Family LIMONIIDAE

 

Limoniids share many characteristics with tipulids, but are normally smaller in size, though exceptions occur. The family is found worldwide with approximately 11,000 species having been described. In extant forms, immatures are found in aquatic or semiaquatic environments where they feed on decaying plant matter, fungi, and algae. Adult males of certain species may swarm.
The separation of Limoniidae, Cylindrotomidae, and Tipulidae is retained in this catalog following the treatment of these families by contemporary European workers.
A possible limoniid has been recorded from the Lower Cretaceous amber of Lebanon by Whalley (1981) and undetermined material has been recorded from the Late Cretaceous amber of France by Schlüter (1978: 93) and the Cretaceous amber of Siberia (Zherichin & Sukacheva, 1973: 20; Zherichin, 1978: 82). Other undetermined records include material from the Miocene of Sado Island Japan (Fujiyama, 1985: 37), a possible new genus close to Pseudolimnophila Alexander and Eutonia Wulp from the Paleocene of Alberta, Canada (Wighton, 1982: 578), and two specimens probably representing new genera from the Lower Cretaceous deposits of the Koonwarra Fossil Bed, Australia (Jell & Duncan, 1986: 169).

Ref.: Meunier (1906f, revision of amber taxa); Scudder (1894, review of North American fossils); Alexander (1931, review of Baltic amber taxa).

 

Genus ADELPHOMYIA Bergroth

*ADELPHOMYIA Bergroth, 1891: 134. Type species: Adelphomyia helvetica Bergroth, 1891 [= Limnobia punctum Meigen, 1818], by monotypy.

disjunctula Cockerell & Haines, 1921a: 83. PA: UK (England) (Eocene/Oligocene) [C].

 

Genus ALLARITHMIA Loew

ALLARITHMIA Loew, 1850b: 36. Type species: Allarithmia palpata Loew, 1850, by monotypy.

palpata Loew, 1850b: 38. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus ANTOCHA Osten Sacken

*TAPHROPHILA Rondani, 1856: 185. Type species: Limnobia inusta Meigen, 1818 [misidentification], by original designation.
*ANTOCHA Osten Sacken, 1860: 219. Type species: Antocha saxicola Osten Sacken, 1860, by subsequent designation of Coquillett (1910: 508).
[In the interest of stability and in accordance with usage, the name Antocha is here retained for species in this genus, whereas Hutson & Vane-Wright (1969) recommend to consider Taphrophila as senior synonym of Antocha. The original type designation of Taphrophila is based on a misidentification while the description of Taphrophila refers to characters typical of Antocha. In this case, a subsequent type-designation is required. Acceptance of the original (misidentified) type species of Taphrophila cannot be recommended because it would result in Taphrophila becoming a junior synonym of Limonia and a senior synonym of Atypophthalmus.]

principialis Scudder, 1894: 208 (46). NE: USA (Oligocene) [C].

 

Genus ARCHIMESOTIPULA Lin

ARCHIMESOTIPULA Lin, 1986: 87. Type species: Archimesotipula antefortis Lin, 1986, by original designation.
ARCHIMOSOTIPULA. Incorrect original spelling of Archimesotipula (Lin, 1986: 87).

antefortis Lin, 1986: 88. PA: China (Lower Jurassic) [C].
antifortis. Incorrect original spelling of antefortis (Lin, 1986: 111).

 

Genus ARCHITIPULA Handlirsch

ARCHITIPULA Handlirsch, 1906a: 490. Type species: Architipula seebachiana Handlirsch, 1906, by subsequent designation of Rohdendorf (1962: 313).
PROTIPULA Handlirsch, 1906a: 491. Type species: Protipula crassa Handlirsch, 1906, by monotypy.
PARATIPULA Bode, 1953: 305. Type species: Paratipula acuta Bode, 1953, by original designation.
EOASILIDEA Bode, 1953: 315. Type species: Eoasilidea fragmentosa Bode, 1953, by original designation.

Subgenus ARCHITIPULA Handlirsch

seebachiana Handlirsch, 1906a: 490. PA: Germany (Lower Jurassic) [C].

Subgenus GRIMMENIA Krzeminski & Zessin

GRIMMENIA Krzeminski & Zessin, 1990: 41. Type species: Architipula prima Krzeminski & Zessin, 1990, by original designation.

prima Krzeminski & Zessin, 1990: 41. PA: Germany (Lower Jurassic) [C].
secunda Krzeminski & Zessin, 1990: 42. PA: Germany (Lower Jurassic) [C].

Unplaced to Subgenus of ARCHITIPULA Handlirsch

acuta Bode, 1953: 305 (Paratipula). PA: Germany (Lower Jurassic) [C].
aequabilis Bode, 1953: 309 (Paratipula). PA: Germany (Lower Jurassic) [C].
analiramosa Bode, 1953: 306 (Paratipula). PA: Germany (Lower Jurassic) [C].
arculifera Bode, 1905: 244. (Phryganidium). PA: Germany (Lower Jurassic) [C].
areolata Handlirsch, 1939: 110. PA: Germany (Lower Jurassic) [C].
asiatica Rohdendorf, 1964: 147 (1974: 156). PA: Kirghizistan (Lower Jurassic) [C].
austeni Jarzembowski, 1991: 100. PA: UK (England) (Lower Cretaceous) [C].
basiminuta Bode, 1953: 308 (Paratipula). PA: Germany (Lower Jurassic) [C].
bedeisimilis Bode, 1953: 304. PA: Germany (Lower Jurassic) [C].
bodei Handlirsch, 1939: 112. PA: Germany (Lower Jurassic) [C].
[Name validated by bibliographic reference to Bode (1905: pl. 6, fig. 21.]
brunsvicensis Handlirsch, 1939: 112. PA: Germany (Lower Jurassic) [C].
[Name validated by bibliographic reference to Bode (1905: pl. 6, fig. 22.]
clara Handlirsch, 1939: 111. PA: Germany (Lower Jurassic) [C].
conspicua Handlirsch, 1939: 111. PA: Germany (Lower Jurassic) [C].
crassa Handlirsch, 1906a: 491 (Protipula). PA: Germany (Lower Jurassic) [C].
debilis Handlirsch, 1939: 109. PA: Germany (Lower Jurassic) [C].
dubia Handlirsch, 1939: 111. PA: Germany (Lower Jurassic) [C].
elegans Handlirsch, 1906a: 490. PA: Germany (Lower Jurassic) [C].
formosa Bode, 1953: 307 (Paratipula). PA: Germany (Lower Jurassic) [C].
fragmentosa Bode, 1953: 316 (Eoasilidea). PA: Germany (Lower Jurassic) [C].
geinitzi Handlirsch, 1939: 109. PA: Germany (Lower Jurassic) [C].
minuta Handlirsch, 1939: 108. PA: Germany (Lower Jurassic) [C].
intermedia Handlirsch, 1939: 110. PA: Germany (Lower Jurassic) [C].
lapidaria Handlirsch, 1906a: 492. PA: Germany (Lower Jurassic) [C].
lataelata Bode, 1953: 310 (Paratipula). PA: Germany (Lower Jurassic) [C].
latipennis Handlirsch, 1906a: 491. PA: Germany (Lower Jurassic) [C].
longipes Rohdendorf, 1964: 207 (1974: 232). PA: Kazakhstan (Upper Jurassic) [C].
maior Handlirsch, 1939: 110. PA: Germany (Lower Jurassic) [C].
nana Handlirsch, 1939: 108. PA: Germany (Lower Jurassic) [C].
obliqua Handlirsch, 1939: 110. PA: Germany (Lower Jurassic) [C].
orientalis Kalugina in Kalugina & Kovalev, 1985: 60. PA: Russia (Siberia) (Middle Jurassic) [C].
parva Handlirsch, 1939: 108. PA: Germany (Lower Jurassic) [C].
protipuloides Rohdendorf, 1964: 207 (1974: 231). PA: Kazakhstan (Upper Jurassic) [C].
ptychopteraeformis Bode, 1953: 309 (Paratipula). PA: Germany (Lower Jurassic) [C].
pulla Handlirsch, 1939: 109. PA: Germany (Lower Jurassic) [C].
pusilla Handlirsch, 1939: 107. PA: Germany (Lower Jurassic) [C].
radiata Rohdendorf, 1964: 147 (1974: 155). PA: Kirghizistan (Lower Jurassic) [C].
radiata Rohdendorf, 1962: 312 (1991: 453). Nomen nudum.
robusta Bode, 1953: 308 (Paratipula). PA: Germany (Lower Jurassic) [C].
seebachi Geinitz, 1884: 576 (Phryganidium). PA: Germany (Lower Jurassic) [C].
simplex Handlirsch, 1939: 109. PA: Germany (Lower Jurassic) [C].
stigmatica Handlirsch, 1906a: 491. PA: Germany (Lower Jurassic) [C].
symmetrica Kalugina in Kalugina & Kovalev, 1985: 61. PA: Russia (Siberia) (Middle Jurassic) [C].
turanica Rohdendorf, 1964: 147 (1974: 155). PA: Kirghizistan (Lower Jurassic) [C].
veris Bode, 1953: 307 (Paratipula). PA: Germany (Lower Jurassic) [C].
youngi Krzeminski, 1992c: 43. NE: USA (Upper Triassic) [C].

