]. 

splendidum Meunier, 1907c: 198. Nomen nudum. 

Unidentified sp. - PA: UK (England) (Pliocene) [A] (Reid, 1886: 248).

Genus SCIAPUS Zeller

*PSILOPUS Meigen, 1824: 35. Type species: Dolichopus platypterus Fabricius, 1805, by subsequent designation of Westwood (1840: 134). [Preoccupied by Poli, 1795.] 

*SCIAPUS Zeller, 1842: 831 (new replacement name for Psilopus Meigen). Type species: Dolichopus platypterus Fabricius, 1805, automatic.

combaluzieri Timon-David, 1944: 129. PA: France (Oligocene) [C]. 

erasmius Meuffels & Grootaert, 1999: 290 (new replacement name for amabilis Statz). PA: Germany (Oligocene) [C].

amabilis Statz, 1940: 139. PA: Germany (Oligocene) [C]. [Preoccupied by Parent, 1929.] 

rottensis Statz, 1940: 139. PA: Germany (Oligocene) [C].

Genus SEPTOCELLULA Hong

SEPTOCELLULA Hong, 1981: 106. Type species: Septocellula asiatica Hong, 1981, by original designation.

asiatica Hong, 1981: 109. PA: China (Eocene) [A]. 

Genus SINODOLICHOPODITES Hong

SINODOLICHOPODITES Hong, 2002: 377. [Unavailable name; genus-group name based on an unavailable species-group name.]

liaoningensis Hong, 2002: 377. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus SUNODOLICHOPODITES Hong

SUNODOLICHOPODITES Hong, 2002: 385. [Unavailable name; genus-group name based on an unavailable species-group name.]

longus Hong, 2002: 386. PA: China (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specification of a type depository.]

Genus SYMPYCNITES Grimaldi & Cumming

SYMPYCNITES Grimaldi & Cumming, 1999: 67. Type species Sympycnites primaevus Grimaldi & Cumming, 1999, by original designation.

primaevus Grimaldi & Cumming, 1999: 67. PA: Lebanon (Lower Cretaceous) [A].

Genus SYSTENUS Loew

*SYSTENUS Loew, 1857: 34. Type species: Rhaphium adpropinquans Loew, 1857 [= Rhaphium pallipes Roser, 1840], by subsequent designation of Foote et al. (1965: 517).

ciliatus Meunier, 1907e: 222 [1907g: 245]. PA: Baltic Region (Eocene) [A]. 

ciliatus Meunier, 1907c: 198. Nomen nudum.

Genus THINOPHILUS Wahlberg

*THINOPHILUS Wahlberg, 1844: 37. Type species: Rhaphium flavipalpe Zetterstedt, 1843, by monotypy.

piraticus Meunier, 1907e: 222 [1908f: 58]. PA: Baltic Region (Eocene) [A]. 

piraticus Meunier, 1907c: 198. Nomen nudum.

Genus THRYPTICUS Gerstaecker

*THRYPTICUS Gerstaecker, 1864: 43. Type species: Thrypticus smaragdinus Gerstaecker, 1864, by monotypy.

gestuosus Meunier, 1907d: 209 [1907h: 262]. PA: Baltic Region (Eocene) [A]. 

gestuosus Meunier, 1907c: 198. Nomen nudum. 

gulosus Meunier, 1907d: 209 [1907h: 262]. PA: Baltic Region (Eocene) [A]. 

gulosus Meunier, 1907c: 198. Nomen nudum. 

molestus Meunier, 1907d: 209 [1907h: 261]. PA: Baltic Region (Eocene) [A]. 

molestus Meunier, 1907c: 198. Nomen nudum. 

sobrius Meunier, 1907d: 209 [1907h: 261]. PA: Baltic Region (Eocene) [A]. 

sobrius Meunier, 1907c: 198. Nomen nudum.

Genus WANGIA Hong

WANGIA Hong, 2002: 354. Type species: Septocellula trichopoda Hong, 1981, by original designation. [Preoccupied by Fowler, 1954.]

trichopoda Hong, 1981: 116 (Septocellula). PA: China (Eocene) [A].

Genus WHEELERENOMYIA Meunier

WHEELERENOMYIA Meunier, 1907c: 199. Type species: Wheelerenomyia eocenica Meunier, 1907, by monotypy.

bickeli Negrobov & Selivanova, 2003: 61 (Amesorhaga). PA: Baltic Region (Eocene) [A].

corcula Meunier, 1907e: 222 (Neurigona). PA: Baltic Region (Eocene) [A]. 

corcula Meunier, 1907c: 198 (Saucropus). Nomen nudum.

eocenica Meunier, 1907c: 198 [1907e: 222, 1908f: 58]. PA: Baltic Region (Eocene) [A].

longicerca Negrobov & Selivanova, 2003: 63 (Amesorhaga). PA: Baltic Region (Eocene) [A].

originarius Meunier, 1907d: 210 [1908a: 8] (Nematoproctus). PA: Baltic Region (Eocene) [A]. 

originarius Meunier, 1907c: 198 (Nematoproctus). Nomen nudum. 

pacatus Meunier, 1907d: 210 [1908b: 21] (Nematoproctus). PA: Baltic Region (Eocene) [A]. 

pacatus Meunier, 1907c: 198 (Nematoproctus). Nomen nudum. 

parvulus Meunier, 1907d: 210 [1908b: 22] (Nematoproctus). PA: Baltic Region (Eocene) [A]. 

parvulus Meunier, 1907c: 198 (Nematoproctus). Nomen nudum. 

parvus Meunier, 1907d: 210 [1908b: 21] (Nematoproctus). PA: Baltic Region (Eocene) [A]. 

parvus Meunier, 1907c: 198(Nematoproctus). Nomen nudum. 

pellucidus Meunier, 1907d: 210 [1908b: 22] (Psilopus). PA: Baltic Region (Eocene) [A]. 

pellucidus Meunier, 1907c: 198 (Psilopus). Nomen nudum. 

peratticus Meunier, 1907d: 210 [1908b: 22] (Psilopus). PA: Baltic Region (Eocene) [A]. 

peratticus Meunier, 1907c: 198 (Psilopus). Nomen nudum. 

persastutulus Meunier, 1907d: 210 [1908b: 22] (Psilopus). PA: Baltic Region (Eocene) [A].

persastutulus Meunier, 1907c: 198 (Psilopus). Nomen nudum. 

subparvus Meunier, 1916a: 278 (Nematoproctus). PA: Baltic Region (Eocene) [A].

quadrispinosa Negrobov & Selivanova, 2003: 60 (Amesorhaga). PA: Baltic Region (Eocene) [A].

vladimiri Negrobov & Selivanova, 2003: 58 (Amesorhaga). PA: Baltic Region (Eocene) [A].

Genus XANTHOCHLORUS Loew

*XANTHOCHLORUS Loew, 1857: 42. Type species: Medeterus ornatus Haliday, 1832, by subsequent designation of Coquillett (1910: 620).

tenellus Statz, 1940: 138. PA: Germany (Oligocene) [C]. [Preoccupied by Wiedemann,, 1828; no new name proposed here.]

Unplaced Species of DOLICHOPODIDAE

pusilla Meunier, 1904c: 73 [1904d: 62] (Sciara). PA: Baltic Region (Eocene) [A]. [Preoccupied by Meigen, 1818; no new replacement name proposed here.]