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Family GLOSSINIDAE
Members of the calypterate family Glossinidae are the fabled
"tsetse"** of Africa. Adults of extant forms are principal vectors of
sleeping sickness. Females are larviparous. After the
larva is laid on the ground, it immediately pupates.
The current distribution of the family is restricted to the
African continent, though the fossil record of these flies is known only from
the Oligocene of the North American continent. Many workers have postulated the
fly's possible hosts in the geologic past and various reasons for such a
disjunct distribution. One of the more colorful of these was related by Cole
& Schlinger (1969) who stated "A very rough estimate has been given
concerning these ancient glossinid flies, which were
said to have subsisted on dinosaur blood for a hundred million years. When
their big bloodbanks vanished, they turned to the
nearest available substitute, the alligator."
Lambrecht (1980) has more recently hypothesized
the geological and evolutionary history of the family. Grimaldi (1992) is the
most current worker to review the fossil record and revised the systematics of
the fossils and discussed the cladistic biogeography and paleobiogeography of
the group.
Refs.:
Cockerell (1918, review of fossil taxa); Lambrecht (1980,
paleoecology of Glossina); Grimaldi (1992, review of fossil taxa).
Genus GLOSSINA
Wiedemann
*GLOSSINA
Wiedemann, 1830: 253. Type species: Glossina longipalpis Wiedemann,
1830, by monotypy.
PALOESTRUS
Scudder, 1892: 18. Type species: Paloestrus
oligocenus Scudder, 1892, by monotypy.
COCKERELLITHA
Townsend, 1938: 166. Type species: Glossina osborni Cockerell, 1909, by
original designation.
LITHOGLOSSINA
Townsend, 1938: 166. Type species: Glossina armatipes Cockerell, 1917 [=
Glossina osborni Cockerell, 1909], by original designation.
oligocenus Scudder, 1892: 19 (Paloestrus).
NE: USA (Oligocene) [C].
veterna Cockerell, 1916b: 70. NE: USA
(Oligocene) [C].
osborni
Cockerell, 1909e: 128. NE: USA (Oligocene) [C].
armatipes
Cockerell, 1917a: 19. NE: USA (Oligocene) [C].
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** The word "tsetse" is from
the Tswana language of southern Africa and means, literally, "fly".
This means that the many scientists and others who have commonly called these
flies "tsetse flies" are really calling them "fly flies".