 

Genus ATARBA Osten Sacken

*ATARBA Osten Sacken, 1869: 127. Type species: Atarba picticornis Osten Sacken, 1869, by monotypy.

vectensis Cockerell, 1915c: 492. PA: UK (England) (Eocene/Oligocene) [C].

 

Genus AUSTROLIMNOPHILA Alexander

AUSTROLIMNOPHILA Alexander, 1920c: 4 (as Limnophila subgenus). Type species: Limnophila eutaeniata Bigot, 1888, by original designation.

Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzemio(n,´)ski, 1992a: 107).

 

Genus CALOBAMON Loew

CALOBAMON Loew, 1850b: 36, 38. Type species: Calobamon longipes Keilbach, 1982, by subsequent monotypy in Keilbach (1982: 326).

longipes Keilbach, 1982: 326. PA: Baltic Region (Eocene/Oligocene) [A].
[Species name valid by bibliographic reference to characters in Loew (1850b: 36 for the genus Calobamon).]

 

Genus CHEILOTRICHIA Rossi

*CHEILOTRICHIA Rossi, 1848: 12. Type species: Erioptera imbuta Meigen, 1818, by monotypy.

Subgenus EMPEDA Osten Sacken

*EMPEDA Osten Sacken, 1869: 183 (as genus). Type species: Empeda stigmatica Osten Sacken, 1869, by subsequent designation of Coquillett (1910: 537).

axillaris Alexander, 1931: 110 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
diacantha Alexander, 1931: 109 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].
minuta Meunier, 1899e: 173 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
prolifica Meunier, 1906f: 375 (Empeda). PA: Baltic Region (Eocene/Oligocene) [A].
elongata Meunier, 1906f: 376 (Empeda). PA: Baltic Region (Eocene/Oligocene) [A].
platyphylla Alexander, 1931: 107 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].
rectistyla Alexander, 1931: 111 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].
schummeli Meunier, 1917a: 98 (Empeda). PA: Baltic Region (Eocene/Oligocene) [A].
subabortiva Alexander, 1931: 110. (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of CHEILOTRICHIA Rossi

duplicata Alexander, 1931: 108 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].
ferruginea Cockerell, 1921c: 457 (Empeda). PA: UK (England) (Eocene/Oligocene) [C].
hyalina Cockerell, 1921c: 457 (Empeda). PA: UK (England) (Eocene/Oligocene) [C].

 

Genus CLADONEURA Scudder

CLADONEURA Scudder, 1894: 212 (50). Type species: Cladoneura willistoni Scudder, 1894, by monotypy.

willistoni Scudder, 1894: 213 (51). NE: USA (Oligocene) [C].

 

Genus CLADURA Osten Sacken

*CLADURA Osten Sacken, 1860: 229. Type species: Cladura flavoferruginea Osten Sacken, 1860, by monotypy.

integra Scudder, 1894: 216 (54). NE: USA (Oligocene) [C].
maculata Scudder, 1894: 215 (53). NE: USA (Oligocene) [C].

 

Genus CRETOLIMONIA Kalugina

CRETOLIMONIA Kalugina, 1986: 115. Type species: Cretolimonia popovi Kalugina, 1986, by original designation.

popovi Kalugina, 1986: 115. PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C].

 

Genus CRITONEURA Loew

CRITONEURA Loew, 1850b: 36. Type species: Limnophila pentagonalis Meunier, 1906, by subsequent monotypy in Meunier (1906f: 361).
[Critoneura originally proposed without validly diagnosed species. The first included species was pentagonalis Meunier (1906f: 361 [footnote]) who validated the name by giving characters to differentiate it.]

longipes Loew, 1850b: 38. Nomen nudum.
pentagonalis Meunier, 1906f: 361 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].
pentagonalis Loew, 1850b: 38. Nomen nudum.

 

Genus CYTTAROMYELLA Meunier

CYTTAROMYELLA Meunier, 1915d: 229. Type species: Cyttaromyella bastini Meunier, 1915, by monotypy.

bastini Meunier, 1915d: 229. PA: Germany (Oligocene) [C].

 

Genus DACTYLOLABIS Osten Sacken

*DACTYLOLABIS Osten Sacken, 1860: 240 (as Limnophila subgenus). Type species: Limnophila montana Osten Sacken, 1860, by monotypy.

Subgenus IDIOLABIS Alexander

IDIOLABIS Alexander, 1931: 59. Type species: Dactylolabis terebrella Alexander, 1931, by original designation.

terebrella Alexander, 1931: 59. PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of DACTYLOLABIS Osten Sacken

tenuis Statz, 1944b: 99. PA: Germany (Oligocene) [C].
vetusta Alexander, 1931: 60. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus DICRANOMYIA Stephens

*DICRANOMYIA Stephens, 1829a: 53. Type species: Limnobia modesta Meigen, 1818, by subsequent designation of Coquillett (1910: 533).

antennifera Théobald, 1937a: 344. PA: France (Oligocene) [C].
excavata Cockerell & Haines, 1921a: 83 [1921b: 110]. PA: UK (England) (Eocene/Oligocene) [C].
exhumata Cockerell, 1922: 17 (new replacement name for excavata Cockerell). PA: UK (England) (Eocene/Oligocene) [C].
excavata Cockerell, 1921c: 458. PA: UK (England) (Eocene/Oligocene) [C]. [Preoccupied by Cockerell & Haines, 1921.]
faecarius Scudder, 1894: 206 (44) (Rhamphidia). NE: USA (Oligocene) [C].
fontainei Scudder, 1894: 200 (38). NE: USA (Oligocene) [C].
fragilis Scudder, 1894: 199 (37). NE: USA (Oligocene) [C].
graciosa Meunier, 1916b: 489. PA: Baltic Region (Eocene/Oligocene) [A].
inferna Scudder, 1894: 198 (36). NE: USA (Oligocene) [C].
lobata Meunier, 1906f: 363. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
loewi Scudder, 1894: 207 (45) (Rhamphidia). NE: USA (Oligocene) [C].
longipes Scudder, 1894: 197 (35). NE: USA (Oligocene) [C].
perpendicularis Savtschenko, 1967: 469 (Limonia). PA: Baltic Region (Eocene/ Oligocene) [A].
primitiva Scudder, 1877a: 748. NE: USA (Eocene) [C].
rhodolitha Cockerell, 1908b: 228. NE: USA (Eocene) [C].
rohweri Brown, 1985: 99 (Limonia). NE: USA (Oligocene) [C].
rostrata Scudder, 1877a: 749. NE: USA (Eocene) [C].
decrepita Scudder, 1877a: 752 (Tipula). NE: USA (Eocene) [C].
saxetana Scudder, 1894: 206 (44) (Rhamphidia). NE: USA (Oligocene) [C].
sergio Krzeminski & Gentilini, 1992: 93. PA: Italy (Miocene) [C].
sinuata Meunier, 1916b: 490. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
stagnorum Scudder, 1894: 197 (35). NE: USA (Oligocene) [C].
stigmosa Scudder, 1877a: 746. NE: USA (Eocene) [C].
tecta Scudder, 1877a: 752 (Tipula). NE: USA (Eocene) [C].
thybotica Henriksen, 1922: 21 (Rhamphidia). PA: Denmark (Eocene) [C].
undulata Cockerell & Haines, 1921a: 83 [1921b: 110]. PA: UK (England) (Eocene/Oligocene) [C].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzemio(n,´)ski, 1992a: 107).

