Family PTYCHOPTERIDAE

 

 

Ptychopteridae, or false crane flies, are small to medium-sized flies very similar in general habitus to crane flies (Tipulidae and related families). More than 60 described species are known worldwide. Immatures of living forms are found in saturated mud along stream edges.

The family is an ancient lineage, especially as it recently had the diverse fossil assemblage of taxa in the Eoptychopteridae transferred to it (Lukashevich, 2008). Examination of type material of many species in Lukashevich et al. (1998) clarified the relationships of many of the genera in the fossil Ptychopteridae, including Eoptychoptera Handlirsch, which now has a number of synonymous genera described by Handlirsch and Bode.

 

In addition to the taxa listed below, a single species has been described by Kalugina (1989) from the Jurassic deposits of Siberia, but its generic affinities are unknown and it is listed in this catalog as unplaced to genus. A pupa of an undetermined species of Ptychoptera from the Jurassic of Siberia was recorded by Brauer et al. (1889), but needs to be examined to confirm the generic identification. It is provisionally listed here.



Ref.: Lukashevich et al. (1998, revision of Eoptychopteridae), Lukashevich (2008, review of fossil taxa).



 

 

Genus ARCHITENDIPES Rohdendorf

 

ARCHITENDIPES Rohdendorf, 1962: 317 (1991: 459). Type species: Architendipes tshernovskiji Rohdendorf, 1962, by original designation.


PALAEOTENDIPES Rohdendorf, 1962: 317 (1991: 459). Type species: Palaeotendipes alexii Rohdendorf, 1962, by original designation.

tshernovskiji Rohdendorf, 1962: 317 (1991: 459). PA: Kirghizstan (Lower Jurassic) [C].



            
alexii Rohdendorf, 1962: 317 (1991: 459). PA: Kirghizstan (Lower Jurassic) [C].

 

 

Genus BOLBOIA Kalugina

 

BOLBOIA Kalugina, 1989: 73 (1990: 70). Type species: Bolboia mira Kalugina, 1989, by original designation.

mira Kalugina, 1989: 75: (1990: 71). PA: Russia (Siberia) (Upper Jurassic/Lower Cretaceous) [C].

 

 

Genus BRODLIKA Lukashevich, Coram & Jarzembowski

 

BRODLIKA Lukashevich, Coram & Jarzembowski, 2001: 457. Type species: Brodlika mitchelli Lukashevich, Coram & Jarzembowski, 2001, by original designation.

mitchelli Lukashevich, Coram & Jarzembowski, 2001: 457. PA: UK (England) (Lower Cretaceous) [C].

 

 

Genus CRENOPTYCHOPTERA Kalugina

 

CRENOPTYCHOPTERA Kalugina in Kalugina & Kovalev, 1985: 41. Type species: Crenoptychoptera antica Kalugina, 1985, by original designation.

antica Kalugina in Kalugina & Kovalev, 1985: 42. PA: Russia (Siberia) (Middle Jurassic) [C].


bavarica Krzeminski & Ansorge, 1995: 2. PA: Germany (Upper Jurassic) [C].

conspecta Lukashevich, 1995: 168. PA: Russia (Siberia) (Lower/Middle Jurassic) [C].

decorosa Hao, Dong & Ren, 2009: 106. PA: China (Middle Jurassic) [C].

defossa Kalugina in Kalugina & Kovalev, 1985: 42. PA: Russia (Siberia) (Middle Jurassic) [C].


dobbertinensis Ansorge in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 332. PA: Germany (Lower Jurassic) [C].

gronskayae Kalugina, 1989: 71 (1990: 68). PA: Russia (Siberia) (Lower Cretaceous) [C].

liturata Lukashevich, 2011: 28(623). PA: Mongolia (Upper Jurassic) [C].

vicina Hao, Dong & Ren, 2009: 108. PA: China (Middle Jurassic) [C].

vulgaris Hao, Dong & Ren, 2009: 107. PA: China (Middle Jurassic) [C].

 

 

Genus DOPTYCHOPTERA Lukashevich

 

DOPTYCHOPTERA Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 337. Type species: Doptychoptera baisica Lukashevich, 1998, by original designation.

baisica Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 338. PA: Russia (Siberia) (Lower Cretaceous) [C].