 

Genus DICRANOPTYCHA Osten Sacken

*DICRANOPTYCHA Osten Sacken, 1860: 217. Type species: Dicranoptycha germana Osten Sacken, 1860, by subsequent designation of Coquillett (1910: 533).
STIBADOCERITES Zeuner, 1941: 96. Type species: Stibadocerites europaeus Zeuner, 1941, by monotypy.

anna Krzeminski & Gentilini, 1992: 91. PA: Italy (Miocene) [C].
electrina Alexander, 1931: 40. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
europaea Zeuner, 1941: 96 (Stibadocerites). PA: UK (Scotland) (Upper Cretaceous) [C].
lignitica Statz, 1934: 99. PA: Germany (Oligocene) [C].
rottensis Statz, 1944b: 95. PA: Germany (Oligocene) [C].

 

Genus DICRANOTA Zetterstedt

*DICRANOTA Zetterstedt, 1838: 851. Type species: Dicranota guerini Zetterstedt, 1838, by monotypy.

cockerelli Brown, 1985: 99. NE: USA (Oligocene) [C].

 

Genus DICTYOTIPULA Rohdendorf

DICTYOTIPULA Rohdendorf, 1962: 312 (1991: 452). Type species: Dictyotipula densa Rohdendorf, 1962, by original designation.

densa Rohdendorf, 1962: 312 (1991: 452). PA: Kirghizistan (Lower Jurassic) [C].

 

Genus DIPLARCHITIPULA Rohdendorf

DIPLARCHITIPULA Rohdendorf, 1962: 313 (1991: 452). Type species: Diplarchitipula multimedialis Rohdendorf, 1962, by original designation.

destructa Rohdendorf, 1964: 144 (1974: 154). PA: Kirghizistan (Lower Jurassic) [C].
multimedialis Rohdendorf, 1962: 313 (1991: 452). PA: Kirghizistan (Lower Jurassic) [C].

 

Genus DIPTERODICTYA Rohdendorf

DIPTERODICTYA Rohdendorf, 1961: 92. Type species: Dipterodictya tipuloides Rohdendorf, 1961, by original designation.

tipuloides Rohdendorf, 1961: 92. PA: Kirghizistan (Lower Jurassic) [C].

 

Genus ELEPHANTOMYIA Osten Sacken

*ELEPHANTOMYIA Osten Sacken, 1860: 220. Type species: Limnobiorhynchus canadensis Westwood, 1835 [misidentification, = Elephantomyia westwoodi Osten Sacken, 1869], by original designation.

baltica Alexander, 1931: 58. PA: Baltic Region (Eocene/Oligocene) [A].
brevipalpa Loew, 1851: 400 (Toxorhina). PA: Baltic Region (Eocene/Oligocene) [A].
brevipalpa Loew, 1850b: 37 (Toxorhina). Nomen nudum.
longirostris Loew, 1851: 400 (Toxorhina). PA: Baltic Region (Eocene/Oligocene) [A].
longirostris Loew, 1850b: 37 (Toxorhina). Nomen nudum.
pulchella Loew, 1851: 400 (Toxorhina). PA: Baltic Region (Eocene/Oligocene) [A].
pulchella Loew, 1850b: 37 (Toxorhina). Nomen nudum.
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzemio(n,´)ski, 1992a: 107).

 

Genus EOTIPULA Handlirsch

EOTIPULA Handlirsch, 1906a: 491. Type species: Eotipula parva Handlirsch, 1906, by subsequent designation of Carpenter (1992: 402).

coarctata Handlirsch, 1939: 113. PA: Germany (Lower Jurassic) [C].
defuncta Handlirsch, 1939: 112. PA: Germany (Lower Jurassic) [C].
lapidaria Handlirsch, 1906a: 492. PA: Germany (Lower Jurassic) [C].
longa Handlirsch, 1939: 112. PA: Germany (Lower Jurassic) [C].
mortua Handlirsch, 1939: 112. PA: Germany (Lower Jurassic) [C].
parva Handlirsch, 1906a: 491. PA: Germany (Lower Jurassic) [C].

 

Genus EOTIPULINA Kalugina

EOTIPULINA Kalugina in Kalugina & Kovalev, 1985: 52. Type species: Eotipulina sibirica Kalugina, 1985, by original designation.

complanata Kalugina in Kalugina & Kovalev, 1985: 54. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
fumosa Kalugina in Kalugina & Kovalev, 1985: 54. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
sibirica Kalugina in Kalugina & Kovalev, 1985: 52. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].
?Unidentified sp.-PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C] (Kalugina, 1986: 116).

 

Genus EOTIPULOPTERA Kalugina

EOTIPULOPTERA Kalugina in Kalugina & Kovalev, 1985: 55. Type species: Eotipuloptera savtshenkoi Kalugina, 1985, by original designation.

savtshenkoi Kalugina in Kalugina & Kovalev, 1985: 55. PA: Russia (Siberia) (Middle/Upper Jurassic) [C].

 

Genus EPIPHRAGMA Osten Sacken

*EPIPHRAGMA Osten Sacken, 1860: 238 (as Limnophila subgenus). Type species: Limnophila pavonina Osten Sacken, 1860 [= Limnobia fasciapennis Say, 1823], by subsequent designation of Coquillett (1910: 539).

spilopterum Cockerell, 1921c: 458. PA: UK (England) (Eocene/Oligocene) [C].

 

Genus ERIOPTERA Meigen

*ERIOPTERA Meigen, 1803: 262. Type species: Erioptera lutea Meigen, 1804, by subsequent designation of Coquillett (1910: 540).
[In the interests of stability and in accordance to usage, prior designations of type species by Blanchard in Audouin et al. (1848: pl. 163) and Rondani (1856) that would render Erioptera a senior synonym of Molophilus Curtis are set aside here. Suspension of I.C.Z.N. rules is required to validate the subsequent type-designation by Coquillett (1910).]

abbreviata Keilbach, 1982: 325. Nomen nudum.
anomala Keilbach, 1982: 325. Nomen nudum.
arcuata Alexander, 1931: 105. PA: Baltic Region (Eocene/Oligocene) [A].
berendtii Keilbach, 1982: 325. Nomen nudum.
danae Heyden, 1870: 252. PA: Germany (Oligocene) [C].
fasciipennis Statz, 1944b: 97. PA: Germany (Oligocene) [C].
gracilis Meunier, 1899e: 173. PA: Baltic Region (Eocene/Oligocene) [A].
hybrida Keilbach, 1982: 325. Nomen nudum.
oligocaenica Meunier, 1922a: 509 [1923c: 610]. PA: Germany (Oligocene) [C].
pagasti Statz, 1944b: 97. PA: Germany (Oligocene) [C].
perspicillata Meunier, 1922a: 509 [1923c: 610]. PA: Germany (Oligocene) [C].
socia Keilbach, 1982: 325. Nomen nudum.

 

Genus GERANOMYIA Haliday

*GERANOMYIA Haliday, 1833: 154. Type species: Geranomyia unicolor Haliday, 1833, by monotypy.

messiniana Gentilini, 1986: 181. PA: Italy (Miocene) [C].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).
Unidentified sp.-PA: France (Oligocene) [C] (Scudder, 1894: 170).

 

Genus GNOPHOMYIA Osten Sacken

*GNOPHOMYIA Osten Sacken, 1860: 223. Type species: Gnophomyia tristissima Osten Sacken, 1860, by subsequent designation of Coquillett (1910: 547).

debilis Heer, 1849: 201 (Limnobia). PA: Croatia (Miocene) [C].
gentilinii Krzeminski & Krzeminska, 1990: 496. PA: Italy (Eocene) [C].
inferna Alexander, 1931: 122. PA: Baltic Region (Eocene/Oligocene) [A].
magna Meunier, 1906f: 371. PA: Baltic Region (Eocene/Oligocene) [A].
minuta Meunier, 1906f: 369 (Tricyphona). PA: Baltic Region (Eocene/Oligocene) [A].
parvicellula Alexander, 1931: 122. PA: Baltic Region (Eocene/Oligocene) [A].
procera Meunier, 1906f: 370. PA: Baltic Region (Eocene/Oligocene) [A].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzemio(n,´)ski, 1992a: 107).

 

Genus GONOMYIA Meigen

*GONOMYIA`Meigen,`1818:`147.`Type species: Limnobia tenella Meigen, 1818, by monotypy.
[Originally proposed in synonymy; available by established use prior to 1961.]
*GONIOMYIA Osten Sacken, 1869: 176 (unjustified emendation of Gonomyia Meigen). Type species: Limnobia tenella Meigen, 1818, automatic.

Subgenus ELECTROGONOMYIA Alexander

ELECTROGONOMYIA Alexander, 1931: 118. Type species: Gonomyia pinetorum Alexander, 1931, by original designation.

pinetorum Alexander, 1931: 118. PA: Baltic Region (Eocene/Oligocene) [A].

Subgenus GONOMYIA Meigen

oligocenica Alexander, 1931: 119. PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of GONOMYIA Meigen

andrea Krzeminski & Gentilini, 1992: 92. PA: Italy (Miocene) [C].
munda Statz, 1944b: 96. PA: Germany (Oligocene) [C].
sturi Heyden, 1870: 253 (Limnobia). PA: Germany (Oligocene) [C].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).
Unidentified`sp.-PA:`Italy`(Eocene)`[C] (Krzeminski & Krzeminska, 1990: 496).

 

Genus GYMNASTES Brunetti

*GYMNASTES Brunetti, 1911: 281. Type species: Gymnastes violaceus Brunetti, 1911, by monotypy.

fasciatipennis Cockerell, 1921c: 456. PA: UK (England) (Eocene/Oligocene) [C].

 

Genus GYNOPLISTIA Macquart

*GYNOPLISTIA Macquart, 1835: 649. Type species: Gynoplistia annulata Macquart, 1835, by subsequent designation of Theischinger (1993: 5).
*GYNOPLISTIA Westwood, 1835: 280. Type species: Gynoplistia nervosa Westwood, 1835, by subsequent designation of Brunetti (1918: 332). [Preoccupied by Macquart, 1835.]
[Macquart apparently recorded taxa from a manuscript or proof sheet of Westwood in the supplement of his 1835 work and gave characters to validate them, while crediting Westwood for nothing more than the names. Macquart's work predates Westwood's and the new names must take the date and authorship from Macquart (1835: 649). See Evenhuis (1990c: 3) for further details. Theischinger's (1993) type species designation is from the 2 included species in Gynoplistia Macquart.]

mitchelli Jarzembowski, 1991: 102. PA: UK (England) (Lower Cretaceous) [C].

 

Genus HAPLOTIPULA Bode

HAPLOTIPULA Bode, 1953: 311. Type species: Haplotipula majalis Bode, 1953, by original designation.

cubitoramosa Bode, 1953: 312. PA: Germany (Lower Jurassic) [C].
majalis Bode, 1953: 311. PA: Germany (Lower Jurassic) [C].

 

Genus HELIUS Lepeletier & Serville

*MEGARHINA Lepeletier & Serville in Latreille et al., 1828: 585. Type species: Limnobia longirostris Meigen, 1818, by monotypy.
*HELIUS Lepeletier & Serville, 1828: 831 (alternate original name for Megarhina Lepeletier & Serville). Type species: Limnobia longirostris Meigen, 1818, by monotypy.
[The formation of the name Helius is considered here to be Latin and not vernacular as by Stone et al. in Alexander (1965: 50­51).]
*RHAMPHIDIA Meigen, 1830: 281. Type species: Limnobia longirostris Meigen, 1818, by subsequent designation of Westwood (1840: 129).

botswanensis`Rayner`&`Waters,`1990:`310.`AF:`Botswana`(Upper`Cretaceous)`[C].
constenius Krzeminski, 1991a: 312. NE: USA (Eocene/Oligocene) [C].
formosus Krzeminski, 1993a: 600. PA: Baltic Region (Eocene/Oligocene) [A].
minutus Loew, 1850b: 37. PA: Baltic Region (Eocene/Oligocene) [A].
succinea Meunier, 1906f: 367 (Antocha). PA: Baltic Region (Eocene/Oligocene) [A].
abditus`Krzeminski,`1985:`116.`PA:`Baltic`Region (Eocene/Oligocene) [A].
pulcher Loew, 1850b: 37 (Rhamphidia). PA: Baltic Region (Eocene/Oligocene) [A].
tenerus Statz, 1944b: 95. PA: Germany (Oligocene) [C].
weigandi Statz, 1934: 97 (Elephantomyia). PA: Germany (Oligocene) [C].

 

Genus HEXATOMA Latreille

*HEXATOMA Latreille, 1809: 260. Type species: Hexatoma nigra Latreille, 1809, by monotypy.

dimidiata Henriksen, 1922: 20 (Eriocera). PA: Denmark (Eocene) [C].
minuta Alexander, 1931: 84. PA: Baltic Region (Eocene/Oligocene) [A].
palpata Meunier, 1906f: 387 (Eriocera). PA: Baltic Region (Eocene/Oligocene) [A].
plastica Alexander, 1931: 86 (Eriocera). PA: Baltic Region (Eocene/Oligocene) [A].
succini Osten Sacken, 1869: 252 (Anisomera). PA: Baltic Region (Eocene/Oligocene) [A].
succini Loew, 1850b: 37 (Anisomera). Nomen nudum.

 

Genus HOTTIPULA Evenhuis

SINOTIPULA`Hong,`1983:`106.`Type`species:`Sinotipula huabeiensis Hong, 1983, by original designation. [Preoccupied by Alexander, 1935.] New synonymy.
HOTTIPULA Evenhuis, 1933: in this work (new replacement name for Sinotipula Hong). Type species: Sinotipula huabeiensis Hong, 1983, automatic.

huabeiensis Hong, 1983: 101 (Sinotipula). PA: China (Middle Jurassic) [C]. New combination.

 

Genus IDIOGNOPHOMYIA Alexander

*IDIOGNOPHOMYIA Alexander, 1956: 403 (as Gnophomyia Osten Sacken subgenus). Type species: Gnophomyia capicola Alexander, 1956, by original designation.

seiverti Brown, 1985: 99 (Gnophomyia). NE: USA (Oligocene) [C]. New combination.

 

Genus LEPTOTIPULOIDES Carpenter

LEPTOTIPULA Bode, 1953: 312. Type species: Leptotipula fastigata Bode, 1953, by original designation. [Preoccupied by Alexander, 1917.]
LEPTOTIPULOIDES Carpenter, 1986: 576 (new replacement name for Leptotipula`Bode).`Type`species: Leptotipula fastigata Bode, 1953, automatic.

fastigata Bode, 1953: 312 (Leptotipula). PA: Germany (Lower Jurassic) [C].