 

 

Genus EOLIMNOBIA Handlirsch

 

EOLIMNOBIA Handlirsch, 1906a: 489. Type species: Eolimnobia geinitzi Handlirsch, 1906, by monotypy.

geinitzi Handlirsch, 1906a: 489. PA: Germany (Lower Jurassic) [C].

 

 

Genus EOPTYCHOPTERA Handlirsch

 

EOPTYCHOPTERA Handlirsch, 1906a: 489. Type species: Eoptychoptera simplex Handlirsch, 1906, by monotypy.

PROPTYCHOPTERA Handlirsch, 1906a: 489. Type species: Proptychoptera liasina Handlirsch, 1906, by monotypy.

METAXYBITTACUS Bode, 1953: 287. type species: Metaxybittacus eximius Bode, 1953, by original designation.

ACRITORHYPHUS Bode, 1953: 299. Type species: Acritorhyphus ramosus Bode, 1953, by original designation.

PALAEOLIMNOBIA Bode, 1953: 301. Type species: Palaeolimnobia maculosa Bode, 1953, by original designation.

aequidistans Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 324. PA: Kirghizistan (Lower Jurassic) [C].

ansorgi Ren & Krzeminski, 2002: 208. PA: China (Middle Jurassic) [C].

altaica Kalugina, 1988: 83 (Proptychoptera). PA: Mongolia (Lower Cretaceous) [C].


asiatica Lukashevich, 1994: 111 (Proptychoptera). PA: Kazakhstan (Upper Jurassic) [C].


aucta Lukashevich, 1994: 113 (Proptychoptera). PA: Kazakhstan (Upper Jurassic) [C].


britannica Lukashevich, Coram & Jarzembowski, 2001: 454. PA: UK (England) (Lower Cretaceous) [C].

cretacea Kalugina, 1989: 72 (1990: 69) (Proptychoptera). PA: Russia (Siberia) (Lower Cretaceous) [C].


elevata Lukashevich, 2000b: 29. PA: Russia (Siberia) (Upper Jurassic/Lower Cretaceous) [C].

eximia Bode, 1953: 288 (Metaxybittacus). PA: Germany (Lower Jurassic) [C].

      maculosa Bode, 1953: 301 (Palaeolimnobia). PA: Germany (Lower Jurassic) [C].

gronskayae Lukashevich, 1995: 168. PA: Russia (Siberia) (Lower Cretaceous) [C].

jurassica Ren & Krzeminski, 2002: 208. PA: China (Middle Jurassic) [C].

longifurcata Lukashevich, Coram & Jarzembowski, 2001: 454. PA: UK (England) (Lower Cretaceous) [C].

magna Lukashevich, 1994: 113 (Proptychoptera). PA: Kazakhstan (Upper Jurassic) [C].


maxima Kalugina in Kalugina & Kovalev, 1985: 41 (Proptychopterina). PA: Russia (Siberia) (Middle Jurassic) [C].


modica Lukashevich, 1994: 114 (Proptychoptera). PA: Russia (Siberia) (Middle Jurassic) [C].


praemaculata Kalugina in Kalugina & Kovalev, 1985: 40 (Proptychoptera). PA: Russia (Siberia) (Middle Jurassic) [C].


shurabica Lukashevich, 2000b: 27. PA: Kirghizistan (Lower/Middle Jurassic) [C].

simplex Handlirsch, 1906a: 489. PA: Germany (Lower Jurassic) [C].

      liasina Handlirsch, 1906a: 489 (Proptychoptera). PA: Germany (Lower Jurassic) [C].


      maculata Handlirsch, 1920: 204 [1939: 106] (Proptychoptera). PA: Germany (Lower Jurassic) [C].


      megapolitana Handlirsch, 1939: 106 (Proptychoptera). PA: Germany (Lower Jurassic) [C].


      similis Handlirsch, 1939: 106 (Proptychoptera). PA: Germany (Lower Jurassic) [C].

      ramosus Bode, 1953: 299 (Acritorhyphus). PA: Germany (Lower Jurassic) [C].

      minor Bode, 1953: 300 (Proptychoptera). PA: Germany (Lower Jurassic) [C].


      ovalis Bode, 1953: 301 (Proptychoptera). PA: Germany (Lower Jurassic) [C].


tempestila Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 326. PA: Mongolia (Upper Jurassic) [C].

vitrea Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 325. PA: Kirghizistan (Lower Jurassic) [C].