 

Genus LIASSOTIPULA Tillyard

LIASSOTIPULA Tillyard, 1933: 74. Type species: Liassotipula anglicana Tillyard, 1933, by original designation.

anglicana Tillyard, 1933: 75. PA: UK (England) (Lower Jurassic) [C].

 

Genus LIMNOCEMA Scudder

LIMNOCEMA Scudder, 1894: 201 (39). Type species: Limnocema marcescens Scudder, 1894, by subsequent designation of Cockerell (1909d: 84).

lutescens Scudder, 1894: 203 (41). NE: USA (Oligocene) [C].
marcescens Scudder, 1894: 202 (40). NE: USA (Oligocene) [C].
mortoni Scudder, 1894: 204 (42). NE: USA (Oligocene) [C].
sternbergi Cockerell, 1927a: 162. NE: USA (Oligocene) [C].
styx Scudder, 1894: 203 (41). NE: USA (Oligocene) [C].

 

Genus LIMNOPHILA Macquart

*LIMNOPHILA Macquart, 1834: 95. Type species: Limnobia pictipennis Meigen, 1818, by subsequent designation of Westwood (1840: 128).
HETEROPOECILOSTOLA Meunier, 1899f: 358. Type species: Limnophila brevipetiolata Meunier, 1906, by present designation.
[Heteropoecilostola originally proposed without included named species. Type designated herein is the first inclusion of a named species within the genus in accordance with the Code.]
HETEROLIMNOPHILA Meunier, 1906f: 376 (as Limnophila subgenus; unjustified emendation of Heteropoecilostola Meunier). Type species: Limnophila brevipetiolata Meunier, 1906, automatic (by designation of the type species for Heteropoecilostola in this work).

aliena Cockerell, 1921c: 461. PA: UK (England) (Eocene/Oligocene) [C].
brevipetiolata Meunier, 1906f: 378 . PA: Baltic Region (Eocene/Oligocene) [A].
cyclospila Cockerell & Haines, 1921a: 83 [1921b: 109]. PA: UK (England) (Eocene/Oligocene) [C].
deleta Cockerell, 1921c: 459. PA: UK (England) (Eocene/Oligocene) [C].
elegans Théobald, 1937a: 287. PA: France (Oligocene) [C].
furcata Giebel, 1856: 245 (Limnobia). PA: Baltic Region (Eocene/Oligocene) [A].
incognita Cockerell, 1921c: 462. PA: UK (England) (Eocene/Oligocene) [C].
marklae Wighton, 1980: 623. NE: Canada (Paleocene) [C].
pterotrichia Statz, 1934: 103. PA: Germany (Oligocene) [C].
rhenana Meunier, 1923c: 610. PA: Germany (Oligocene) [C].
rogersii Scudder, 1894: 218 (56). NE: USA (Oligocene) [C].
ruinarum Scudder, 1894: 220 (58). NE: USA (Oligocene) [C].
samlandica Alexander, 1931: 73. PA: Baltic Region (Eocene/Oligocene) [A].
skwarrae Alexander, 1931: 74. PA: Baltic Region (Eocene/Oligocene) [A].
statzi Evenhuis, 1994: in this work (new replacement name for robusta Statz). PA: Germany (Oligocene) [C].
robusta Statz, 1934: 102. PA: Germany (Oligocene) [C]. [Preoccupied by Wahlgren, 1905.]
strigosa Scudder, 1894: 219 (57). NE: USA (Oligocene) [C].
umbonata Statz, 1944b: 99. PA: Germany (Oligocene) [C].
vasta Scudder, 1894: 219 (57). NE: USA (Oligocene) [C].
veterana Statz, 1934: 102. PA: Germany (Oligocene) [C].

 

Genus LIMONIA Meigen

*LIMONIA`Meigen,`1803:`262. Type species: Tipula tripunctata Fabricius, 1781, automatic [by designation of Westwood (1840: 129) for Limnobia].
*AGELIA Rafinesque, 1815: 130 (unnecessary new replacement name for Limonia`Meigen).`Type species: Tipula tripunctata Fabricius, 1781, automatic.
*LIMNOBIA Meigen, 1818: 116 (unnecessary new replacement name for Limonia`Meigen).`Type species: Tipula tripunctata Fabricius, 1781, by subsequent designation of Westwood (1840: 129).
ATARACTA Loew, 1850b: 36. Type species: Ataracta grandis Meunier, 1899, by subsequent monotypy in Meunier (1899e: 172).
CORETHRIUM Westwood, 1854: 387. Type species: Corethrium pertinax Westwood, 1854, by monotypy.

albertensis Krzeminski & Teskey, 1987: 891. NE: Canada (Upper Cretaceous) [A].
antiqua`Statz, 1934: 98 (Limnobia). PA: Germany (Oligocene) [C].
cingulata Heer, 1849: 199 (Limnobia). PA: Croatia (Miocene) [C].
congruens Keilbach, 1982: 319. Nomen nudum.
deferi`Théobald,`1937a: 236 (Limnobia). PA: France (Oligocene) [C].
deleta`Giebel,`1856:`246 (Limnobia). PA: Baltic Region (Eocene/Oligocene) [A]. New combination.
flagellata Alexander, 1931: 38. PA: Baltic Region (Eocene/Oligocene) [A].
formosa`Heer,`1849:`198 (Limnobia). PA: Switzerland (Miocene) [C].
gracilipes Keilbach, 1982: 318. Nomen nudum.
gracilis Keilbach, 1982: 318. Nomen nudum.
grandis Meunier, 1899e: 172 (Ataracta). PA: Baltic Region (Eocene/Oligocene) [A].
horioni Statz, 1934: 98 (Limnobia). PA: Germany (Oligocene) [C].
latipennis Keilbach, 1982: 319. Nomen nudum.
luca Krzeminski & Gentilini, 1992: 94. PA: Italy (Miocene) [C].
meunieri Alexander, 1931: 36. PA: Baltic Region (Eocene/Oligocene) [A].
murchisoni Heer, 1856: 30 (Limnobia). PA: France (Oligocene) [C].
curtisi Giebel, 1856: 245 (Limnobia). PA: France (Oligocene) [C].
pertinax Westwood, 1854: 387 (Corethrium). PA: UK (England) (Lower Cretaceous) [C].
remota Keilbach, 1982: 319. Nomen nudum.
rostrata Keilbach, 1982: 319. Nomen nudum.
sphenoptera Keilbach, 1982: 319. Nomen nudum.
spilota Cockerell, 1921c: 459 (Limnobia). PA: UK (England) (Eocene/Oligocene) [C].
tenuis Heer, 1849: 200 (Limnobia). PA: Croatia (Miocene) [C].
theobaldi Piton, 1940: 214 (Limnobia). PA: France (Oligocene) [C].
vetusta Heer, 1849: 200 (Limnobia). PA: Croatia (Miocene) [C].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).

 

Genus LIPSOTHRIX Loew

*LIPSOTHRIX Loew, 1873a: 67. Type species: Lipsothrix nobilis Loew, 1873, by monotypy.
ELECTROLABIS Alexander, 1931: 58. Type species: Electrolabis extincta Alexander, 1931, by original designation.

extincta Alexander, 1931: 58 (Electrolabis). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus MACALPINA Krzeminski & Teskey

MACALPINA Krzeminski & Teskey, 1987: 890. Type species: Macalpina incomparabilis Krzeminski & Teskey, 1987, by original designation.

incomparabilis Krzeminski & Teskey, 1987: 890. NE: Canada (Upper Cretaceous) [A].

 

Genus MESOTIPULA Handlirsch

MESOTIPULA Handlirsch, 1920: 205 [1939: 107]. Type species: Mesotipula brachyptera Handlirsch, 1920, by monotypy.

brachyptera Handlirsch, 1920: 205 [1939: 107]. PA: Germany (Lower Jurassic) [C].
curvata Handlirsch, 1939: 107. PA: Germany (Lower Jurassic) [C].
mirabilis Krzeminski & Zessin, 1990: 40. PA: Germany (Lower Jurassic) [C].
siggae Krzeminski & Zessin, 1990: 40. PA: Germany (Lower Jurassic) [C].
vicina Handlirsch, 1939: 109 (Architipula). PA: Germany (Lower Jurassic) [C].

 

Genus MIKROTIPULA Bode

MIKROTIPULA Bode, 1953: 314. Type species: Mikrotipula dixaeformis Bode, 1953, by original designation.

dixaeformis Bode, 1953: 314. PA: Germany (Lower Jurassic) [C].

 

Genus MOLOPHILUS Curtis

*MOLOPHILUS Curtis, 1833: pl. 444. Type species: Molophilus brevipennis Curtis, 1833 [= Erioptera atra Meigen, 1804], by original designation.

Subgenus MIOMOLOPHILUS Krzeminski

MIOMOLOPHILUS Krzeminski, 1992b: 108. Type species: Molophilus theischingeri Krzeminski, 1992, by original designation.

theischingeri`Krzeminski, `1992b:`108. NT: Dominican Republic (Oligocene/Miocene) [A].

 

Genus NEMUMEIA Kalugina

NEMUMEIA Kalugina, 1986: 115. Type species: Nemumeia ponomarenkoi Kalugina, 1986, by original designation.

ponomarenkoi Kalugina, 1986: 116. PA: Mongolia (Upper Jurassic/Lower Cretaceous) [C].

 

Genus ORMOSIA Rondani

*ORMOSIA Rondani, 1856: 180. Type species: Erioptera nodulosa Macquart, 1826, by original designation.
GONOMYIELLA Meunier, 1899d: 335. Type species: Ormosia electrella Alexander, 1931, by present designation.
[Gonomyiella originally proposed without included species. The type designated is from the first included species within the genus in accordance with the Code.]
GONOMYELLA. Incorrect original spelling of Gonomyiella (Meunier, 1899d: 334).

beurleni Alexander, 1931: 102. PA: Baltic Region (Eocene/Oligocene) [A].
electrella Alexander, 1931: 101. PA: Baltic Region (Eocene/Oligocene) [A].
skwarrae Alexander, 1931: 100. PA: Baltic Region (Eocene/Oligocene) [A].
tornquisti Alexander, 1931: 102. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus ORYCTOGMA Scudder

ORYCTOGMA Scudder, 1894: 194 (32). Type species: Oryctogma sackenii Scudder, 1894, by monotypy.

sackenii Scudder, 1894: 195 (33). NE: USA (Oligocene) [C].

 

Genus OZOTIPULA Bode

OZOTIPULA Bode, 1953: 313. Type species: Ozotipula tarda Bode, 1953, by original designation.

tarda Bode, 1953: 313. PA: Germany (Lower Jurassic) [C].

 

Genus PALAEOMACROMASTIX Meunier

PALAEOMACROMASTIX Meunier, 1917a: 100. Type species: Palaeomacromastix bornhardti Meunier, 1917, by monotypy.

bornhardti Meunier, 1917a: 100. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PALAEOPOECILOSTOLA Meunier

PALAEOPOECILOSTOLA Meunier, 1899d: 334. Type species: Lasiomastix longicornis Meunier, 1906, by subsequent monotypy in Handlirsch (1907c: 995).

fastuosa Meunier, 1906f: 381 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].
longicornis Meunier, 1906f: 377 (Lasiomastix). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
ssp. longicornis s. str. PA: Baltic Region (Eocene/Oligocene) [A].
longicornis Loew, 1850: 37 (Limnophila). Nomen nudum.
robusta Meunier, 1906f: 383 (Limnophila). PA: Baltic Region (Eocene/ Oligocene) [A]. [Preoccupied by Wahlgren, 1905.]
palaea Bergroth, 1913: 583 (Tipula; new replacement name for robusta Meunier). PA: Baltic Region (Eocene/Oligocene) [A].
ssp. parallela Alexander, 1931: 48. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
speciosa Meunier, 1906f: 384 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PHYLIDOREA Bigot

*PHYLIDOREA Bigot, 1854: 456. Type species: Limnobia ferruginea Meigen, 1818, by subsequent designation of Coquillett (1910: 590).

pulchripennis Meunier, 1906f: 380 (Limnophila). PA: Baltic Region (Eocene/ Oligocene) [A]. New combination.
servilis Alexander, 1931: 72 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A]. New combination.

 

Genus PHYLLOLABIS Osten Sacken

*PHYLLOLABIS Osten Sacken, 1877: 202. Type species Phyllolabis claviger Osten`Sacken,`1877,`by`subsequent`designation`of Coquillett (1910: 590).

andreei Alexander, 1931: 52. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PILARIA Sintenis

*PILARIA Sintenis, 1889: 398. Type species: Limnobia pilicornis Zetterstedt, 1851 (as "Limnophila") [= Limnobia meridiana Staeger, 1840], by subsequent designation of Coquillett (1910: 591).
[The wording in Sharp (1890: 281) is here interpreted as not constituting a valid type designation.]

baltica Alexander, 1931: 82. PA: Baltic Region (Eocene/Oligocene) [A].
batheri Alexander, 1931: 81. PA: Baltic Region (Eocene/Oligocene) [A].
castellariana Krzeminski & Gentilini, 1992: 88. PA: Italy (Miocene) [C].
electrina Cockerell & Clark, 1918: 115 (Limnophila). PA: UK (England) (Eocene/Oligocene) [C].
elongata Meunier, 1906f: 381 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus POLYMERA Wiedemann

*POLYMERA Wiedemann, 1820b: 40. Type species: Chironomus hirticornis Fabricius, 1805, by monotypy.

magnifica Meunier, 1906f: 385. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus PRAEARCHITIPULA Kalugina

PRAEARCHITIPULA Kalugina in Kalugina & Kovalev, 1985: 56. Type species: Praearchitipula notabilis Kalugina, 1985, by original designation.

lata Kalugina in Kalugina & Kovalev, 1985: 59. PA: Russia (Siberia) (Middle Jurassic) [C].
notabilis`Kalugina`in Kalugina & Kovalev, 1985: 57. PA: Russia (Siberia) (Middle Jurassic) [C].
parvula Kalugina in Kalugina & Kovalev, 1985: 57. PA: Russia (Siberia) (Middle Jurassic) [C].
spasskia`Kalugina`in Kalugina & Kovalev, 1985: 59. PA: Russia (Siberia) (Middle Jurassic) [C].
trinervis`Kalugina`in Kalugina & Kovalev, 1985: 57. PA: Russia (Siberia) (Middle Jurassic) [C].

 

Genus PRONOPHLEBIA Scudder

PRONOPHLEBIA Scudder, 1877a: 750. Type species: Pronophlebia rediviva Scudder, 1877, by monotypy.

rediviva Scudder, 1877a: 750. NE: USA (Eocene) [C].


Genus PROTOLIMNOPHILA Riek

PROTOLIMNOPHILA Riek, 1954: 58. Type species: Protolimnophila superba Riek, 1954, by original designation.

superba Riek, 1954: 58. AU: Australia (Eocene) [C].

 

Genus PSEUDOLIMNOPHILA Alexander

*PSEUDOLIMNOPHILA Alexander, 1919: 917 (as Limnophila subgenus). Type species: Limnophila luteipennis Osten Sacken, 1860, by subsequent designation of Alexander (1920a: 849).