 

 

Genus EOPTYCHOPTERINA Kalugina

 

EOPTYCHOPTERINA Kalugina in Kalugina & Kovalev, 1985: 37. Type species: Eoptychopterina rohdendorphi Kalugina, 1985, by original designation.

abbreviata Lukashevich, 1994: 107. PA: Kazakhstan (Upper Jurassic) [C].


adnexa Hao, Ren & Shih, 2009b: 223. PA: China (Lower Cretaceous) [C].

angularis Lukashevich, 1994: 107. PA: Kazakhstan (Upper Jurassic) [C].


antica Hao, Ren & Shih, 2009b: 223. PA: China (Lower Cretaceous) [C].

baisica Kalugina, 1989: 68 (1990: 67). PA: Russia (Siberia) (Lower Cretaceous) [C].


camura Lukashevich, Coram & Jarzembowski, 2001: 456. PA: UK (England) (Lower Cretaceous) [C].

daiensis Kalugina, 1989: 68 (1990: 64). PA: Russia (Siberia) Upper Jurassic/ Lower Cretaceous) [C].


demissa Lukashevich, Coram & Jarzembowski, 2001: 455. PA: UK (England) (Lower Cretaceous) [C].

dimidiata Lukashevich, Coram & Jarzembowski, 2001: 456. PA: UK (England) (Lower Cretaceous) [C].

elenae Ren & Krzeminski, 2002: 209. PA: China (Middle Jurassic) [C].

gigantea Zhang, 2004a: 174. PA: China (Middle Jurassic) [C].

glaber Lukashevich, 1994: 110. PA: Russia (Siberia) (Middle Jurassic) [C].


kaluginae Lukashevich, 2004: 69 (304). PA: Mongolia (Upper Jurassic) [C].

karatavica Lukashevich, 1994: 106. PA: Kazakhstan (Upper Jurassic) [C].


mediata Hao, Ren & Shih, 2009b: 224. PA: China (Lower Cretaceous) [C].

omissa Lukashevich, 2004: 69 (304). PA: Kazakhstan (Upper/Middle Jurassic) [C].

petri Lukashevich, 2004: 70 (305). PA: Russia (Siberia) (Upper Jurassic/Lower Cretaceous) [C].

postica Liu, Shih & Ren, 2012: 56. PA: China (Middle Jurassic) [C].

rohdendorphi Kalugina in Kalugina & Kovalev, 1985: 39. PA: Russia (Siberia) (Lower/Middle Jurassic) [C].


transbaicalica Kalugina in Kalugina & Kovalev, 1985: 39. PA: Russia (Siberia) (Upper Jurassic) [C].


undensis Kalugina, 1989: 66 (1990: 63). PA: Russia (Siberia) (Upper Jurassic/ Lower Cretaceous) [C].

 

 

Genus ETOPTYCHOPTERA Handlirsch

 

ETOPTYCHOPTERA Handlirsch, 1909: fig. 5. Type species: Etoptychoptera tertiaria Handlirsch, 1909, by monotypy.

tertiaria Handlirsch, 1909: fig. 5 [1910: 122]. NE: Canada (Oligocene) [C].

 

 

Genus LEPTYCHOPTERA Lukashevich & Azar

 

LEPTYCHOPTERA Lukashevich & Azar, 2003: 196. Type species: Leptychoptera dimkina Lukashevich & Azar, 2003, by original designation.

 

Subgenus BURMAPTYCHOPTERA Lukashevich

 

BURMAPTYCHOPTERA Lukashevich in Lukashevich & Grimaldi, 2004: 360. Type species: Leptychoptera reburra Lukashevich, 2004, by original designation.

calva Lukashevich in Lukashevich & Grimaldi, 2004: 361. OR: Burma (Upper Cretaceous) [A].

reburra Lukashevich in Lukashevich & Grimaldi, 2004: 361. OR: Burma (Upper Cretaceous) [A].

 

Subgenus LEPTYCHOPTERA Lukashevich & Azar

 

dimkina Lukashevich & Azar, 2003: 197. PA: Lebanon (Lower Cretaceous) [A].

vovkina Lukashevich & Azar, 2003: 202. PA: Lebanon (Lower Cretaceous) [A].