Subgenus BALTILIMNOPHILA Krzeminski

BALTILIMNOPHILA Krzeminski, 1990a: 53. Type species: Limnophila elegantissima Meunier, 1906, by original designation.

elegantissima Meunier, 1906b: 104 (Limnophila). PA: Baltic Region (Eocene/ Oligocene) [A].
pulcherrima`Krzeminski,`1990a:`55. PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of PSEUDOLIMNOPHILA Alexander

ambigua Alexander, 1931: 70. PA: Baltic Region (Eocene/Oligocene) [A].
concinna Meunier, 1906f: 380 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].
continuata Meunier, 1906f: 379 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].
exigua Meunier, 1906f: 377 (Prionolabis). PA: Baltic Region (Eocene/Oligocene) [A].
inculpata Alexander, 1931: 68. PA: Baltic Region (Eocene/Oligocene) [A].
loewiella Alexander, 1931: 63 (new replacement name for maculata Meunier). PA: Baltic Region (Eocene/Oligocene) [A].
maculata Meunier, 1916b: 491 (Limnophila; as concinna subspecies). PA: Baltic Region (Eocene/Oligocene) [A]. [Secondary junior homonym of Limonia maculata Meigen, 1804; loewiella valid under I.C.Z.N. Code Article 59(b).]
pinicola Alexander, 1931: 67. PA: Baltic Region (Eocene/Oligocene) [A].
producta Meunier, 1906f: 376 (Prionolabis). PA: Baltic Region (Eocene/Oligocene) [A].
siciliana Krzeminski & Skalski, 1983: 303. PA: Sicily (Miocene) [A].
speirsae Wighton, 1980: 622. NE: Canada (Paleocene) [C].
timida Alexander, 1931: 65. PA: Baltic Region (Eocene/Oligocene) [A].
vulcana Meunier, 1906f: 381 (Limnophila). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus RHABDOMASTIX Skuse

*RHABDOMASTIX Skuse, 1890a: 828. Type species: Rhabdomastix ostensackeni Skuse, 1890, by monotypy.

Subgenus PALAEOGONOMYIA Meunier

PALAEOGONOMYIA Meunier, 1899f: 359. Type species: Gonomyia pulcherrima Meunier, 1906, by subsequent designation of Alexander (1931: 112).

borussica Meunier, 1906f: 374 (Gonomyia). PA: Baltic Region (Eocene/Oligocene) [A].
brevis Alexander, 1931: 117. PA: Baltic Region (Eocene/Oligocene) [A].
elegantula Meunier, 1906f: 372 (Gonomyia). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
klebsi Alexander, 1931: 113. PA: Baltic Region (Eocene/Oligocene) [A].
pulchella Meunier, 1906f: 373 (Gonomyia). PA: Baltic Region (Eocene/Oligocene) [A].
pulcherrima Meunier, 1906f: 372 (Gonomyia). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
ssp. graciosa Meunier, 1906f: 373 (Gonomyia; as species). PA: Baltic Region (Eocene/Oligocene) [A].
ssp. pulcherrima s. str. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
pulchra Meunier, 1906f: 374 (Gonomyia). PA: Baltic Region (Eocene/Oligocene) [A].

Subgenus SACANDAGA Alexander

*SACANDAGA Alexander, 1911: 349 (as genus). Type species: Sacandaga flava Alexander, 1911, by original designation.

praecursor Cockerell, 1927a: 161. NE: USA (Oligocene) [C].
Unidentified sp.-PA: Germany (Miocene) [A] (Schumann & Wendt, 1989: 41).

Unplaced to Subgenus of RHABDOMASTIX Skuse

frigida Scudder, 1894: 212 (50) (Gonomyia). NE: USA (Oligocene) [C].
labefactata Scudder, 1894: 210 (48) (Gonomyia). NE: USA (Oligocene) [C].
oligocaenica Statz, 1944b: 98. PA: Germany (Oligocene) [C].
primogenitalis Scudder, 1894: 211 (49) (Gonomyia). NE: USA (Oligocene) [C].
profundi Scudder, 1894: 210 (48) (Gonomyia). NE: USA (Oligocene) [C].
scudderi Cockerell, 1920a: 247 (Gonomyia). NE: USA (Eocene) [C].

 

Genus RHADINOBROCHUS Scudder

RHADINOBROCHUS Scudder, 1894: 223 (61). Type species: Rhadinobrochus extinctus Scudder, 1894, by monotypy.

extinctus Scudder, 1894: 223 (61). NE: USA (Oligocene) [C].

 

Genus RHIPIDIA Meigen

*RHIPIDIA Meigen, 1818: 153. Type species: Rhipidia maculata Meigen, 1818, by monotypy.

brodiei Cockerell, 1917b: 373. PA: UK (England) (Eocene/Oligocene) [C].
extincta Unger, 1841: 425. PA: Croatia (Miocene) [C].
picta Heer, 1849: 197. PA: Croatia (Miocene) [C].
propinqua Heer, 1849: 198. PA: Croatia (Miocene) [C].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).


Genus RYTA Nazarov

RYTA Nazarov, 1994: 106. Type species: Ryta berestiana Nazarov, 1994, by original designation.

berestiana Nazarov, 1994: 107. PA: Belarus (Eocene) [A].


Genus SPILADOMYIA Scudder

SPILADOMYIA Scudder, 1877a: 750. Type species: Spiladomyia simplex Scudder, 1877, by monotypy.

simplex Scudder, 1877a: 750. NE: USA (Eocene) [C].

 

Genus STYRINGOMYIA Loew

STYRINGIA Berendt, 1845: 57. Nomen nudum.
STYRINGIA Bronn, 1847: 597. Nomen nudum.
*STYRINGOMYIA Loew, 1845: 6. Type species: Styringomyia venusta Loew, 1845, by monotypy.
MESOMYITES Cockerell, 1917b: 377. Type species: Mesomyites concinnus Cockerell, 1917, by original designation.
*SYRINGOMYIA Alexander, 1972: 841 (unjustified emendation of Styringomyia Loew). Type species: Styringomyia venusta Loew, 1845, automatic.

concinna Cockerell, 1917b: 377 (Mesomyites). PA: UK (England) (Eocene/Oligocene) [C].
extensa Cockerell & Haines, 1921a: 83 [1921b: 112]. PA: UK (England) (Eocene/Oligocene) [C].
gracilis Loew, 1850b: 38. PA: Baltic Region (Eocene/Oligocene) [A].
[Species-group name valid by characters given in the key (Loew, 1850b: 36). Though Loew mentions both venusta and gracilis under Styringomyia (p. 38), the key is to Baltic amber fossils only and gracilis is the only Baltic amber species of Styringomyia mentioned by Loew.]
pulchella Loew, 1845: 7. Nomen nudum.
venusta Loew, 1845: 6. OR: "Ostindische" (Holocene) [K].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).

 

Genus SYMPLECTA Meigen

*SYMPLECTA Meigen, 1830: 282. Type species: Limnobia punctipennis Meigen, 1818 [= Limonia hybrida Meigen, 1804], by subsequent designation of Westwood (1840: 128).

Subgenus PSILOCONOPA Zetterstedt

*PSILOCONOPA Zetterstedt, 1837: 64 [1838: 847] (as genus). Type species: Psiloconopa meigeni Zetterstedt, 1837, by monotypy.
MIOPSILOPTERA Gentilini, 1984: 175 (as genus). Type species: Miopsiloptera savtschenkoi Gentilini, 1984, by original designation.

gracilis Théobald, 1937a: 345 (Ilisia). PA: France (Oligocene) [C].
savtschenkoi Gentilini, 1984: 175 (Miopsiloptera). PA: Italy (Miocene) [C].

 

Genus TANYMERA Loew

TANYMERA Loew, 1850b: 36. Type species: Tanymera gracilicornis Osten Sacken, 1869, by subsequent monotypy in Osten Sacken (1869: 193).
[Tanymera originally proposed without included species. The type species is the first inclusion of a named species by Osten Sacken (1869: 193).]
TANYMERA Alexander, 1931: 75. Type species: Tanymera fritschi Alexander, 1931, by original designation. [Preoccupied by Loew, 1850.]

annulata Meunier, 1899e: 173. PA: Baltic Region (Eocene/Oligocene) [A].
arguta Alexander, 1931: 76. PA: Baltic Region (Eocene/Oligocene) [A].
berendti Alexander, 1931: 79. PA: Baltic Region (Eocene/Oligocene) [A].
crassicornis Meunier, 1899e: 173. PA: Baltic Region (Eocene/Oligocene) [A].
fritschi Alexander, 1931: 78. PA: Baltic Region (Eocene/Oligocene) [A].
gracilicornis Osten Sacken, 1869: 193. PA: Baltic Region (Eocene/Oligocene) [A].
terminans Alexander, 1931: 78. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus TANYSPHYRA Loew

TANYSPHYRA Berendt, 1845: 57. Nomen nudum.
TANYSPHYRA Bronn, 1847: 597. Nomen nudum.
TANYSPHYRA Loew, 1850b: 36, 38. Type species: Tanysphyra gracilis Loew, 1850 [preoccupied, = Limnophila meunieri Cockerell, 1909], by monotypy.
POECILOSTIELLA Meunier, 1899d: 334. Type species: Tanysphyra gracilis Loew, 1850 [preoccupied, = Limnophila meunieri Cockerell, 1909], by present designation.
[Poecilostiella originally proposed without included species. The present type designation is from the first included species herein in accordance with the Code.]

meunieri Cockerell, 1909b: 58 (Limnophila; new replacement name for gracilis Loew). PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
gracilis Loew, 1850b: 38. PA: Baltic Region (Eocene/Oligocene) [A]. [Preoccupied by Wiedemann, 1828 in Limnophila; meunieri valid by IC.Z.N. Code 59(b).]

 

Genus TASIOCERA Skuse

*TASIOCERA Skuse, 1890a: 815. Type species: Tasiocera tenuicornis Skuse, 1890, by subsequent designation of Alexander (1920b: 52).

Subgenus DASYMOLOPHILUS Goetghebuer

*DASYMOLOPHILUS Goetghebuer in Goetghebuer & Tonnoir, 1920: 132 (as Molophilus subgenus). Type species: Erioptera murina Meigen, 1818, by original designation.

circumcincta Meunier, 1906f: 368 (Erioptera). PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus THAUMASTOPTERA Mik

*THAUMASTOPTERA Mik, 1866: 302. Type species: Thaumastoptera calceata Mik, 1866, by monotypy.

electra Alexander, 1931: 41. PA: Baltic Region (Eocene/Oligocene) [A].
ryszardi Krzeminski, 1985: 115. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus TOXORHINA Loew

TOXORHINA Berendt, 1845: 57. Nomen nudum.
TOXORHINA Bronn, 1847: 597. Nomen nudum.
*TOXORHINA Loew, 1850b: 36, 37. Type species: Toxorhina fragilis Loew, 1851, by subsequent designation of Osten Sacken (1869: 113).
TOXORRHINA, error for Toxorhina Loew.

Subgenus CERATOCHEILUS Wesché

*CERATOCHEILUS Wesché, 1910: 358 (as genus). Type species: Ceratocheilus winnsampsoni Wesché, 1910 [= Styringomyia cornigera Speiser, 1908], by subsequent designation of Brunetti (1920: 230).

caucasiensis Krzeminski & Freiwald, 1991: 155. PA: Russia (Stavropol) (Miocene) [C].
eridana Meunier, 1917a: 96 (Ceratocheilus). PA: Baltic Region (Eocene/Oligocene) [A].
Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Krzeminski, 1992a: 107).

Unplaced to Subgenus of TOXORHINA Loew

madagascariensis Meunier, 1906c: 98. AF: Madagascar (Holocene) [K].
madagascariensis Meunier, 1905d: 89. Nomen nudum.

 

Genus TRENTEPOHLIA Bigot

*TRENTEPOHLIA Bigot, 1854: 456, 473. Type species: Limnobia trentepohlii Wiedemann, 1828 (as "Trentepohlia limnobioides"), by monotypy.

Subgenus MONGOMA Westwood

*MONGOMA Westwood, 1881: 364 (as genus). Type species: Mongoma fragillima Westwood, 1881, by monotypy.

cruciferella Cockerell, 1917b: 373 (Mongoma). PA: UK (England) (Eocene/Oligocene) [C]. New combination.
pallescens Cockerell, 1921c: 460 (Mongoma). PA: UK (England) (Eocene/Oligocene) [C]. New combination.

Subgenus TRENTEPOHLIA Bigot

dampfiana Alexander, 1931: 123. PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of TRENTEPOHLIA Bigot

Unidentified sp.-NT: Dominican Republic (Oligocene/Miocene) [A] (Davis, 1989: 548; Krzeminski, 1992a: 107).

 

Genus TRICHONEURA Loew

TRICHONEURA Berendt, 1845: 57. Nomen nudum.
TRICHONEURA Bronn, 1847: 597. Nomen nudum.
*TRICHONEURA Loew, 1850b: 36. Type species: Trichoneura vulgaris Loew, 1850, by monotypy.
SACKENIELLA Meunier, 1894d: clxxviii (1895a: clxxviii). Type species: Trichoneura vulgaris Loew, 1850, by subsequent monotypy in Speiser (1900: 317).
SACKANIELLA. Incorrect original spelling of Sackeniella [Meunier, 1894d: clxxviii (1895a: clxxviii)].

Subgenus TRICHONEURA Loew

gracilistylus Alexander, 1931: 57. PA: Baltic Region (Eocene/Oligocene) [A].
longicornis Keilbach, 1982: 320. Nomen nudum.
ritzkowskii Krzeminski, 1990b: 189. PA: Baltic Region (Eocene/Oligocene) [A].
vulgaris Loew, 1850b: 36. PA: Baltic Region (Eocene/Oligocene) [A]; PA: Germany (Miocene) [A].
decipiens Meunier, 1906f: 387. PA: Baltic Region (Eocene/Oligocene) [A].
prolifica Meunier, 1916b: 492 (as vulgaris subspecies). PA: Baltic Region (Eocene/Oligocene) [A].

Unplaced to Subgenus of TRICHONEURA Loew

canadensis Krzeminski & Teskey, 1987: 888. NE: Canada (Upper Cretaceous) [A].
grubei Keilbach, 1982: 320. Nomen nudum.
heini Keilbach, 1982: 320. Nomen nudum.

 

Genus TRICYPHONA Zetterstedt

*TRICYPHONA Zetterstedt, 1837: 65 [1838: 551]. Type species: Limonia immaculata Meigen, 1804, by monotypy.

electrina Alexander, 1931: 95. PA: Baltic Region (Eocene/Oligocene) [A].
sepulchralis Alexander, 1931: 94. PA: Baltic Region (Eocene/Oligocene) [A].
succinea Alexander, 1931: 94. PA: Baltic Region (Eocene/Oligocene) [A].

 

Genus ULA Haliday

*ULA Haliday, 1833: 153. Type species: Ula mollissima Haliday, 1833, by monotypy.
HAPLONEURA Loew, 1850b: 36. Type species: Haploneura hirtipennis Osten Sacken, 1869, by subsequent monotypy in Osten Sacken (1869: 275).

hirtipennis Osten Sacken, 1869: 275 (Haploneura). PA: Baltic Region (Eocene/ Oligocene) [A].
rottensis Statz, 1934: 101. PA: Germany (Oligocene) [C].

 

Genus XUTIPULA Hong

XUTIPULA Hong, 1983: 108. Type species: Xutipula longipetalis Hong, 1983, by original designation.

longipetalis Hong, 1983: 108. PA: China (Middle Jurassic) [C].

 

Unplaced Species of LIMONIIDAE

ferrea Cockerell, 1921c: 460 (Gonomyia). PA: UK (England) (Eocene/Oligocene) [C].
grisea Cockerell, 1921c: 461 (Gonomyia). PA: UK (England) (Eocene/Oligocene) [C].
indecisa Cockerell & Haines, 1921a: 83 [1921b: 111] (Gonomyia). PA: UK (England) (Eocene/Oligocene) [C].
lutescens Cockerell, 1921c: 461 (Gonomyia). PA: UK (England) (Eocene/Oligocene) [C].

 

This page last revised 17 November 2001