 

 

Genus NEDOPTYCHOPTERA Lukashevich

 

NEDOPTYCHOPTERA Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 338. Type species: Neodoptychoptera karavatica Lukashevich, 1998, by original designation.

karavatica Lukashevich in Lukashevich, Ansorge, Krzeminski & Krzeminska, 1998: 339. PA: Kazakhstan (Upper Jurassic) [C].

 

 

Genus PROBITTACOMORPHA Freiwald & Willman

 

PROBITTACOMORPHA Freiwald & Willman, 1992: 182. Type species: Probittacomorpha christenseni Freiwald & Willman, 1992, by original designation.


brisaci Krzeminski, Kania & Nel, 2012: 82. PA: France (Miocene) [A].

christenseni Freiwald & Willman, 1992: 182. PA: Denmark (Paleocene/Eocene) [C].



 

 

Genus PROPTYCHOPTERINA Kalugina

 

PROPTYCHOPTERINA Kalugina in Kalugina & Kovalev, 1985: 40. Type species: Proptychopterina handlirschi Kalugina, 1985, by original designation.

amota Lukashevich, 1994: 120. PA: Russia (Siberia) (Middle Jurassic) [C].


evecta Lukashevich, 1994: 119. PA: Kazakhstan (Upper Jurassic) [C].


gracilis Lukashevich, 1994: 117. PA: Kazakhstan (Upper Jurassic) [C].


handlirschi Kalugina in Kalugina & Kovalev, 1985: 41. PA: Russia (Siberia) (Middle Jurassic) [C].


immensa Lukashevich, 2000b: 29. PA: Russia (Siberia) (Upper Jurassic/Lower Cretaceous) [C].

makarova Lukashevich, 2000b: 29. PA: Russia (Siberia) (Lower Jurassic) [C].

mongolica Kalugina, 1988: 84. PA: Mongolia (Lower Cretaceous) [C].


oleynikovi Kalugina, 1989: 71 (1990: 67). PA: Russia (Siberia) (Upper Jurassic/ Lower Cretaceous) [C].


opinata Lin & Lukashevich, 2006: 292. PA: China (Middle Jurassic) [C].

sharategica Kalugina 1992: 113 (1993: 146). PA: Mongolia (Upper Jurassic) [C].


tenera Lukashevich, 2000b: 29. PA: Russia (Siberia) (Upper Jurassic/Lower Cretaceous) [C].

yeniseica Lukashevich, 1994: 119. PA: Russia (Siberia) (Middle Jurassic) [C].

 

 

Genus PTYCHOPTERA Meigen

 

LIRIOPE Meigen, 1800: 14. Suppressed by I.C.Z.N. (1963: 339).


*PTYCHOPTERA Meigen, 1803: 262. Type species: Tipula contaminata Linnaeus, 1758, by subsequent designation of Latreille (1810: 442).


eocenica Podenas, 2007: 195. PA: Baltic Region (Eocene) [A].

miocenica Cockerell, 1910c: 280 (Bittacomorpha). NE: USA (Oligocene) [C].


Unidentified sp. - PA: Russia (Siberia) (Middle/Upper Jurassic) [C] (Brauer et al., 1889: 20).


 

 

Genus PTYCHOPTERULA Handlirsch

 

PTYCHOPTERULA Handlirsch, 1909: 269. Type species: Ptychoptera deleta Novák, 1877, by monotypy.


deleta Novák, 1877: 88 (Ptychoptera). PA: Czech Republic (Oligocene) [C].



 

 

Genus SINOPTYCHOPTERITES Hong

 

SINOPTYCHOPTERITES Hong, 2002a: 151. [Unavailable name; genus-group name based on an unavailable species-group name.]

paludus Hong, 2002a: 153. PA: Chine (Eocene) [A]. [Unavailable name; species-group name proposed after 1999 without specifying a type depository.]

 

 

Genus ZHIGANKA Lukashevich

 

ZHIGANKA Lukashevich, 1995: 169. Type species: Zhigankia comitans Lukashevich, 1995, by original designation.

comitans Lukashevich, 1995: 170. PA: Russia (Siberia) (Lower Cretaceous) [C].

woolgari Lukashevich, Coram & Jarzembowski, 2001: 169. PA: UK (Lower Cretaceous) [C].

 

 

Unplaced Species of PTYCHOPTERIDAE

 

mesozoica Kalugina, 1989: 75 (1990: 73) (Ptychoptera). PA: Russia (Siberia) (Lower Cretaceous) [C].